Ento 301
Ento 301
Ento 301
What is Entomology?
It is the branch of science that deals with the study of insects, their characteristics, physiology,
behavior as well as their relationship with other organisms and environment.
What is an insect?
The term ‘insect’ is derived from the Latin word insectum, which means ‘cut into’. Insects are
the organisms with segmented bodies. They are divided in mainly three parts: head, thorax and
abdomen.
● a segmented body
● The typical insect mouth has a pair of lower jaws (maxillae) and upper jaws (mandibles)
which are designed to bite.
● They have different types of mouth parts such as, chewing, sucking and piercing.
● Most insects have one or two pairs of wings although some insects such as lice, fleas,
bristletails and silverfish are completely wingless.
Role of Insects:
Insects are both harmful and beneficial for the environment. The harmful insects are considered
pests to man as pests of cultivated crops, animals, stored products, carries of human diseases
and pests of household and industrial articles.
They are also helpful in many ways such as producers of honey, lac, silk, dyes, etc., pollinators
of crops and as natural enemies of crop pests. They also decompose dead and waste to recycle
the nutrients. Some of the insects are considered to be as friends of human being
Types of Insects:
I) Productive insects
Silk worm: - The silk worm filament secreted from the salivary gland of the larva helps us in
producing silk.
Honey bee: - Provides us with honey and many other byproducts like bees wax and royal
jelly.
Lac insects: - The secretion from the body of these scale insects is called lac. Useful in
Parasites: These are small insects which feed and live on harmful insects by completing
their life cycle in a host and kill the host insect. E.g. egg, larval and pupal parasitoids
Predators: These are large insects which capture and devour harmful insects.E.g.
Coccinellids and Preying Mantid.
Pollinators: Many cross pollinated plants depend on insects for pollination and fruit set.
E.g. Honey bees, aid in pollination of sunflower crop.
Scavengers: Insects which feed on dead and decaying matter are called scavengers.They
important for maintaining hygiene in the surroundings.E.g. Carrion beetles, Rove beetles
feed on dead animals and plants.
Harmful Insects
Each cultivated plant damage by many insect pests which feed on them reduces the
yield of the crop. E.g. cotton bollworm, Rice stem bores.
Storage pests
Insects feed on stored products and cause economic loss. E.g. Rice weevil, Pulse beetle.
Cattle are affected by pests like Horse fly, Flesh fly, and Lice. These insects suck blood of the
animals and sometimes cause and spread the diseases.
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Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
INSECT INTEGUMENT
The vertebrates have internal skeleton known as endoskeletonwhile in insects it is
located outside the body forming exoskeleton.
Insect cuticle provides space for attachment of muscles of antenna and mouthparts,
called as tentorium.
Insect body wall iscalled as Integument or Exoskeleton. It is the external covering of
the body which is ectodermal in origin. It is rigid, flexible, lighter, stronger and
variously modified in different body parts to suit different modes of life.
Integument consists of 3 layers:-
1) Cuticle (Upper)
2) Epidermis (or) hypodermis(Middle)
3) basement membrane (Inner)
1. Cuticle:-
It is outermost thick layer of integument secreted by epidermis.
It is non-cellular.
It is divided in to two regions:-Epicuticle (Upper) and Procuticle(Inner)
A. Epicuticle:
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
B. Procuticle:
Composition of cuticle:
♣ Two major components of insect cuticle are; i) Chitin,ii) Proteins.
i) Chitin:
It is a nitrogenous polysaccharide.
It consists of high molecular weight polymer of anhydro-N-acetyl glucosamine
residues joined by β-glycosidic linkages.It is water insoluble but soluble in dilute
acids, alkalies and organic solvents.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
3. Basement membrane:
It is the basal part of the body wall formed from degenerated epidermal cells and
appears as non-living amorphous (shapeless) granular layer of integument.
It is about 0.5µ in thickness and consists of fibrous protein, glycosaminoglycans
which are polymers of disaccharides.
The basement membrane forms a continuous sheet beneath the epidermis, where
muscles are attached and become continuous with sarcolemma of the muscles.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
Cuticular Cuticular
out growths invagination
Apophyses
Cuticular Aculei
processes
Spines
I. Cuticular appendages:
These are the outgrowths of the cuticle / integument connected with it by means of a
membranous joint. They arise from modified epidermal cells. These are classified in
to setae and spurs.
♣ Commonly known as hairs and arise from a cup like alveolus or pit. Setae are hollow
structures developed as extension of exocuticle and are produced by a single enlarged
hypodermal cell called ‘ trichogen’ cell. Articular membrane is usually produced by a
second hypodermal cell called ‘tormogen’ cell.
♣ Setae have role of taxonomic importance and vary with species to species. Study of
arrangement of setae is known as ‘chaetotaxy’.
(2) Spurs:
♣ Occur on the legs of many insects and differ from setae in beingmulticellular in
origin.
II. Cuticular processes:
They have no membranous articulation; They are of two types
(1) Microtrichia / fixed hairs / aculei:
♣ These are minute hair like structures found on wings of Mecoptera and certain
Diptera.
(2) Spines:
♣ Outgrowths of the cuticle which are more or less thorn like in form.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
MOULTING PROCESS
Process of periodical shedding of old cuticle and formation of new cuticle this process
is known as moulting. Itis a complex process which involves: - Apolysis, Ecdysis and
Sclerotization.
Apolysis : [Apo = formation ; Lysis = dissolution]
The dissolution of old cuticle and formation of new one is known as apolysis.
It starts with repeated mitotic division of epidermal cells resulting in increase in
number and size of epidermis, Because of this change, the epidermal cells exert
tension on cuticular surface and as a result get separated them from the cuticle.
Due to separation of epidermis from the cuticle a sub cuticular space is created and
the epidermal cells starts producing their secretion i.e. moulting fluid and cuticular
material into this space.
Ecdysis :
The stage where the insect has both newly formed epi and procuticle.
The ecdysial membrane starts splitting along the line of weakness due to muscular
activity of the inner developing insect and also because of swallowing of air & water
resulting in the distention of the gut and also due to the pumping of blood from
abdomen to thorax through muscular activity.
After the breakage of old cuticles, the new instar comes out bringing its head followed
by thorax, abdomen and appendages.
Sclerotization :
After shedding of old cuticle the new cuticle which is soft, milky white coloured
becomes dark and hard through the process known as tanning (or) sclerotization.
Three types of hormones involved in the process of moulting which are as follows-
JH: Juvenile Hormone: Produced from corpora allata of brain that helps the insects
to be in immature stage.
MH: Moulting hormone: Produced from prothoracic glands of brain that induces the
process of moulting.
Eclosion Hormone: Released from neurosecretory cells in the brain that help in the
process of ecdysis or eclosion.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
INSECT HEAD
It is the foremost part in insect body consisting of 6 segments that are fused to form a
head capsule.
The head segments can be divided in to two regions i.e. procephalon and
gnathocephalon (mouth).
Head is attached or articulated to the thorax through neck or cervix.
Head capsule is sclerotized and the head capsule excluding appendages formed by the
fusion of several sclerites is known as cranium.
Inside the head, an endoskeletal structure called the tentoriumwhich give supports to
the brain, and provides a rigid origin for muscles of the mandibles and other
mouthparts.
Head is concerned with feeding and sensory perception.
Segment Appendages
I Pre antennary segment Procephalon Pair of compound eyes&
three ocelli (Simple eyes)
The head remain vertical and is at right angle to the long axis of the body and mouth
parts are ventrally placed and projected downwards. This is also kwown as Orthopteroid type.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
CockroachGrass hopper
The head remains in the same axis to body and mouth parts are projected forward.
This is also known as Coleopteroid type.Eg: beetles
Beetles
It is same as prognathous but mouthparts are directed backward and held inbetween the fore
legs. .This is also kwown as Hemipteroid or Opisthorhynchous type.Eg: bugs
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
Suture: The sclerites separated from each other by means of thin impressed line called
suture.(Sometimes referred as a sulcus).
2.Clypeus:It is situated above the labrum, separated by fronto-clypeal suture & also
separated from gena by clypogenal suture.
3.Frons: It is unpaired, facial part of the head capsule lying between the arms of
epicranial suture.
4. Gena: It is the area extending from below the compound eyes to just above the
mandibles. It is separated from frons by frontoganal suture and from clypeus clypogenal
suture.
5. Epicraniun: It forms the upper part of the head extending from frons to the neck.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
6. Vertex: It is the top portion of epicranium which lies behind the frons or the area
between the two compound eyes.
7. Epicranial Suture: Starting from the dorsal portion of the epicranium is an inverted Y
shape suture known as epicranial suture or ecdysial line. The head capsule breaks open
along this line at the time of moulting.
8. Occiput: It is an inverted “U” shaped structure representing the area between the
epicranium and post occiput.
9. Post occiput: It is the extreme posterior part of the insect head that remains before the
neck region.
10. Occular sclerites: These are cuticular ring like structures present around each
compound eye.
11. Antennal sclerites: These form the basis for the antennae and present around the
scape.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
5. Post occipital suture: It is the only real suture in insect head. Posterior end of the
head is marked by the post occipital suture to which the sclerites are attached. As this
suture separates the head from the neck, hence named as real suture.
6. Genal suture: It is the sutures present on the lateral side of the head i.e. gena.
7. Occular suture: It is circular suture present around each compound eye.
8. Antennal suture: It is a marginal depressed ring around the antennal socket.
INSECT THORAX
It is the middle part of the body consisting of 3segmentssuch as prothorax,
mesothorax and metathorax, each possessing a pair of legs and a pair of wings on
meso and meta thoracic segment. Meso and meta thoracic segments bear a pair of
wings each together known as pterothorax (Ptera = wings).
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
Sutures – The sclerites are separated from each other by means of thin impressesd
lines called as sutures.
Sclerites forming these regions are called as tergites, sternites and pleurites,
respectively.
INSECT ABDOMEN
The abdomen in the embryo usually consists 11 segments.
The abdominal segments are sometimes designated as uromeres.
The terminal region of abdomen is called telson which bears anus.
The 1st abdominal segment gets fused to metathorax forming propodeum. (In ants,
bees and wasps).
The first eight abdominal segments carry a pair of spiracles each.
Thorax generally concerned with reproduction and metabolic activity.
Appendages of abdomen–
1. Non reproductive appendages –
a) Cerci –They are present on 11th segment in most of the insects. It is
present inmale cockroach, silverfish, grasshopperCerciusuallyact as
tactile organ or sound receptors in grasshopper. They become a part of
male genitalia in caddis fly. In earwigs, cerci are modified into
defensive organ.
b) Prolegs in insect larvae- The larvae of Lepidoptera bear five pairs of
abdominal legs called Prolegs on 3rd 4th 5th 6th and 10th segments.
These Prolegs bear spines like structures called crochets, on terminal
ends to grip the plant surfaces. In case of larvae of sawfly there are
eight pairs of Prolegs but are without crochets.
c) Abdominal gills-
It is present in aquatic insects for respiration. eg. Nymph of odonata.
d) Cornicles: Aphids have a pair of short tubes known as cornicles or
siphonculi projecting from dorsum of fifth or sixth abdominal segment.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
They permit the escape of waxy fluid which perhaps serves for
protection against predators.
2. Reproductive appendages –
It includes abdominal Segments from 1 to 7 are pregenital
segments, 8th and 9th are known as genital segments as they
form genital appendages i.e. ovipositor in females and aedeagus
or penis in males. 10thand11thsegments are known as
postgenital segments. These organs are specially concern with
mating in male and deposition of eggs in females. They are
collectively called as external genitalia or gonapophysis.
Male external genitalia-The9th sternum bears two styli and
pair of claspers which help to hold female during copulation.
The aedeagus lies between claspers.
Female external genitalia-It has a special egg laying organs
called ovipositor for egg lying on 8th and 9th segments.The
ovipositor of house fly& fruit fly is called pseudoovipositor.
INSECT ANTENNAE
Antennae are a pair of sensory preoral appendages arising from the 2nd or antennal
segment of the head possessing nerves coming from deutocerebrum of the brain.
Antennae are also called feelers.
They are well developed in adults and poorly developed in immature stages.
Antennae are absent in order protura and class Arachnida whereas 2 pairs of
antenna (antennules) are present in class Crustacea.
Aantennal socket (antennifer) is provided with an antennal suture. The base of socket
is connected to the edge of the socket by an articulatory membrane. This permits free
movement of antennae
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
2) Pedicel: It is the 2nd or middle segment of antenna that forms a joint between scape and
flagellum. It consists of the special auditory organ known as “Jhonston’s organ”.
3) Flagellum: It is the last antennal segment which consists of many segments that varies in
shape and size.
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
Term Mean
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Theory Notes on Fundamentals of Entomology
Prof. Mochi A. S. (K. K. Wagh College of Agriculture, Nashik)
Functions of Antennae
1. To feel and find the its way
2. To detect danger
3. To find food
9. It possesses hydro fuse hairs to form air funnel eg. (Water beetle)
11. Helps in mating by holding opposite sex eg. (Flea, Spring tails)
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Theory Notes on Fundamentals of Entomology
Insect Digestive System
Definition:
The digestive system of insects is a specialized mechanism responsible for the
breakdown and absorption of food.
Importance:
It enables insects to obtain essential nutrients for growth, development, and
survival.
1. General Structure
A. Foregut
1. Mouthparts: Designed for food ingestion and manipulation.
2. Esophagus: Tube-like structure connecting the mouth to the midgut.
3. Crop: Temporary storage organ for food.
B. Midgut
1. Stomach: Enzymatic digestion and nutrient absorption take place.
2. Gastric ceca: Outgrowths from the stomach, involved in enzyme production
and digestion.
C. Hindgut
1. Intestine: Absorption of water, ions, and remaining nutrients.
2. Rectum: Stores and eliminates waste.
2. Digestive Processes
A. Ingestion
1. Mastication: Food particles are broken down by the mandibles or other
mouthparts.
2. Salivation: Saliva aids in the initial digestion of carbohydrates.
B. Digestion
1. Mechanical digestion: Food is physically broken down into smaller pieces.
2. Chemical digestion: Enzymes break down complex molecules into simpler
forms.
C. Absorption
1. Nutrient absorption: Occurs in the midgut, where nutrients are transported
across the gut wall into the insect's body.
2. Water absorption: Takes place in the hindgut, where excess water is
reabsorbed.
D. Elimination
1. Waste elimination: Undigested material and metabolic waste products are
expelled through the rectum as feces.
5. Conclusion
A. The digestive system of insects is a diverse and specialized structure, enabling
them to exploit a wide range of food sources.
B. Understanding insect digestion is crucial for pest management, insect
conservation, and studying insect-plant interactions.
5. Reproductive Strategies:
Insects exhibit a wide range of reproductive strategies, including sexual
reproduction, parthenogenesis (development of offspring without fertilization),
and various forms of reproductive behavior.
Some insects display complex reproductive behaviors such as mating rituals,
territoriality, and mate guarding.
Insect reproductive strategies are often influenced by environmental factors,
including resource availability, population density, and seasonal changes.
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Home > Tutorials > Thorax > Legs
Legs
Most insects have three pairs of walking legs one pair on each thoracic segment. Each leg
contains five structural components (segments) that articulate with one another by means of hinge
joints:
1. Coxa The term pretarsus
2. Trochanter refers to the terminal
3. Femur segment of the tarsus and
4. Tibia any other structures
5. Tarsus attached to it, including:
ungues a pair of claws
arolium a lobe or
adhesive pad between the
claws
empodium a large
bristle (or lobe) between
the claws
pulvilli a pair of adhesive pads
Leg Adapations and Modifications:
Characteristic Appearance Example(s)
Ground beetles
Cursorial adapted for running and
Cockroaches
Raptorial adapted for catching and holding prey Praying mantids
Diving bugs
Natatorial adapted for swimming and
Water beetles
Fossorial adapted for digging in soil Mole crickets
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Saltatorial adapted for jumping Grasshoppers
Return to THORAX | Go to WINGS
© 2016 by John R. Meyer
Last Updated: 24 March 2016
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Insect Mouth parts
Labrum / upper lip
Mandible Hypopharynx
Upper jaw orTongue
Maxilla
Lower Jaw
-”upper lip”
Cardo
Stipes
Palp-
to taste food
Lacinia
Galea
4. Labium – Lower Lip
(2nd Maxilla)
Submentum
Post-mentum
Mentum
Prementum
Glossa
Palp
Paraglossa