PESTS OF STORED PRODUCTS

A ‘pest of stored products’ can refer to any organism that infests and damages
stored food, books and documents, fabrics, leather, carpets, and any other dried or
preserved item that is not used shortly after it is delivered to a location, or moved
regularly. Technically, these pests can include microorganisms such as fungi and
bacteria, arthropods such as insects and mites, and vertebrates such as rodents and
birds. Stored product pests are responsible for the loss of millions of dollars every
year in contaminated products, as well as destruction of important documents and
heritage artifacts in homes, offices and museums. Many of these pests are brought
indoors in items that were infested when purchased. Others originate indoors when
susceptible items are stored under poor storage conditions, or when stray individual
pests gain access to them.
Storage pests often go unnoticed because they infest items that are not regularly
used and they may be very small in size. Infestations are noticed when the pests
emerge from storage, to disperse or sometimes as a result of crowding or after
having exhausted a particular food source, and search for new sources of food and
harborage. Unexplained occurrences of minute moths and beetles flying in large
numbers near stored items, or crawling over countertops, walls and ceilings,
powdery residues below and surrounding stored items, and stale odors in pantries
and closets can all indicate a possible storage pest infestation. Infestations in stored
whole grains or beans can also be detected when these are soaked in water, and
hollowed out seeds rise to the surface, along with the adult stages of the pests, and
other debris. Other telltale signs are clumping or webbing of particles. It is
important to recognize these pests and locate the sources of infestation as soon as
possible, to prevent their establishment and spread.
This section will cover insects and mites commonly encountered in storage and
these can be broadly grouped into stored food (or pantry products), and stored
fabric (or closet) pests. However, many of these pests are not restricted to either
group and will infest food, fabric, as well as many other similar items of plant or
animal origin.

PESTS OF STORED FOOD/PANTRY PESTS

Pantry pests covered in this section are grouped into insects (beetles, moths) and
non-insects (mites). The larvae of these moths and beetles are all pale-white or
cream in color and may be difficult to distinguish, but the adults are easier to
identify. The adult beetles will mostly be found with their grubs, feeding on the
stored products, but the adult moths do not feed and will only lay eggs. Mites are
very minute and will need magnification to be detected.
Generally, none of these pests cause harm when consumed in small quantities
along with food materials. However, large infestations can significantly alter the
appearance, taste, flavor and quality of food, as well as create favorable conditions
for growth of secondary fungal and bacterial pathogens. Allergic reactions can be
produced in sensitive people. Certain general pests such as ants are often attracted
to food stored in pantries and may be toxic if consumed in large numbers.

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INSECTS: BEETLES

Common name(s): Bean weevil, seed beetle, pulse beetle

Scientific name, classification: Acanthoscelides spp., Callosobruchus spp., Order:
Coleoptera, Family: Chrysomelidae.
Distribution: Worldwide.
Description and ID characters: Small
beetles, ⅛ - ¼ inch in length, with stout,
rounded oblong bodies, generally colored
grayish brown or dark gray with mottled
patterns. Although they are called ‘weevils’,
they lack the characteristic elongated snout
found in true weevils (Family Curculionidae).
Larvae are small plump, yellowish-cream
colored, legless grubs.
Best identifying features: Short elytra
(hardened front wings that form a shell over
the top of the body) with flattened ends,
which do not cover the tip of the abdomen. Bean beetles
Head is small, black and bent downwards. Photo: Patrick Marquez, Bugwood.org
Entire body is clothed with fine, short hairs.
Eyes are large and protruding. Antennae are well developed, often as long as or
longer than the front legs, with flattened leaf-like segments towards the tip. Adult
beetles are capable of flight, and will fly out in large numbers when infested beans
are disturbed.
Pest status: Important pest of stored legumes such as peas and beans, and
occasionally grains and other foods. Will also attack legume pods on standing crops
in the field.
Damage/injury: Adults and grubs bore into seeds and feed on the contents. Neat,
circular exit holes can be seen on
damaged seeds. Heavy infestation causes
the stored beans to heat up, resulting in
mold growth and change in flavor and
taste. They can chew through plastic and
paper bags to reach stored food.
Life history: Eggs are laid and stuck on
to the surface of beans seeds, and can be
seen as minute white spots. Each seed
may have several eggs on its surface.
Eggs can be laid on the beans in the field, Eggs, exit holes and adults on beans Photo:
before harvest. Emerging larvae bore Originality1988
into the seeds and feed on the kernels,
and feed on several seeds before pupating. Larvae ready to pupate will chew a
round hole on a seed to emerge as adults, and then pupate inside the seed. Pupae
are small and dark brown in color. Duration of life cycle is highly dependent on
temperatures, and takes 3-4 weeks at temperatures above 30oC; longer during
cooler months. 5-6 generations are possible in a year.
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Common name(s): Rice weevil
Scientific name, classification: Sitophilus oryzae,
Order: Coleoptera, Family: Curculionidae. Two
related species, the maize weevil or greater rice
weevil Sitophilus zeamais, and the wheat weevil or
granary weevil Sitophilus granarius are also very
common; the 3 species are often mistaken for each
other.
Distribution: Worldwide.
Description and ID characters: Small dark Rice weevil adult
Photo: Joseph Berger, Bugwood.org
brown, reddish-brown or black elongated weevils,
about 1/16 inch in length. Larvae are small plump,
yellowish-cream colored grubs.
Best identifying features: Pronounced snout
projecting in front of the head, bearing elbowed
antennae with club-shaped tips. Microscopic Rice weevil adult
Photo: Natasha Wright, Bugwood.org
examination shows four reddish-brown or orange
patches on the corners of the wings in rice weevils
and maize weevils (marks on top of wings are
fainter), which are also both capable of flight. The
granary weevil lacks these marks, and is not capable
Maize weevil adult
of flight. Adults of all species will go motionless, Photo: Natasha Wright, Bugwood.org
withdrawing their legs close to their bodies and play
dead when disturbed. Grubs of all species are
small, ‘C’ shaped, plump cream colored, legless
larvae.
Pest status: Important pest of several stored food
grains, mainly rice, wheat, corn and their products Granary weevil adult
such as breakfast cereals, macaroni, noodles, etc. Photo: Jennifer C. Giron Duque

Adult rice and maize weevils can attack standing

grain crops in the field before harvest. All species can also attack other food
materials such as dried fruits and nuts.
Damage/injury: Adults chew on grain and feed on
the contents leaving the hulls if present, while the
grubs develop within them. Together, they reduce
the infested grain over time to a coarse, stale
smelling powder. Individual infested grains have
irregular feeding marks, and some are partially
hollowed out. Heavy infestation causes the stored
grains to heat up, resulting in mold growth and Rice weevil larvae
change in flavor and taste. They can chew through Photo: Pest and Disease Image Library
plastic and paper bags to reach stored food.
Life history: Females chew a small hole on the surface of a grain, lays a single egg
within it and seals the hole with a waxy secretion. The hatching grub feeds within
the grain till pupation, and emerges as an adult by breaking open the almost
hollowed out grain. Duration of life cycle is highly dependent on temperatures,

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and takes 3-4 weeks at temperatures above 30oC; longer during cooler months. Up
to 10 generations are possible in a year.

Common name(s): Cigarette beetle

Scientific name, classification: Lasioderma
serricorne, Order: Coleoptera, Family:
Anobiidae.
Distribution: Worldwide.
Description and ID characters: Small
reddish-brown, stout and rounded beetle,
about 1/16 inch in length, with an oval
outline.
Best identifying features: Head and thorax
Cigarette beetle adults
bent downwards almost perpendicular to the Photo: Pest and Disease Image Library
body, head is not visible when viewed from
the top. Elytra are almost smooth, or with very faint grooves. Body is clothed with
fine, short hairs. Antennae are short and of uniform thickness throughout their
length. Larvae are small plump, creamy-white, legless grubs with a small yellowish-
brown head, and covered with yellowish-brown silky hairs.
Cigarette beetles are very often confused with the drugstore beetle, which belongs
to the same family and often attacks similar stored products. However, drugstore
beetles have elytra with deep lengthwise grooves; their antennae end in a 3-
segmented club and their larvae have shorter hairs. If viewed together, drugstore
beetles may be slightly larger and more elongated, while cigarette beetles may be
shorter and rounded.
Both these beetles are also confused with another common beetle belonging to
family Anobiidae, the common furniture beetle Anobium punctatum, which may be
longer than both, darker in color, with a pronounced hump or upward projection
on their pronotum, resembling a hood.
Pest status: Important pest of stored tobacco and various other dried herbs and
spices, as well as many other stored products such as dried fruits and nuts, cereals,
oilcakes, flour, bone meal, dried flowers, etc.
Damage/injury: Tobacco is the preferred
food of cigarette beetles, which gives them
their common name. All harvested, dried
and processed forms of tobacco are
attacked, which include raw dry bales,
refined cigarettes, cigars, and chewing
tobacco. Adult beetles and grubs bore
through the leaves and leaving holes and
ultimately reducing the tobacco to a fine
Cigarette beetle pupa, adult and larva on a
powder. They also attack and destroy damaged cigar
many other stored products in a similar Photo: Clemson Univ.-USDA
manner.
Life history: Eggs are laid loosely on the stored product, and the hatching grubs
and adults together feed on the material till it is exhausted. Pupation is also takes
place among the food product. Mature larvae create a small cocoon with particles
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of the substrate and their saliva, and pupate within it. Duration of life cycle is
highly dependent on temperatures, and takes 3-4 weeks at temperatures above
30oC; longer during cooler months. 5-6 generations are possible in a year.

Common name(s): Drugstore beetle, bread

beetle, biscuit beetle
Scientific name, classification: Stegobium
paniceum, Order: Coleoptera, Family:
Anobiidae.
Distribution: Worldwide.
Description and ID characters: Small
reddish-brown, elongated beetle, about ⅛ -
1/ inch in length, with elongated-oval
16
outline. Drugstore beetle adults
Best identifying features: Head and thorax Photo: Pest and Disease Image Library
bent downwards almost perpendicular to the
body, head is not visible when viewed from the top. Elytra have distinct and deep
lengthwise grooves. Body is clothed with fine, short hairs. Antennae are short and
end in a 3-segmented club. Larvae are small, plump, creamy-white, legless grubs
covered with short, silky hairs.
Drugstore beetles are very often confused with the cigarette beetle, which belongs
to the same family and often attacks similar stored products. However, cigarette
beetles have elytra that are almost smooth or with very faint grooves; their
antennae are of uniform thickness throughout the length, and their larvae are
covered with long, yellowish-brown silky hairs. If viewed together, drugstore
beetles may be slightly larger and more elongated, while cigarette beetles may be
shorter and rounded.
Both these beetles are also confused with another common beetle belonging to
family Anobiidae, the common furniture beetle Anobium punctatum, which may be
longer than both, darker in color, with a pronounced hump or upward projection
on their pronotum, resembling a hood.
Pest status: Important pest of various dried herbs
and medicinal plants, as well as various other dried
herbs and spices, tobacco, dried fruits and nuts,
cereals, book bindings, leather, etc.
Damage/injury: These beetles have a preference
for dried herbs and other medicinal plants and are
common in prescription drug preparations, giving
them their common name. However, they actually
attack a wider range of stored products than
cigarette beetles, and are even known to bore
through aluminum, tin and lead sheets. They also
infest bird or rodent nests near homes and
buildings. Adult beetles and grubs bore through
the stored product leaving small holes and Larvae of cigarette (above) and
drugstore (below) beetles
ultimately reducing it to a fine powder. Adult Photo: Pest and Disease Image Library
beetles often chew small shot-holes on packaging
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of stored products to emerge, causing some of the powdery remains within to spill
out.
Life history: Eggs are laid loosely on the stored product, and the hatching grubs
and adults together feed on the material till it is exhausted. Pupation is also takes
place among the food product. Mature larvae create a small cocoon with particles
of the substrate and their saliva, and pupate within it. Duration of life cycle is
highly dependent on temperatures, and takes 3-4 weeks at temperatures above
30oC; longer during cooler months. 5-6 generations are possible in a year.

Common name(s): Khapra beetle, cabinet beetle

Scientific name, classification: Trogoderma
granarium, Order: Coleoptera, Family:
Dermestidae.
Distribution: Worldwide.
Description and ID characters: Small dark
brown, stout, rounded oval shaped beetle,
about 1/16 inch in length.
Best identifying features: Closer
observation will reveal that the body is
covered with short, dense hair. Indistinct
reddish-brown or lighter brown patterns may
Khapra beetle adults
be present on the elytra. The head may be Photo: Pest and Disease Image Library
very slightly visible when viewed from the
top, it is bent downwards and is almost fully
covered by the pronotum. Antennae are short
and end in a 3- or 5-segmented club.
Larvae are about ¼ inch long, with a
yellowish-white, distinctly segmented body
covered with short, stiff hairs. The hairs
towards the tail end are longer. The larvae
turn golden or reddish-brown as they mature.
Pest status: Important pest of stored Khapra beetle larva
Photo: James D. Young, USDA APHIS PPQ
products. One of the world’s most destructive
and invasive stored product pests, on quarantine list in many countries including
the U.S. Any suspected occurrences should be reported to the state regulatory
agencies.
Damage/injury: Khapra beetles
attack a large number of stored
products of plant and animal origin,
but prefer grain and cereal products
such as whole grains, flour, noodles,
and similar items. The larvae are
more damaging than the adults. In
stored grains, they are known to
feed partially on one grain before
starting on the next one, and thus a Khapra beetle adult and larvae damaging stored wheat
Photo: Ministry of Ag. And Regional Development
single beetle can damage a large
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quantity of the grain. Adult beetles readily chew through plastic and paper
packaging to reach food sources. Heavy infestations can cause stored products to
heat up leading to mold growth, and reduction in flavor and quality of food items.
Life history:
Adult beetles rarely fly and feed very little during their short lifespan of about 14-30
days. Eggs are laid loosely in the food source and the hatching larvae immediately
begin to feed. They pass through several molts before turning into adults.
Development is highly influenced by environmental conditions, and the larvae can
remain dormant for many years under unfavorable conditions. They can crawl into
tiny spaces, which also protect them from insecticides or other treatments aimed at
stored product pests. Khapra beetles prefer low humidity (less than 2%) and are
poorly adapted to high humidity. This increases the risk of their spread in the hot
and dry conditions in the arid southwest. Therefore it is very important to
recognize the pest and take appropriate quarantine measures if they are noticed.
Introduction and spread is almost always by human activities because the adults
rarely fly.

Common name(s): Flour beetles

Scientific name, classification: Tribolium spp., Order: Coleoptera, Family:
Tenebrionidae. The confused flour beetle T. confusum and the red flour beetle T.
castaneum are the most commonly occurring species.
Distribution: Worldwide.
Description and ID characters: Small,
slender, flattened and elongated, reddish
brown beetles, about ⅛ inch in length.
Best identifying features: Both species
are flattened, elongated reddish-brown
bodies; head and thorax have minute
punctures, elytra have lengthwise ridges
and punctures between the ridges. The
thorax is more-or-less rectangular in shape, Confused flour beetle
and the margins lack teeth. Photo: Sarefo
Confused flour beetles have antennae
that gradually enlarge towards the tip and
end in a 4-segmented club. The thorax is
slightly broader towards the head than
towards the abdomen, but edges are almost
straight.
Red flour beetles have antennae that Red flour beetle
Photo: Natasha Wright
abruptly enlarge at the tip to form a 3-
segmented knob. The thorax has rounded edges, and slightly wider at the middle.
Pest status: Important pest of a wide range of stored grains and their products
such as cereals and flour.
Damage/injury: Do not usually damage whole grains, but found more in broken
bits and flour, or grain previously infested by other pests. The damage is less due to
feeding, but more due to contamination of the stored product with large numbers
of various life stages, dead bodies, cast skins and fecal pellets. They also produce
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pungent substances during their activities that can seriously affect the quality of
grain and result in reduced marketability and consumption by humans and
livestock. Heavy infestations also cause heat buildup in storage containers,
resulting in secondary mold growth.

Red flour beetles infesting corn Red flour beetle life stages
Photo: Western Australia Dept. of Ag. Photo: Pest and Disease Image Library

Life history: Eggs are laid loosely on the stored product, and the hatching grubs
and adults together feed on the material till it is exhausted. Pupation is also takes
place among the food product and pupae are not enclosed in cocoons. Duration of
life cycle is highly dependent on temperatures, and takes 6 weeks at temperatures
above 30oC; longer during cooler months. Up to 10 generations are possible in a
year. Adults are very active and long lived, mostly surviving up to 1 year or more.

Common name(s): Saw-toothed grain beetle

Scientific name, classification: Oryzaephilus surinamensis, Order: Coleoptera,
Family: Silvanidae. A related species, the
merchant grain beetle O. mercator is also very
common; the 2 species are often mistaken for
each other.
Distribution: Worldwide.
Description and ID characters: Small, slender,
flattened and elongated beetle, colored dark-
brown, about ⅛ inch in length.
Best identifying features: 6 saw-tooth like
projections along both side margins of the
pronotum (thorax). Short connecting segment
between the thorax and abdomen. Elytra are
elongated with rounded ends and rows of minute
puncture-like depressions, and completely cover Saw-toothed grain beetle
the abdomen. Prominent eyes and long antennae. Photo: Pest and Disease Image Library
Larvae are yellowish-white with brown head, and
elongated worm-like body with sparse hairs. 3 pairs of short legs can be seen on
the thorax.
The merchant grain beetle is almost identical, except for larger eyes that are located
further away from the neck than in the saw-toothed beetle.
Pest status: Important pest of a wide range of stored products of plant or animal
origin.

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Damage/injury: Attack a large number of stored
products within the pantry and surrounding
rooms. Feeding on germ region of stored seeds is
reported to reduce germination. Adult beetles and
grubs bore through the stored product, and
leaving holes and ultimately reducing it to a fine Saw-toothed grain beetle larva
Photo: Mohammed El Damir
powder. Adult beetles readily chew through plastic
and paper packaging to reach food sources.
Heavy infestations can cause stored products to heat up leading to mold growth,
and reduction in flavor and quality of food items.
Life history: Eggs are laid loosely on the stored product, and the hatching grubs
and adults together feed on the material till it is exhausted. Most of the larvae are
found in the top 2-3 inches of the stored material. Pupation is also takes place
among the food product. Mature larvae create a small cocoon with particles of the
substrate and their saliva, and pupate within it. Duration of life cycle is highly
dependent on temperatures, and takes 3-4 weeks at temperatures above 30oC;
longer during cooler months. 5-6 generations are possible in a year.

Common name(s): Spider beetles

Scientific name, classification: Ptinus spp.,
Mezium spp., Gibbium spp., Order: Coleoptera,
Family: Anobiidae, subfamily Ptininae (sometimes
as family Ptinidae). The smooth spider beetle
Gibbium aequinoctiale, the American spider beetle
Mezium americanum and the whitemarked spider
beetle Ptinus fur, are common species.
Distribution: Worldwide.
Description and ID characters: Small, globular
Smooth spider beetle
brown or black beetles, some with patterns, about Photo: Sarefo
⅛ - ¼ inch in length.
Best identifying features: Spider-like appearance due to relatively large, globular
pear-shaped abdomens and long legs. Elytra
fully covers the abdomen. Head is pointed
downwards and is hidden when viewed from the
top.
Smooth spider beetles are slightly over ⅛ inch
in length, with shiny dark reddish-brown,
abdomen and long, light-brown legs and
antennae.
American spider beetles are about ⅛ inch in
length. Head and thorax are dull yellow and
hairy, thorax is almost cylindrical with blunt
projections on either side, elytra are glossy dark American spider beetle
reddish brown or black. Antennae and legs are Photo: Pest and Disease Image Library

long and slender, and pale yellow or light brown

in color.
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Whitemarked spider beetles are about ¼ inch
in length, and light reddish brown in color, with
patches of white hairs on the elytra. Females have
2 white patches on each elytron that join to form
two transverse bands. Legs and antennae are long,
slender and pale yellow in color.
Larvae of all species are small, plump and C-
shaped, legless grubs with light brown heads.
Adult spider beetles are sometimes confused with
bed bugs, but can be easily distinguished on closer
examination. Whitemarked spider beetle
Pest status: Scavenger and occasional pest of a Photo: Pest and Disease Image Library
wide range of stored products, predominantly
cereal products.
Damage/injury: Spider beetles are general
scavengers and feed on a wide variety of food
items. Occasional infestation of wooden
structures near food sources is reported, in wall
voids and drop ceilings. Larvae cause a typical
"scarring" of the wood in buildings in the
formation of pupal cells prior to pupation. They
do not bite or sting humans or pets, or spread Spider beetle larva
Photo: Pest and Disease Image Library
diseases.
Life history: Eggs are laid in stored products, on outside of packaging or in debris
found in cracks and crevices of storage structures. Larvae feed on the available
food sources and bore into nearby wood or cardboard structures to pupate. Some
species overwinter in the larval stage, and pupate in the following spring, while
some are active year round. 2-3 overlapping generations are possible in a year.

Common name(s): Yellow mealworm

Scientific name, classification: Tenebrio molitor,
Order: Coleoptera, Family: Tenebrionidae.
Distribution: Worldwide.
Description and ID characters: Some of the
largest stored product insects. Adults are shiny
dark brown or black beetles, about ½ - ¾ inch
long. They usually emerge in early summer and fly
to lights.
Best identifying features: The thorax in adults is
covered with fine punctures and the elytra have Yellow mealworm adult beetle
longitudinal grooves. Photo: Clemson Univ.-USDA

Larvae are slender, pale yellow at first and turning

dark yellow and about 1 inch long when mature, with each segment bordered
yellowish brown.
Pest status: Occasional pests of stored grains and other stored products.
Damage/injury: Mealworms feed externally on stored grains and do not bore into
them or cause webbings. Therefore they are not considered serious pests, but they
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can cause damage and contaminate the stored
products with their feces and cast skins. However a
severe infestation can destroy large quantities of
stored products especially when unnoticed for a
long period of time.
Life history: Adults emerge in late spring or early
summer. Eggs are laid and glued to the food source. Yellow mealworm
Photo: Clemson Univ.-USDA
Larvae feed continuously through the summer and
into the fall and overwinter as larvae. They pupate in spring, and emerge as adults
in about 2 weeks. Due to overwintering in the larval stage, only 1 generation
usually occurs in a year.

INSECTS: MOTHS

Common name(s): Angoumois grain moth

Scientific name, classification: Sitotroga cerealella, Order: Lepidoptera, Family:
Gelechiidae.
Distribution: Worldwide.
Description and ID characters: Adult
moths are small, slender, dark tan, buff or
straw-colored moth, about 1⁄3 inch in
length and wingspan of ½ inch.
Best identifying features: Back edges of
both pairs of wings have a fringe of hairs.
Both pairs of wings end in a finger-like
projection, which is more distinct in the
hind wings.
Angoumois grain moth adult. Note fringes
Larvae are creamy white with a dark along back edges of wings and finger-like
head. Pupae are reddish brown and projection at the tip of wings
enclosed in a silken cocoon. Photo: CSIRO
Damaged grains show circular holes, similar to those made by certain beetle pests
of stored grains such as the rice weevil or maize weevil. Locating the adults is
necessary to confirm the source of damage.
Pest status: Occasional pest of stored grains and other food products, and also of
grains (especially corn) in the field.
Damage/injury: Damage is caused entirely by the larvae. They bore into the
kernel of grains and usually spend the entire larval stage and pupation within the
kernel. Damaged kernels are hollow and often have a round exit hole through
which the adult emerged. Infested grains will not germinate. They have an
unpleasant smell which makes them unfit for consumption by humans or livestock.
Infestation can increase moisture content and temperature of the stored products,
attracting mold and other microorganisms. The adult moths do not feed on or
damage stored products, but only visit to lay eggs. Stored ear corn is much more
susceptible to attack than shelled corn. Infestations can start in the field before the
grains are harvested, or can be acquired from existing stored grains.

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Life history: Eggs are laid and glued to the surface of the grains, mostly while still
in the field. The hatching larvae immediately bore into a grain kernel and seal the
entrance hole. They live within the grain and continue to feed and hollow out the
contents, passing through about 3 molts. Mature larvae chew a circular hole on the
surface of the kernel, seal it with silk and then pupate in a silken cocoon within the
grain. The adult moth emerges and flies out through the exit hole, and the cycle
repeats. 4-5 overlapping generations usually occur in a year; many more in
favorable conditions.

Angoumois grain moth larva in damaged grain (left); newly emerged adult near exit hole on a
grain in a ear of corn (right). Photos: Clemson Univ.-USDA Coop. Extension

Common name(s): Indian meal moth

Scientific name, classification: Plodia interpunctella, Order: Lepidoptera, Family:
Pyralidae.
Distribution: Worldwide.
Description and ID characters:
Adults are small shiny, moths, about ¼
inch in length with a wingspan of about
¾ inch. They are the more commonly
seen stage and are attracted to lights.
Best identifying features: In newly
emerged moths, the head and thorax
are reddish brown in color. The top
half of forewings are light-gray or tan,
but bottom half is reddish brown or Indian meal moth (left) and larva (right)
Photo: Pest and Disease Image Library
coppery border with dark stripes.
Edges of the forewings have fringes of
dark gray hairs. Hind wings, usually
seen only during flight, are uniformly
light-gray or tan. Most adult moths
soon lose their characteristic coloration
as their delicate wings brush against
surfaces, and therefore may appear
uniformly tan or gray in color (see
lower picture). Head is pointed, with
Indian meal larvae and pupae
prominent eyes and long slender Photo: Purdue Univ. Extension

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antennae. The moth rests with its forelegs slightly raised, giving the body a sloping
appearance. Larvae vary from pale white, pale pink, light green to almost brown in
color, and are about ½ inch in length. They have 5 pairs of short stubby ‘prolegs’
on the abdomen, which enable them to move about easily in their substrate. Pupae
are brown colored, about ¼ - ¾ inch in length, and may or may not be enclosed in
a silken cocoon.
The adult moths are the indicators of an
infestation although they do not feed on or
damage stored products. Larvae can move away
from their food source, even climbing up walls,
to different locations to pupate, and this can be
misleading when trying to find the source of
infestation.
Pest status: Important pest of a variety of
stored products. Most common moth pest of
stored grains.
Damage/injury: Damage is caused entirely by
the larvae, by feeding on the food source, and Indian meal moth webbing
Photo: Clemson Univ.-USDA
spinning large amounts of silken webs. The
webs serve to protect the larvae from
desiccation, but they attract fecal pellets, debris from the damaged food source, and
cast larval skins, leading to far greater contamination than by the feeding itself. The
webbings can cause great nuisance during processing of grains, and clogging of mill
machinery. Infestation can also increase moisture content and temperature of the
stored products, attracting mold and other microorganisms. Most larvae are
located in the top 2-3 inches of the stored product. Adult moths cannot enter
closed packages, and need an opening such as a hole or a seam to reach the food
source to lay their eggs. The adults feed on nectar and do not damage stored
products.
Life history: Adults mate in 2-3 days after emergence and eggs are laid directly on
the food source. Larvae continue to feed within the food source, and can also
move to other nearby sources. Pupation can also be within the food source, or in
other distant locations. Duration of life cycle is highly dependent on temperatures,
but is usually completed in 4-5 weeks under optimal conditions. Several
overlapping generations are possible in a year.

Common name(s): Mediterranean flour moth

Scientific name, classification: Ephestia kuehniella, Order: Lepidoptera, Family:
Pyralidae.
Distribution: Worldwide.
Description and ID characters: Adult
moths are small, slender, shiny pale gray
moths, about ½ inch in length and
wingspan of ¾ to 1 inch. Larvae are small
white or pinkish colored larvae, with dark Flour moth
head and small black spots all over the Photo: Mark Dreiling

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body. Pupation is in silken cocoons, often far away from the food source. Pupae
are reddish brown in color.
Best identifying features: Forewings are pale pinkish gray with several dark spots,
a broad dark band towards the middle, and two broken zig-zag lines towards the
tips. Hind wings are uniformly pale gray or white, with a thin dark border all
around. The moth rests with its forelegs slightly raised, giving the body a sloping
appearance.
Pest status: Occasional pest of stored grains
and other food products.
Damage/injury: Damage is caused entirely
by the larvae, by feeding on the food source,
and spinning silken webs. The webs along with
fecal pellets, debris from the damaged food
source, and cast larval skins, add to the Flour moth webbing
contamination caused by the larvae and their Photo: Clemson Univ.-USDA
feeding. Infestation can also increase moisture
content and temperature of the stored products, attracting mold and other
microorganisms. They can also cause hindrances during processing of grains.
Adult moths cannot enter closed packages, and need an opening such as a hole or a
seam to reach the food source to lay their eggs. The adults feed on nectar and do
not damage stored products.
Life history: Female moths lay their eggs directly on the food source. Larvae on
hatching immediately begin to spin their webs and feed voraciously on the food
source till they are ready to pupate. Mature larvae will leave the immediate area
near the food source to pupate. Life cycle, as with all stored product pests, is highly
dependent on temperature, and is usually completed in about 7 weeks under
optimal conditions. 4-5 overlapping generations can occur in a year.

INSECTS: PSOCIDS

Common name(s): Psocid

Scientific name, classification: Liposcelis spp.
is a common species, Order: Psocoptera,
Family: Liposcelididae.
These tiny, delicate, soft-bodied insects may
occasionally be seen in stored products such as
cereals, or paper and cardboard containers used
to store these items. Damage is rarely
significant. (See detailed description under
‘Minor Household Pests’).
Psocids on wheat grains
Photo: Pest and Disease Image Library

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NON-INSECTS: MITES

Common name(s): Grain mite, flour mite

Scientific name, classification: Different genera. Order: Acariformes-
Trombidiformes (Subclass Acari/Acarina), Family: Acaridae. Acarus siro and
Glycophagus destructor are cosmopolitan
species.
Distribution: Worldwide.
Description and ID characters: Tiny,
microscopic arthropods with eight legs,
about 1⁄32 inch or lesser in length.
Best identifying features: Need
magnification to identify. Might be
barely visible with the naked eye, as
minute, shiny, soft creamy white
rounded particles. Colors may vary with
the food source, from pale gray to pink. Grain mites on wheat grains
Pest status: Occasional pest of a Photo: Pest and Disease Image Library
variety of stored products.
Damage/injury: The mites feed on
different stored products of plant or
animal origin including grain, flour,
cereal products, cheese, dried fruit, hay,
leather, dried herbs, etc. In severe
infestations, the mites can be seen as a
layer of fine powder over the host
material and they give off a ‘minty’ or
similar sweet-sickly odor when crushed.
This renders food material unsuitable Grain mite among cereal particles
for consumption. Grain mites are Photo: Canadian Grain Commission
known to transmit pathogenic microbes
and produce substances called ‘allergens’ that can cause allergic reactions in some
people.
Life history: Adult females lay eggs in the food material, and which hatch into
larvae (with 6 legs), which then mature into nymphs and later adults (with 8 legs).
Life cycle from egg to adult can be completed in about 14 days at normal room
temperatures. These mites prefer warm, humid environments and thrive in poorly
maintained pantries or food storage areas which have high moisture levels.

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PESTS OF FABRIC/CLOSET PESTS

BEETLES

Common name(s): Carpet beetle

Scientific name, classification: Anthrenus spp., Attagenus spp., Order: Coleoptera,
Family: Dermestidae. The black carpet beetle Attagenus unicolor, the furniture carpet
beetle Anthrenus verbasci and the varied carpet beetle Anthrenus flavipes are common
species in the southwest, and throughout the U.S.
Distribution: Worldwide.
Description and ID characters: Small, dark, rounded oval shaped beetles, about
⅛ inch in length, some with patterns, resembling lady beetles. Larvae are short,
plump worm-like grubs covered with long hairs.
Best identifying features:
Black carpet beetles are shiny black or dark
brown, with brown legs. Larvae are light or reddish
brown, and elongated, about ¼ to ½ inch in length,
much longer than the adults. They are smooth and
shiny and carrot-shaped, the body broader towards
the head and tapering towards the other end, ending
in a tuft of long hairs. The rest of the body is also Black carpet beetle adult and larva
Photo: Clemson Univ.-USDA
covered with shorter, sparser hair.
Furniture carpet beetles are patterned with black,
white, brown and tan scales, which fall off as they
age, and older adults often look uniformly dark
brown or black, with white undersides. Larvae are
pale yellow, turning dark brown with distinct light
and dark stripes, as they grow. Their bodies are
broader towards the head and taper towards the
rear, ending in a tuft of hairs. The rest of the body is Furniture carpet beetle adult, larva
also covered with long, dense tufts of hair. Photo: André Karwath
Varied carpet beetles are similar to furniture
carpet beetles, with black, white, brown and yellow
patterns, that may fade as they age making the
adults look uniformly brown or black. Larvae are
plump dark yellowish brown grubs, broader
towards the rear end, slightly larger than adults, and
covered with long, stiff bristly hairs. If disturbed
they will freeze and hold their hair upright,
appearing like a small clump of fibers on their
substrate.
Larvae of all species leave numerous cast skins Varied carpet beetle adult and larvae
Photo: Clemson Univ.-USDA
which are also covered with bristles or hairs, and
they damage larger continuous areas, and these can help to determine if the causes
of damage are carpet beetles or clothes moths. Clothes moth larvae leave behind
smaller, thin and delicate cast skins and webbings or cases, and their damage is in

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the form of small holes distributed over the fabric, which are not as extensive as
damage by carpet beetles.
Pest status: Occasional pest of fabrics and similar items, rarely of stored food. Can
produce allergic reactions in sensitive people.
Damage/injury: Damage is caused entirely by the larvae, by feeding on the food
source. They attack various kinds of fabrics and similar products, with a preference
for those of animal origin, such as silk, wool, leather, furs, feathers, hair, felt,
bristles in brushes, taxidermy mounts, etc.
Fabrics of plant origin such as cotton and
jute are also readily attacked, as well as
upholstery, carpets and tapestries and
similar materials. Blended fabrics
containing synthetic and natural fabrics are
attacked, but purely synthetic fabrics such
as polyester or nylon are not usually
attacked. Damage occurs more in dark Carpet beetle damage on a sweater
hidden and undisturbed spots such as Photo: Clemson Univ.-USDA
cracks and crevices of furniture, inside
rolls of carpets or rugs, or in folds and hems of clothing. Larvae are also known to
attack various food substances.
Some people are allergic to the larvae, and their hair or bristles, and experience
intense irritation, itching and other symptoms on the skin and respiratory tracts. In
most cases, these reactions were the result of individuals becoming sensitized to the
larvae by long-term exposure.
The adults do not feed on fabrics or stored goods, but fly outdoors to feed on
plants with abundant pollen and nectar, and therefore are often transported
indoors with flowers. They come indoors only to lay eggs. If they cannot gain
access indoors, the beetles are known to infest outdoor habitats such as nests of
birds and other small animals. Most infestations indoors are caused by adults that
are brought in on infested material, or cut flowers. Once indoors, the adults can fly
within a home or building, seeking spots to lay their eggs, such as in air ducts,
closets, under furniture or under baseboards.
Life history: Eggs are laid directly on the food source, the preferred ones being
woolen carpets or furs. The hatching larvae feed voraciously, and remain in dark,
undisturbed spots on the food source. Larval periods can vary widely in duration,
sometimes extending up to 3 years, but are completed in about 3 months under
favorable conditions. As they near pupation, the larvae move away from the
immediate food source, to escape cannibalism, which is exhibited when several
larvae exist in close contact. Many species will overwinter or undergo a period of
dormancy just before pupation. Pupation may be in or near the food source, or in a
concealed spot, or even within the last larval skin, if no shelter is available. Adults
usually emerge in late summer to early fall. They live for about 2 weeks, within
which they mate, locate suitable spots and lay eggs. Up to 2 overlapping
generations may occur in a year.

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 A coat damaged by case-making clothes moths
Photo: Dawn Gouge

A floor rug damaged by carpet beetles. Note the larva towards the center of the picture.
Photo: Dinakar Radhakrishnan

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MOTHS

Common name(s): Clothes moth

Scientific name, classification: Tinea spp., Tineola spp., Order: Lepidoptera,
Family: Tineidae. The webbing clothes moth Tineola bisselliella and the case-making
clothes moth Tinea pellionella are most common.
Distribution: Worldwide.
Description and ID characters: Small, slender and shiny yellowish-gray moths,
about ¼ inch in length and wingspan of ½
inch, much smaller than moths that infest
stored foods.
Best identifying features:
Webbing clothes moths are more
yellowish with a golden sheen, and their
wings are fringed with golden hairs. The Webbing clothes moth
head has a tuft of reddish brown hairs. Photo: Olaf Leillinger

Case-making clothes moths are more

grayish or silvery in color, with faint dark
spots on their wings. The head has a tuft of
yellow hairs.
Adults of both species are weak flyers and
try to avoid light. They are usually found in
dark, undisturbed areas of closets. When Case-making clothes moth
disturbed, they try to escape by running, Photo: ©Entomart
rather than flying and if necessary they will
flutter.

Webbing clothes moth larva creating web Case-making clothes moth larvae with cases
Photo: Guido Gerding Photo: ©Entomart

Larvae of both species are similar in appearance, small white or cream colored
caterpillars with a dark brown head, about ½ inch in length. Webbing clothes moth
larvae are mostly found under a web of silk, fabric debris and feces, whereas case-
making clothes moth larvae are always enclosed in a silken, spindle shaped case that
they carry around as they move.
Larvae leave behind small, delicate cast skins and webs or cases, and small holes
distributed over the fabrics. These can help to distinguish their damage from that

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caused by carpet beetles, which leave behind larger, shell-like cast skins covered
with long bristles or hairs, and large contiguous areas of damaged fabric.
Pest status: Occasional pest of fabrics
and similar items, but rarely of stored
food.
Damage/injury: Damage is caused
entirely by the larvae, by feeding on the
food source. All kinds of fabric including
cotton, silk, wool, furs, feathers, hair, felt,
bristles in brushes, taxidermy mounts,
upholstery, carpets and tapestries and
similar materials of animal or plant origin
are attacked. Wool is probably their most
preferred food. Blended fabrics
containing synthetic fabrics along with Webbing clothes moth, larva, pupa, damage
cotton, silk or wool are also attacked, but Photo: Clemson Univ.-USDA
purely synthetic fabrics such as polyester
or nylon are not usually attacked. Damage occurs more in hidden spots on the
fabrics, such as under collars and cuffs, in folds and hems, cracks and crevices of
furniture and rolls of carpets or rugs. Fabrics stained with perspiration, urine, or
food have higher chances of being attacked. The adults do not feed, and exist
solely to reproduce. They visit stored products only to lay eggs and do not cause
any other damage. Clothes moths have also been reported to live in bird nests, and
also feed on cobwebs and other fibers.
Life history: Adults mate soon after their emergence, and the females lay clusters
of eggs, which they stick to the food source with a glue-like substance. They die
soon after egg laying, males may live slightly longer than females but for a month at
most. The eggs hatch into larvae that immediately begin to feed, and the webbing
larvae make webs and the case-making larvae make their cases. The webs and cases
are mostly similar in color to the substrate color, because they contain fibers from
the substrate. Webbing larvae extend their web tunnels as they move about, and
the webs accumulate their feces, debris
and fibers. The case-making larvae
remain within their case throughout their
larval period, the case increasing in size
as they grow. They carry the case around
with them as they move on the substrate,
extending their heads out from either end
of the case to feed. Webbing larvae Case-making clothes moth larval case
pupate in silken cocoons on the food Photo: Kathryn Robinson

source, but case-making larvae often

move away from their feeding sites to
pupate in cracks or crevices nearby.
Life cycles are temperature dependent,
but are usually completed in 4-6 months,
and up to 2 generations are possible in a Case-making clothes moth pupa
year. Photo: ©Entomart

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Note: Mothballs should not be used to control fabric pests. Mothballs contain
either naphthalene or paradichlorobenzene, both of which are toxic and can
produce harmful effects when inhaled.

Sources, further information:

Food and fabric pests
http://www.ipm.ucdavis.edu/PMG/menu.house.html#DESTROY
Recognizing and controlling insect pests of stored products
http://citybugs.tamu.edu/factsheets/household/food-fabric/ent-2007/
Stored product pests in the home
http://extension.uga.edu/publications/detail.cfm?number=B1378

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