Evolition 12

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Table 1.

The Difference Between Anagenesis and Cladogenesis

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1.3 Summery of Anagenesis Vs Cladogenesis
Evolution is a process in which change in the heritable characteristics of biological populations
occurs over successive generations. Anagenesis and cladogenesis are two types of such
mechanisms. The key difference between anagenesis and cladogenesis is that anagenesis is a
process that undergoes phyletic evolution within a single species of organism. Cladogenesis is a
type of branching evolution that splits the ancestral species into several new species. In
anagenesis, a single gene pool transforms into another gene pool. However, in cladogenesis, a
single gene pool splits into different gene pools.

2. Definition of Cladogram

A cladogram is the graphical representation of the hypothetical relationship (phylogenetic


relationship) between different groups of organisms. It is used in the phylogenetic analysis of
organisms to determine the evolutionary relationship between them.

The cladogram is derived from Greek words clados and gramma where ‘clados’ means branch
and ‘gramma’ means characters. It is an unscaled representation of a phylogenetic analysis where
only the topography of the diagram matters. However, it doesn’t have any time axis and is
instead a simple diagram that summarises a pattern of characters among different organisms.
Although a cladogram includes hypothetical ancestors to derive a relationship, it is the starting
point for further analysis.
3.1 Features of a cladogram

 The trees that result from the cladistic analysis are relative statements of relationship and do
not indicate ancestors or descendants. E.g., it hypothesizes that Birds and Mammals are related
but not that Mammals evolved from Birds or that Birds evolved from Mammals.
 In a cladogram, branch lengths are not proportional to the number of evolutionary changes and
thus have no phylogenetic meaning.
 The external taxa of a cladogram line up neatly in a row or a column.
 Cladograms are generated by the analysis of morphological characters of the organisms and
DNA or RNA sequencing data. Recently, however, computational phylogenetics is also used in
the combination of the existing characters for the generation of cladograms.
 Cladograms are the assumptions for the preparation of phylogenetic trees.

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 Even though cladograms are of different shapes, they all consist of lines that branch off of
other lines representing the hypothetical ancestors of different organisms.

Figure 3, Parts of a Cladogram

A cladogram is a diagram consisting of the following parts:

Figure 4 Source: BioNinja

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3.1.1 Root
 A root is the initial common ancestors of all the organisms in a cladogram.
 A root is the starting point for any given cladogram. However, the root might also indicate that
it comes from some other larger clades.
3.1.2 Nodes

 Each node is a hypothetical ancestor that gives rise to two or more daughter taxa.
 Nodes indicate the bifurcating branch point of divergence in all cladograms.
 Thus, a node exists in each point where a group of organisms divides or separate into further
different groups.
3.1.3 Clades

 Clades are groups of organisms or genes that include the most recent common ancestor of all
of its members and all of the descendants of that most recent common ancestor.
 A clade is made up of an ancestor and all its descendants.
 It includes a particular node and all of its connected branches.
3.1.4 Taxon / Outgroup

 A taxon or an outgroup is the most distantly related group of animals that isn’t necessarily a
clade. This functions as a point of reference or comparison for the rest of the cladogram.
3.1.5 Branches

 A branch in a cladogram is a line that connects all the other parts of the cladogram.
 The branch length in some cases represents the extent of divergence or the extent of the
relationship among different taxa.
3.2 Constructing Cladograms
How to make a cladogram?
Cladograms can be constructed based on either a comparison of morphological (structural)
features or molecular evidence. Historically, structural features were used to construct
cladograms, but molecular evidence is now more commonly used.
 As discussed, cladograms can be generated either based on the morphological characteristics or
molecular evidence like DNA, RNA or protein sequencing.
 Therefore, on the basis of the characters used in the cladograms, these can be made in two
different ways:
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