Chap-07_Structural Organisation in Animal-II (28)_E
Chap-07_Structural Organisation in Animal-II (28)_E
Chap-07_Structural Organisation in Animal-II (28)_E
Syllabus
Morphology, anatomy and functions of different systems (digestive, circulatory, respiratory, nervous
and reproductive) of an insect (cockroach). (Brief account only).
Chapter Index
EARTHWORM
Indian Earthworms (Pheretima posthuma)
Phylum : Annelida
Class : Oligochaeta
Genus : Pheretima
Species : posthuma
There are several types of Earthworms. The most common genus of Earthworm is Pheretima
in India and Lumbricus in Europe. Pheretima has 500 species, 13 of them are found in India.
Habitat :
Earthworm is reddish brown terrestrial animal which inhabits upper layer of moist soil where it
lives inside burrows during day time.
Earthworm inhabits those soils which have abundant organic matter.
An acre of good moist soil can have upto 50,000 animals. Burrow is made by boring and
swallowing the soil.
Internal Morphollogy
The body wall of earthworms is thin, soft and slimy. From the surface inwards, it consists of
cuticle, epidermis, muscular layers and coelomic epithelium
(1) Cuticle: It is thin and elastic non-cellular protective membrane. It is formed of collagen fibres
secreted by underlying epidermis.
Locomotion
The earthworm does not have specialized locomotory organs.
The locomotion is brought about by the circular and longitudinal muscles of the body wall,
aided by the chitinous curved setae embedded in the skin.
Due to the contraction of the circular muscles of the anterior end the latter becomes thin,
elongated and extends forwards.
At the same time, the setae of the anterior end hold the ground firmly and prevent the animal
from slipping backward.
Now the circular muscles of the anterior end relax and the longitudinal muscles contract.
It causes the shortening and thickening of the anterior segments, and thus, the posterior part of
the body is pulled ahead.
The process is repeated and the worm is able
to move forward with speed. Earthworms
move at the rate of about 15 cm per minute.
Alimentary Canal
It is a straight tube and runs between first and
last segment of the body.
In 1-3 segments buccal cavity is present,
pharynx in 4th segment.
From 5-7 segments oesophagus is present.
Gizzard is present in 8th segment.
From 9-14th segments is a tubular stomach.
Calciferous glands are present in the stomach,
which produce CaCO3 to neutralise the humic
acid.
Intestinestarts from the 15th segment on wards
and continues till the last segment.
A pair of shortand conical Intestinal caecae
projects from the intestine on the 26th
segment. They secrete amylolytic enzyme
which digests starch. Other enzymes are
lipase, cellulase, invertase etc.
Circulatory system
Earthworm is first to evolve a closed circulatory system in the evolution of animals.
The respiratory pigment is haemoglobin which remains dissolved in the plasma, RBC are
absent.
Blood glands are present on 4, 5 and 6th segments, they produce blood cells and haemoglobin.
There are 2 main blood vessels.
Dorsal vessel is the largest vessel and blood flows forwards from postrior end to anterior end.
They have contractile wall, and valves are present. Before the 13th segment, it is distributing
vessel and behind 13th segment, it is a collecting vessel.
Ventral vessel is the main distributing vessel, blood flows backwards from anterior to posterior
end, valves are absent.
Latero-oesophageal vessels are paired vessels which extend from first to thirteenth segment.
EXCRETION
Concept Builder
Earthworm mainly removes the nitrogenous waste in the form of urea in soil. But when plenty of water is availabl
Excretory System
Excretory organs occur as segmentally arranged coiled tubules called nephridia. They deliver
the wastes through a pore to the surface of body wall or into digestive tube.
(i) Three main types of nephridia are:
(a) Pharyngeal nephridia are situated in the
segments 4,5 and 6. They open in the
anterior part of alimentary canal, i.e.,
buccal cavity and pharynx. They are
without nephrostome and are
enteronephric type.
(b) Integumentary nephridia are scattered
in the body wall. They are smallest, v-
shaped, without nephrostome and are
exonephric type. In clitellar segments
they form forests of nephridia.
(c) Septal nephridia are the largest, attached to both faces of each intersegmental septum behind
15th segment.
(ii) Septal nephridia are the only nephridia with nephrostome or funnel. The terminal duct opens
into septal excretory canal. These canals, in turn, open into two supraintestinal excretory
canals. Septal nephridia are enteronephric, finally excretory products are poured into intestine.
Enteronephric condition is an adaptation for the conservation of water or osmoregulation.
(iii) Excretory products of earthworm are urea (about 50%), ammonia (about 40%) and traces of
creatinine. Earthworms are mainly ureotelic.
SENSE ORGANS
Group of specialised cells found in skin and lining of buccal cavity.
Photoreceptors located in prostomium and dorsal epidermis. Perceive light intenSity with the
help of phaosome (Optic organelle).
Thigmoreceptors located in ventral and lateral epidermis.
Olfactoreceptors located in lining of buccal cavity.
Gustatory receptors located in lining of buccal cavity.
RESPIRATION
Earthworm has no special respiratory organs.
Gaseous exchange takes place simply through the skin, which is thin and highly vascular.
Effective gaseous exchange takes place only when the skin is moist.
The skin is kept moist due to the damp earth, secretion of the mucus by the epidermal gland
cell and oozing of coelomic fluid through the dorsal pores.
REPRODUCTION
Earthwonn is hermaphrodite.
There are two pairs of testes present in 10 th and
11th segments. They are surrounded by 2 testes
sacs lying ventrally, one in the 10 th and the other
in 11th segment.
There are 2 pairs of seminal vesicles, one pair in
11th and the other pair in 12th segment.
Maturation of sperms occurs in seminal
vesicles.
The testis sac of 10th segment communicates
with the seminal vesicles of 11 th segment and
testis sac of 11th segment with seminal vesicles
of 12th. From each testis sac, vas deferens carries
the sperms up to 18th segment where they join
the prostatic duct from prostate gland.
Four pairs of spermathecae are present, one pair
in each ofthe6, 7, 8, and 9 segments. They
receive and store the spennatozoa of another
earthworm during copulation.
One pair of ovaries lies in 13th segment, which open through median aperture on 14th segment.
Accessory glands are present on the ventral surface of 17th and 19th segments, which open
through the genital papillae. These are a part of male reproductive system.
Self Assessment
All the following statements are correct about the reproductive system of earthworm, but one is
wrong. Which one is wrong?
(1) There are two pairs of testes one pair in the 10th and one pair in 11th segment
(2) One pair of ovaries attached at the inter-segmental septum of the 12th and 13th segments
(3) Two pairs of accessory glands one pair each in 18th and 19th segments
(4) Four pairs of spermatheca are located in 6th-9th segments, one pair in each segments
The main function of clitellum is
(1) Cocoon formation (2) Locomotion (3) Excretion (4) Copulation
Given below are four statements A to D, select the option in which two statements are correct? A:
In earthworms development is indirect and the larva is trochophore larva.
B : A mutual exchange of sperms occurs between two worms during mating
C: Fertilisation is internal occurs within the cocoon deposited in soil
D : About 3 weeks are required for development, each cocoon produces two to twenty baby
worms with an average of four.
(1) A&B (2) B&C (3) B&D (4) C&D
When plenty of water is available earthworm is
(1) Ureotelic (2) Ammonotelic (3) Uricotelic (4) Aminotelic
Earthworm has a central nerve cord which is
(1) Single (2) Ventral and solid (3) Dorsal and hollow (4) Double and dorsal
Which of the following statement is incorrect about the circulatory system of earthworm?
(1) It exhibits a closed type of blood vascular system
(2) Blood glands are present in 4th, 5th and 6th segments
(3) Haemoglobin is the respiratory pigment which is dissolved in plasma
(4) Ventral vessel is the largest distributing vessel in which the blood flows from posterior to
anterior side
Which of the following is not a correct matching of the part of alimentary canal, its location and
function in earthworm?
Thorax
(i) It comprises of three segments -prothorax, mesothorax and metathorax.
(ii) The three thoracic segments are covered by relatively thicker and larger tergites called nota.
(iii) The notum of prothorax called pronotum is very large and covers the neck also. Each of the
mesonotum and metanotum bears a pair of wings.
Abdomen
It is the largest and the broadest, relatively more
flattened and softer part behind the thorax.
There are ten tergites. In both males and females, the
8th and 9th tergites are mostly covered by the 7th. The
10th tergum is somewhat bowl-shaped and
posteriorly bifurcated into two lobes.
Ventrally, the abdomen has 9 sternites in male and 7
in females.
In females the last sternite (7th) is larger and boat
shaped and together with indistinct 8 th and 9th
sternites it forms a chamber Anal styles like structure
called gynatrium, posterior part of this chamber is
MALE called oothecal chamber. Behind this
chamber, 7th sternite Opening of oothecal chamber bifurcates into two prominent oval plates
called apical lobes. Female gonopore is located between them.
In males, the 9th sternite bears a pair of spine like anal-styles.
Thoracic spiracles are somewhat larger. One pairofthese is between prothoraxand mesothorax
and the other between mesothoraxand metathorax, upon respective pleurites.
The first pair ofabdominal spiracles are dorsolateral upon tergite offirstabdominal segment,
butthe remaining seven pairs are present upon the pleurites of second to eighth segments.
Each spiracle is surrounded by a ring-like sclerite called peritreme.
Mechanism
Several tergo-stemal muscles extend vertically between the tergites and sternites of all
abdominal segments.
Harmonious contractions and relaxations ofthese at regular intervals cause rhythmic expansion
and compression of abdomen leading to inspiration (with relaxation) and expiration (with
contraction) of air.
At rest, the oxygen requirement is less, tracheolar ends get filled with tissue fluid.
The movement of O2 is along the pressure gradient as the tracheolar ends are losing oxygen to
the cells for performing cellular respiration.
O2 requirement increases during activity.
Tracheolar fluid is withdrawn out of Tracheoles.
Alternate expansion and contraction of abdominal cavity occurs involving tergosternal muscles
and abdominal muscles.
High level of CO2 in abdominal cavity make tergo-sternal muscles and abdominal muscles to
contract pushing out the air from tracheal system to the outside through spiracles.
Heart
It is a long elongated tube situated in the mid dorsal line of
thorax and the abdomen immediately beneath the terga.
Heart consists of thirteen chambers.
The last two posterior chambers are very small.
The chambers are separated from one another by deep
constrictions.
The opening of each chamber into another is guarded by
valves which allow blood from behind forward.
The Blood Sinuses
The large body cavity or haemocoel is divided by two
membranous horizontal partitions, into three wide and
flattened sinuses-the dorsal pericardial sinus containing the
'heart', the middle perivisceral sinus containing the gut, and
the ventral perineural sinus or sternal sinus containing the nerve cord.
The partition between pericardial and perivisceral sinuses is called dorsal diaphragm and
between perivisceral and perineural sinuses is called ventral diaphragm.
The sinuses intercommunicate by pores in the respective diaphragms.
A pair of fan like, triangular alary muscles in the floor of the pericardial sinus in each segment
reinforce the dorsal diaphragm by their broad bases and also connect it, by their pointed tips
with the tergite of the segment.
Circulation of Haemolymph
The pumping force that propels the haemolymph is provided by the pulsations of the 'heart'.
The respiratory movements of abdomen and contraction of alary muscles increase this force.
From the pericardial sinus, the haemolymph enters into heart through ostia. When the heart is
filled it contracts from behind forwards. This is its systole phase. Soon the heart becomes
relaxed in its diastole phase. Then the next systole follows after a short time interval called
diastasis. Thus heart pulsates about 50 times/minute.
During systole, the valve like ostia close, preventing back flow of haemolymph into the
pericardial sinus. Therefore, some of its haemolymph is pumped into segmental vessels while
most of it is poured into the head sinus through the terminally opening anterior aorta.
Excretion in Cockroach
Cockroach is uricotelic. In cockroach, following structures help in excretion:
(i) Malpighian tubules
(ii) Fat bodies
(iii) Nephrocytes
(iv) Cuticle
(v) Uricose glands in some species.
Malpighian tubules:
Malpighian tubules are attached at the junction of midgut and hindgut. Excretory products,
dissolved in haemolymph are absorbed by malpighian tubes are discharged into hindgut.
Fat bodies:
Some fat bodies are also present in haemocoel which have mycetocytes, urate cells,
oenocytes and trophocytes.
Nephrocytes :
Nephrocytes present in lateral wall of heart and help in excretion and store nitrogenous waste.
Uricose glands:
In some species, in males, uricose glands are present on the periphery of mushroom glands.
These glands synthesize uric acid . Malpighian tubules are analogous to mammalian kidneys.
Fat bodies are analogous to vertebrate liver.
External Morphology
Skin is made up epidermis and dermis. Mucous glands are present in the dermis and their ducts
open at the surface.
Blood capillaries and pigment cells (chromatophores) are present in the dermis.
Skin is without scales or any other cover or exoskeleton.
Body is divisible into head and trunk, neck is absent. The trunk is provided with a pair of fore
and hind-limbs. The hind-limbs are much larger and muscular than the fore-limbs. Fore-limbs
end in four digits and the hind limbs end in five digits. The digital formula of fore-limbs is
02233. The digital formula of hind limbs is 22343.
Shank or crus is associated with hind limbs.
Sexual dimorphism :
The male ( ) and female ( ) frogs exhibit certain differences in their external features,
which become more pronounced during breeding season.
Generally, male frogs are larger than females.
During breeding season, however, the females become bloated with large ovaries and
numerous ova, and appear considerably larger.
Only the males possess a pair of ventro-Iateral, wrinkled pouch-like vocal sacs located a little
behind the mouth.
These sacs become specially large and distensible, in breeding season.
By inflating these repeatedly with air from the lungs, the males produce a loud croaking sound
meant to call the females for copulation (amplexus).
Internal Morphology
Digestive System
Since the frogs are carnivorous, their
alimentary canal is short in length.
Tadpole larva is herbivorous so
alimentary canal is very long and coiled
in the form of spring.
The mouth is present as a terminal, wide
opening.
It opens into bucco-pharyngeal cavity,
which contains numerous maxillary teeth
arranged along the margin of the upper
jaw and vomerine teeth are present in the
roof of the buccopharyngeal cavity.
The lower jaw is toothless. Salivary
glands Small intestine are absent.
Opening of eustachian tube, vocal sacs
(only in male) gullet and glottis can be
seen clearly in the bucco-pharyngeal cavity.
Respiratory System
Three types of respiration: cutaneous, buccopharyngeal and pulmonary occur.
Cutaneous respiration on land is through the body surface. During hibernation and
aestivation, frog respires only through this method.
Buccopharyngeal respiration occurs through the lining of buccal cavity. It occurs only when
frog is out of water. The mucus membrane of the buccal cavity is moist which dissolves
oxygen whose diffusion occurs into the blood capillaries.
Pulmonary respiration: Lungs in frog are not efficient respiratory organs because only mixed
air enters into them and they mainly function as hydrostatic organs.
Lungs are a pair of thin walled, translucent sacs with inner surface divided into alveoli by
septa. Pulmonary respiration has a maximum frequency of 20/minute.
It occurs when more energy is required. Mouth and gullet are kept closed during pulmonary
respiration.
Respiratory movements in pulmonary respiration are because of buccopharyngeal cavity which
acts as a force pump.
These movements are carried out by set of paired muscles -sternohyal and pterohyal muscles.
Circulatory System
Circulatory system is closed type.
The heart lies enclosed by a thin, transparent, two layered sac, pericardium.
Frog's heart is a three chambered structure made of two upper auricles and a single lower
ventricle.
The two additional chambers connected to the heart of frog are sinus venosus and truncus
arteriosus.
Frogs also possess two well developed portal systems: Renal portal system and Hepatic portal
system. Frog also has two pairs of lymph hearts.
Eye
Eye is guarded by immovable upper eyelid, movable lower eyelid and transparent nictitating
membrane.
Ear
Ear of frog has only middle and internal ear.
Tympanic membrane is present at the body surface.
Middle ear has single bone called columella auris.
Its outer end is attached to the ear drum while inner end to the stapedial plate.
Pressure of air in middle ear is controlled by eustachian tubes.
Membranous labyrinth or internal ear consists of utriculus, sacculus and semicircular canals.
Endolymph fills the membranous labyrinth .
Excretory System
The main organ of excretion is a pair of kidneys.
These are compact, dark red and bean like structures situated little posteriorly in the body
cavity on both sides of vertebral column.
The frog excretes urea, thus is a ureotelic animal.
Urea is carried by blood into the kidney where it is separated and excreted.
Each kidney is composed of several structural and functional units called uriniferous tubules or
nephrons.
Ureter emerges from the kidney as urinogenital ducts in the males.
A common ureter opens into the cloaca.
A thin-walled urinary bladder is present ventral to rectum which also opens in the cloaca.
Reproductive System
Male reproductive organs consist of a pair of yellowish ovoid testes, which are found adhered
to the upper part of kidneys by a double fold of peritoneum called mesorchium.
Vasa efferentia are 10-12 in number and after arising from testes, run through the mesorchium
and enter the kidneys of their side.
In kidneys, these open into Bidder's canal which finally communicates with the urinogenital
duct.
This duct emerges from the kidneys and finally opens into the cloaca.
The cloaca is a small, median chamber that is used to pass faecal matter, urine and sperms to
the exterior.
Ans. Q.51 (4), Q.52 (3), Q.53 (4), Q.54 (4), Q.55 (2), Q.56 (4), Q.57 (3), Q.58 (4), Q.59 (2),
Q.60 (2)
Section – B
Body of nephridia is V-shaped in-
(1) Septal nephridia (2) Integumentary nephridia
(3) Pharyngeal nephridia (4) All the above
Which of the following is not correctly matched-
(1) Lumbricus-Europe (2) Megascolex & Drawida - South India
(3) Pheretima - Shri Lanka, North India (4) Eutyphaeus - N. America
Clitellum of Pheretima is-
(1) Permanent (2) Multilayered (3) Single layered (4) 1 & 2 both
Which structure secretes hormone in Earthworm-
(1) Nerve cord (2) Lymph gland
(3) Cerebral ganglia (4) Chloragogen cells
Answer Key
Section–A
Q.1 3 Q.2 1 Q. 3 4 Q.4 1 Q.5 4 Q.6 2 Q.7 2
Q.8 4 Q.9 4 Q.10 2 Q.11 4 Q.12 3 Q.13 3 Q.14 1
Q.15 2 Q.16 3 Q.17 2 Q.16 1 Q.19 3 Q.20 4 Q.21 2
Q.22 2 Q.23 4 Q.24 2 Q.25 1 Q.26 2 Q.27 2 Q.28 2
Q.29 2 Q.30 4 Q.31 3 Q.32 3 Q.33 1 Q.34 2 Q.35 2
Q.36 3 Q.37 2 Q.38 2 Q.39 2 Q.40 1 Q.41 3 Q.42 3
Q.43 3 Q.44 1 Q.45 2 Q.46 2 Q.47 1 Q.48 3 Q.49 4
Q.50 2 Q.51 4 Q.52 2 Q.53 2 Q.54 3 Q.55 2 Q.56 2
Q.57 1 Q.58 3 Q.59 4 Q.60 2 Q.61 3 Q.62 3 Q.63 3
Q.64 2 Q.65 1 Q.66 4 Q.67 4 Q.68 2 Q.69 2 Q.70 2
Q.71 3 Q.72 3 Q.73 2 Q.74 1 Q.75 2
Section–B
Q.1 2 Q.2 4 Q.3 4 Q.4 3 Q.5 4 Q.6 1 Q.7 4
Q.8 3 Q.9 2 Q.10 3 Q.11 3 Q.12 3 Q.13 4 Q.14 4
Q.15 3 Q.16 2 Q.17 1 Q.18 4 Q.19 1 Q.20 2 Q.21 3
Q.22 3 Q.23 1 Q.24 2 Q.25 4 Q.26 1 Q.27 3 Q.28 4
Q.29 1 Q.30 2 Q.31 1 Q.32 1 Q.33 1 Q.34 3 Q.35 3
Q.36 4 Q.37 3 Q.38 4 Q.39 4 Q.40 2 Q.41 3 Q.42 3
Q.43 1 Q.44 1 Q.45 4 Q.46 4 Q.47 1 Q.48 3 Q.49 3
Q.50 1 Q.51 4 Q.52 1 Q.53 2 Q.54 4 Q.55 3 Q.56 3
Q.57 1 Q.58 1 Q.59 4 Q.60 3 Q.61 2 Q.62 1 Q.63 3
Q.64 1 Q.65 1 Q.66 2 Q.67 4 Q.68 4 Q.69 1 Q.70 2
Q.71 4
Answer Key
Q.1 2 Q.2 2 Q.3 2 Q.4 4 Q.5 4 Q.6 1 Q.7 4
Q.8 3 Q.9 3 Q.10 1 Q.11 4 Q.12 3 Q.13 1 Q.14 3
Q.15 3 Q.16 1 Q.17 2 Q.18 4 Q.19 1 Q.20 1 Q.21 1
Q.22 2 Q.23 1 Q.24 3 Q.25 2 Q.26 2 Q.27 3 Q.28 3
Q.29 2 Q.30 2 Q.31 3 Q.32 2 Q.33 1 Q.34 1 Q.35 3
Q.36 2 Q.37 3 Q.38 1 Q.39 1 Q.40 2 Q.41 3 Q.42 3
Q.43 2 Q.44 3 Q.45 4 Q.46 1 Q.47 2 Q.48 4 Q.49 1
Q.50 2 Q.51 2 Q.52 3 Q.53 2 Q.54 2 Q.55 1 Q.56 1
Q.57 1 Q.58 4 Q.59 2 Q.60 2 Q.61 2 Q.62 4 Q.63 1
Q.64 4 Q.65 4 Q.66 2 Q.67 3 Q.68 2 Q.69 1 Q.70 2
Q.71 2 Q.72 3 Q.73 1 Q.74 1 Q.75 4 Q.76 3 Q.77 3
Q.78 4 Q.79 4 Q.80 3 Q.81 4 Q.82 1 Q.83 1 Q.84 1
Q.85 2 Q.86 4 Q.87 1 Q.88 2
Answer Key
Q.1 1 Q.2 1 Q.3 3 Q.4 3 Q.5 4 Q.6 3 Q.7 1
Q.8 1 Q.9 4 Q.10 2 Q.11 1 Q.12 4 Q.13 3 Q.14 2
Q.15 2 Q.16 3 Q.17 3