Dairy Industry in Sri Lanka - Part I
Dairy Industry in Sri Lanka - Part I
Dairy Industry in Sri Lanka - Part I
Dr. D.H.A.Subasinghe and Prof. H.Abeygunawardena, Faculty of Veterinary Medicine and Animal Sciences,
University of Peradeniya
30 Subasinghe & Abeygunawardena
limitations. These limitations are reported to be due to the prevailing in the South Asian region including Sri Lanka. Proof of
unavailability of land, lack of inputs, lack of establishment and this can be seen from the success of India and Pakistan who have
management experience, low awareness of improved forage taken advantage of this and used primarily the buffalo as a dairy
technology and poor animal production outcome. However, animal in their countries. Approximately 13% of the world's milk
among middle level holdings and large farms, there is a trend for production comes from the buffalo and about 90% of it is
improved pasture and fodder utilization (Premaratne & Premalal, produced in South Asia. The proportion of buffalo milk produced
2008). in India, Pakistan and Nepal are 57%, 77% and 70% respectively.
Other animal nutritionists also have pointed out that poor In Sri Lanka, the lack of good quality forages and non availability
nutrition of the dairy stock in this country has been regarded for of it round the year is a severe handicap faced by dairy farmers
decades as one of the significant limitations for improvement of particularly in the Dry and Dry Intermediate zones in the country.
national milk production (Serasinghe, 2008). Ruminant feeding The buffalo has the unique capacity to utilize poor quality coarse
systems in the country are based mainly on the agro climatic forages effectively in comparison to dairy cattle (Sebastian et al,
zones and depend mostly on natural forages available in the 1970), with 5% higher digestibility of crude fiber and at 4-5%
given region. Traditional cattle and buffalo management in Sri higher efficiency in utilization of metabolic energy for milk
Lanka has used these resources for centuries, for the sustenance production (Mudgal, 1999). This is due to a combination of
of livestock. Generally, only forage crops and weeds grown on several factors such as higher intake of dry matter and longer
uncultivated lands are available for grazing. The natural duration of feed material retained in the digestive tract. The
grasslands include dry zone villus, coconut lands, road sides, variety of micro-flora present in the buffalo rumen is more
jungle-lands, uncultivated or off-season paddy fields. favorable for the digestion of low quality fibrous feeds. Further,
Dairy farming has been further constrained by the seasonal the buffalo has a large and stronger rumen, well developed
fluctuation in the quality and quantity of forages available, salivary glands and the ability to absorb and excrete large
depending on the distribution of rainfall in each region or zone. amounts of water. Besides, it has also the ability to utilize a wide
During the monsoon season rains, there is usually an abundant variety of plant material such as reeds and aquatic weeds
growth of grass, but as the season advances towards the dry available in water logged marshy lands (Perera et al, 2005). The
season quality and quantity of forage supply decreases and re- buffalo could therefore efficiently utilize coarse forages
growth of grass commences only with the onset of the next including crop residues available in the environment and produce
season rains. Farmers in the dry zone are adversely affected by about 5-10 liters milk/day rich in butter fat content (6-10%), and
acute shortage of green forages due to prolonged droughts higher total solids than cow milk. Higher yields could be
frequently experienced. Under such circumstances, farmers have achieved with better feeding and management. Buffalo is
to depend on conserved forages, crop residues and agro- relatively resistant to diseases except highly infectious
industrial byproducts to sustain the production level of the cows. conditions like Haemorrhagic septicaemia and Foot and Mouth
It may be pointed out that half a million ha of coconut land disease. Incidence of mastitis in the buffalo is very low in relation
available in the county is a good resource for crop-stock to dairy cows (Wanasinghe et al, 1985).
integrated dairy production. Another untapped pasture resource Crossbreeding Lanka buffalo with exotic river breeds such as
is the water logged or inundated paddy lands which are not Murrah have shown great promise in increasing the production
presently cultivated, due to soil acidity or high salinity, in some of potential. While the Lanka buffalo and Murrah buffalo under Sri
the districts (Kalutara, Galle, Matara and Hambantota). Natural Lankan conditions have produced 355 liters/lactation and 1,317
grasses and weeds growing in these lands are available for liters/lactation respectively, the crossbreds or F1 animals (50%
grazing throughout the year and is particularly suitable for Murrah) have given 966 L/lactation, while back crosses (75%
buffalo production. Murrah) have given 1,215 L/lactation. Progressive increase in
milk yields have been reported with increase in level of exotic
2.3 Buffalo as a Dairy animal in Sri Lanka blood (Jalatge, 2002). The other advantage of the buffalo is that
River buffalo has been undisputedly accepted as a good dairy the Lanka buffalo can be graded up to 100 percent exotic level
animal in Asia and in some of the European countries such as with no adverse effect of environment, even under dry zone
Italy, Belgium and Bulgaria in more recent times. This is because conditions, unlike temperate crosses of dairy cows under similar
the buffalo has several advantages over dairy cattle for milk situations. The only disadvantage in the buffalo is the late
production under varying environmental situations. This applies maturity and rather sluggish breeding habit that she is known to
well to the environmental, climatic and management conditions possess. However, with improved feeding and management, they
10 6
25
29
Cross bred
Local Cross bred
Pure bred
Local
Pure bred
61 69
Cattle Buffalo
Figure 1. Proportions of Upgraded and Indigenous cattle and buffaloes in Sri Lanka
Source: Ibrahim et al, 1999
Dairy industry in Sri Lanka 31
Table 1 Cattle and Buffalo population: Changing trend in the last decade (1998 - 2007)
Type of livestock 1998 1999 2000 2001 2002 2003 2004 2005 2006 2007
Cattle
Milk Milking 219,300 221,700 213,600 214,600 207,110 211,600 216,050 222,300 229,230 238,610
cows No milk 283,800 286,800 280,400 281,000 271,124 277,400 284,420 288,570 295,840 295,380
Other cows 225,200 223,700 210,300 210,200 202,690 207,600 211,640 215,620 220,990 220,550
Bulls 187,500 190,300 180,000 181,100 174,733 178,800 182,290 185,720 189,110 189,650
Calves 262,600 269,000 263,300 266,400 257,091 263,100 266,500 272,810 279,480 278,800
Total cattle 1,178,400 1,191,500 1,147,600 1,153,200 1,112,948 1.138,700 1,160,900 1,185,020 1,124,650 1,222,990
Buffaloes
Milk Milking 53,300 55,400 53,300 53,400 51,656 51,600 53,060 54,620 55,930 57,580
cows No milk 46,500 48,400 46,500 46,700 45,139 44,800 48,150 49,510 50,400 50,980
Other cows 52,400 80,200 75,800 71,000 70,807 70,300 75,660 77,070 78,910 79,790
Bulls 66,700 66,800 62,100 58,800 56,321 56,000 60,200 61,980 63,000 63,760
Calves 67,700 68,800 66,900 60,500 58,165 57,800 64,530 64,670 65,840 66,810
Total buffalo 316,400 319,500 304,500 290,300 282,087 280,500 301,600 307,750 314,080 318,920
Source: Department of Census & Statistics (Revised annual estimates based on census of agriculture 2002; updated in 2005)
Department of Census & Statistics (Agriculture & Environment Division, National Statistics 2003-2007)
Table 2: Province wise distributions of Cattle and Buffalo population in the last decade (1998 - 2007)
Province 1998 1999 2000 2001 2002 2003 2004 2005 2006 2007
Cattle
Western 58,929 59,585 57,388 57,669 55,656 60,100 57,300 60,590 62,851 62,270
Central 81,866 82,776 79,727 80,116 77,319 81,300 83,200 84,220 83,269 82,540
Southern 76,890 77,744 74,880 75,246 72,619 74,900 75,100 81,000 86,312 86,260
Nothern 158,932 160,699 154,778 155,534 160,096 150,600 160,300 163,700 171,840 178,480
Eastern 172,389 174,306 167,883 168,503 162,814 166,300 175,000 175,100 182,230 186,910
North Eastern 261,113 264,016 254,288 255,530 246,610 254,600 254,900 256,070 259,314 250,650
North Central 190,922 193,046 185,932 186,840 180,318 177,700 182,300 194,839 194,839 194,050
Uva 153,859 155,569 149,836 150,569 145,313 146,500 144,100 147,700 144,403 150,460
Sabaragamuwa 23,499 23,761 22,885 22,997 22,194 26,700 28,500 28,560 29,516 31,370
National total 1,178,400 1,191,500 1,147,600 1,153,200 1,112,948 1,138,700 1,190,900 1,185.020 1,214,574 1,222,990
Buffalo
Western 26,607 26,868 25,607 24,413 23,722 23,670 23,700 24,400 25,400 26,750
Central 26,189 26,447 25,205 24,029 23,349 22,390 24,000 24,200 24,500 24,800
Southern 52,325 52,838 50,357 48,009 46,651 46,600 47,300 48,600 50,800 50,950
Nothern 6,524 6,588 6,278 5,985 5,816 5,740 8,700 8,850 8,850 9,000
Eastern 30,549 30,848 29,400 28,029 27,236 27,190 39,000 39,800 40,500 41,450
North Eastern 87,611 88,468 84,316 80,383 78,109 78,030 80,400 81,000 83,100 83,400
North Central 52,715 53,233 50,733 48,368 46,999 46,970 47,500 48,500 49,600 51,800
Uva 17,463 17,634 16,806 16,023 15,569 15,410 16,000 16,900 15,826 15,070
Sabaragamuwa 16,417 16,577 15,798 15,062 14,636 14,480 14,900 15,400 15,700 15,700
National total 316,400 319,500 304,500 290,300 282,087 280,480 301,500 307,750 314,176 318,920
Table 3 Distribution of cattle and buffaloes on the basis of Agro-ecological Zones 2007
Zone Cattle (No) Percent (%) Buffalos (No) Percent (%) Total No. Percent (%)
Hill & Mid country 154,150 12.6 27,070 8.5 181,220 11.8
Coconut triangle 301,140 24.6 104,800 32.8 405,940 26.3
Wet lowlands 75,010 6.1 34,350 10.8 109,360 7.1
Dry lowlands 692,690 56.7 152,700 47.9 845,390 54.8
National total 1,222,990 100 318,920 100 1,541,910 100
Source: Department of Census & Statistics (Agriculture & Environment Division; Livestock population by types & districts 1998-2007)
32 Subasinghe & Abeygunawardena
have also been minimized. Buffalo has already become a ?Suckling of calves is restricted to twice per day, at milking
popular dairy animal in the sub-urban dairies in Colombo, and times only.
among farmers in Kalutara and Kurunegala districts. Due to the ?Improved health care of calves, including strategic de-
high demand for buffalo milk and curd in the city centers, low worming and vaccination.
feeding cost, availability of inexpensive forage materials in the ?Body condition scoring using a 1-5 scale, as an aid to
respective locations, intensive management system is adopted appropriate feeding decisions.
by the sub-urban dairy farmers. Several recent reports have ?Hygienic milk production and value addition by
pointed out the value of using the buffalo as a dairy animal in Sri conversion to yoghurt, curd and other products.
Lanka (Ranawana, 2008; Subasinghe et al, 1998; Subasinghe et
?Recycling of animal waste and crop residues to provide
al, 2008).
mutual benefits.
Research studies conducted under a multidisciplinary
program have shown that simple low cost feeding systems can ?Maintenance of simple farm records and their use for
be developed using locally available feed resources. The decision making.
strategy used was to combine natural grasses, rice straw and
legume tree fodder with urea molasses multi-nutrient blocks 2.4 Milk production
(UMMB) as feed material, for use by smallholder farmers The cattle and buffalo population in Sri Lanka which
keeping buffalo under an intensive management system contributed to dairy production in the country in 1996 was
(Abeygunawardena et al, 1996). estimated at 1.64 million and 0.7 million respectively. The
The above is a successful example of a smallholder farming estimated population of milk cows and buffalos was around
system available for the predominantly mixed crop-livestock 700,000 and 235,000 respectively (DCS, 1997; Ibrahim et al,
smallholder farmers in the rain fed and irrigated rice growing 1999). Since dairy cattle and buffalo production was based on
regions of Sri Lanka, particularly in Mahaweli System H and C natural grazing, approximately 70% of cattle and 75% of the
and Kurunegala district. In this system buffalo plays a vital buffaloes are found in the Dry and Intermediate zones. Although
complex role in providing milk, draught power and manure for the livestock population data indicate that there has been a decline
the crops. Since the traditional extensive management is under in the population, overall milk production has shown an increase
threat due to intensified land use for crops and settlements, in from 275,000 MT to 330,000 MT from 1987 to 1997. This growth
this system a Smallholder Intensively Managed Buffalo Unit in milk production despite a decrease in the population is attributed
(SIMBU) was developed incorporating improved technologies to an increase in the population of crossbred animals (about 40%)
and integration of crops. This model was subsequently having higher production potential (Ibrahim et al, 1999). More
expanded to groups of other farmers and refined, based on recent data on the cattle and buffalo population estimates (1998 -
location specific needs (Abeygunawardena et al, 1996; Perera, 2007) in Sri Lanka does not indicate any significant growth or
et al, 2005; Subasinghe et al, 2008). The main features of the decline during the said period (DCS, 2008). The cattle and buffalo
SIMBU model were the following: population in 2007 was 1.2 million and 0.32 million respectively
(Table 1)
? Each unit comprised of 3-4 buffaloes and their calves
The domestic milk supply in the country during the 1970s
either purebred (Murrah, Surti, Nili-Ravi) or crosses of
provided nearly 80% of the country's consumption needs.
Indian breeds with milk yields of 4-6 liters/cow/ day.
However, with the opening of the Sri Lankan economy in 1977 and
? Low cost housing with hygienic floor, feed and water the rapid expansion of the human population in the last three
troughs and storage space in the loft for crop residues. decades, milk consumption in the country increased considerably
? Feeding based on grass and leguminous tree fodder, the (Ranaweera, 2007). Increase in per capita consumption has also
latter grown as alleys and as perimeter fence. increased during the same period, pushing up the consumption
? Utilization of rice straw supplemented with urea demand, for example the consumption increased from
molasses multi-nutrient feed blocks (UMMB). 13kg/caput/year to 36kg/caput/year from 1981 to 1997 (Ibrahim et
? Cooling by sprinkling water instead of wallowing, for al, 1999).
heat dissipation. The domestic milk supply in 2006 contributed only 17% of the
? Improved oestrus detection, timing of AI service and total requirement, while the balance was met through imports. Sri
timely pregnancy diagnosis. Lanka has imported around 68,000MT of dairy products in 2006
5 00
700 45 0 Production
Production
600 Collection 400 Imports
500 Imports 350
300
1000MT
400 Consumption
1000MT
250
300 200
200 150
100 100
50
0
1981 1987 1992 1997 1998 2003 2007 0
87 91 95 99 03 07
YEARS YEARS
Figure. 2 Trends in domestic milk production, Collection, Imports Figure. 3 Domestic milk production and imports, 1987-2007
and Consumption (1000 MT), 1981-2007 Source: Adapted from Ibrahim et al, 1999, M/L.D. and DAPH Data
Source: Adapted from Ibrahim, 1999 and Ranawana, 2008 Bank, 2007
Dairy industry in Sri Lanka 33
Table 4 Cattle and buffalo production systems: topography, climate and animal husbandry
Hill & Mid country > 450 > 2000 10 - 32 Pure exotic & crosses; Zero grazing small herds;
Some Zebu crosses some tethering
Coconut triangle < 450 1500-2500 21-28 Crosses of exotic breeds Limited grazing; tethered
under coconut palms
Wet lowlands <450 1875-2500 24-35 Same as above Limited grazing; Medium
size herds; Free grazing
large, nomadic herds
Dry lowlands <450 1000-1750 21-38 Indigenous cattle, Zebu Sedentary small herds
cattle & their crosses in irrigated settlements
Table 5 Annual Milk Production and Collection by the Formal market (1998 2007)
Year Milk Production - Liters ('1000) Milk Collection by Formal Market - Liters ('1000) Milk collected (%)
Table 6 Annual Production, Imports and Availability of Milk and Milk Products, 1998 - 2006
Cow Buffalo Total Availab/ Local Imports Availab/ Local Imports Availab/ Local Imports Availab/
caput Prodn caput Prodn caput Prodn caput
(1000MT) (1000MT) (1000MT) Kg/yr (1000MT) (1000MT) Kg/yr (1000MT) (1000MT) Kg/yr (1000MT) (1000MT) Kg/yr
1998 124.48 25.09 149.57 4.80 8.63 53.56 3.30 4.18 0.26 0.25 1.51 0.32 0.10
1999 126.42 25.50 151.92 4.99 9.52 48.31 3.16 4.24 0.17 0.24 1.54 0.12 0.09
2000 127.74 25.52 153.26 4.96 11.32 56.77 3.73 4.59 0.09 0.25 1.20 0.19 0.08
2001 129.02 25.56 154.58 4.90 15.73 52.13 3.64 4.43 0.08 0.24 1.69 0.14 0.10
2002 129.09 25.64 154.73 5.03 13.67 61.84 3.94 4.42 0.07 0.25 1.74 0.28 0.11
2003 132.22 25.56 157.78 5.47 13.41 63.74 4.10 4.73 0.05 0.24 2.17 0.36 0.13
2004 134.88 25.84 160.72 5.34 13.98 54.02 3.46 5.06 0.16 0.27 2.54 0.38 0.15
2005 136.67 26.12 162.79 5.37 15.73 52.79 3.63 5.27 0.01 0.27 2.57 0.24 0.14
2006 139.34 26.73 166.07 5.33 15.70 68.10 4.21 4.80 - - 2.79 0.16 0.15
Department of Census & Statistics (Agriculture & Environment Division, National Statistics 1985 - 2006)
amounting to approximately Rs. 15 million (Ministry of estimated annual milk production from cows and buffaloes in
Livestock Development, 2007). The total estimated Sri Lanka has grown from around 177 million liters in 1998 to
consumption in 2006 was 640,000MT. Based on the human 202 million liters in 2007. This increase reflects a total growth
population of 19.8 million, individual consumption in 2006 was of 25 million liters (i.e. 14%) or an annual growth of 1.6%
about 32 kg/caput/year. According to available statistics the during the said period (Table 5, 6 & 7).
34 Subasinghe & Abeygunawardena
Table 7 Estimated Milk Production and Imports to Sri Lanka 1998 - 2007 Mahadevan, P. (1951). Genetic improvement in milk
('1000 MT/year) yield due to selection in a herd of Sinhala cattle.
Tropical Agriculturist, 107: 161-169
1 2 3 4 5 6 7 (4+6)
Mahadevan, P. (1952). The performance of Sinhala
Year Cow Buffalo Total * Collected by Imports *** Total
Milk * Milk * Processors ** (LME) consumption cattle. Tropical Agriculturist, CVII : (1,4)233-238.
Mahadevan, P. (1953). Crossbreeding experiences
1998 147 30 177 101 370 547 with Sinhala cattle at Karagoda-Uyangoda.
1999 150 30 180 109 311 491 Tropical Agriculturist, 109: 123-127.
2000 151 30 181 93 349 530 Mahadevan, P. (1970). Genetic methods of improving
2001 153 30 183 102 330 513 cattle production in Ceylon. In: R. R. Appadurai
2002 153 30 183 92 411 595 (ed) The development of the cattle industry in
2003 157 30 187 88 394 581
Ceylon; p.28-36.
2004 160 31 190 99 349 540
2005 162 31 193 100 356 549 Ministry of Livestock and Rural Industries (1995).
2006 165 32 197 109 443 640 Policy and Program; Colombo
2007 170 32 202 114 410 612 Mudgal, V.D. (1999). Milking Buffalo. In:
Smallholder Dairying in the Tropics (eds) L. Felvy,
Source: Ranawana, 2008,* Dept. of Census and Statistics; ** Ministry of Livestock and C. Chantalakhana, ILIRI, Nairobe, Kenya, pp.
Development; DAPH: *** Calculated as LME from Customs returns 101-114.
Perera, B.M.A.O., Abeygunawardena, H., Vale, W.G.
and C. Chantalakhana. (2005). Buffalo in Livestock
Changes in milk production, collection, consumption and imports from and Wealth Creation: Improving the Husbandry of
1981 to 2007 are shown in Fig 2. Domestic production has failed to expand Animals Kept by Poor People in Developing
the market share to meet the country's demand. Hence, the level of imports Countries, (eds.) Owen, E., Katalya, A., Jayasuriya,
has increased (more than double) in the last several years, showing an N. and Smith, T. Livestock Production Program,
increase in the dependency on imports (Fig. 3). Natural Resources International Limited, United
Kingdom, pp. 451-471.
REFERENCES Premadasa, M.A. (1990). Tropical grasslands of Sri
Lanka and India. Journal of Biogeography, 17: 395-
Abeygunawardena, H., Abeywansa, W.D., Ratnayake, D. and Jayatilake, 400
M.W.A.P. (1994). Zebu cattle farming in Sri Lanka: Production Systems Premaratne, S., Premalal, G.G.C. and Jayawardena,
and Production Characteristics. In: Proceedings of the final research V.P. (2003). Sustainable management of grassland
coordinating meeting of an FAO/IAEA Coordinated Research Program resources for ruminant livestock production in Sri
on “Strengthening Research on Animal Production and Disease Lanka. Tropical Agricultural Research and
Diagnosis in Asia through the application of Immunoassay techniques”; Extension, 6: 60-65.
IAEA-TECDOC-736, IAEA, Vienna, Austria, pp 37-51. Premaratne, S. and Premalal, G.G.C. (2008). Country
Abeygunawardena., H, Subasinghe, D.H.A., Perera, A.N.F., Ranawana, Pasture/Forage Resource Profile, Sri Lanka.
S.S.E., Jayatilake, M.W.A.P. and Perera, B.M.A.O. (1996). Transfer of Available at: http://www.srilanka; agpc; fao;
Technology in Smallholder Intensive Buffalo Farming: Results from a country profiles and at: http://www.fao.org/ag/agp/
pilot study in Mahaweli System H. In: The Role of the Buffalo in Rural agpc/doc/counprof/srilanka /srilanka.htm
Development in Asia (eds.) B.M.A.O. Perera, J.A.de S.Siriwardena, Ranawana, Sivali. (2008). Dairy Industry in Sri
N.U.Horadagoda and M.N.M.Ibrahim, pp 67-94. NARESA Press, Lanka: Problems and Prospects. Economic Review,
Colombo. 34, 2-5, Peoples Bank, Sri Lanka
Abeygunawardena, H., Ratnayake, D. and Jayatillake, W.M.A.P. (1997). Ranaweera, N.F.C. (2007). Improved market Access
Characteristics of cattle farming in Sri Lanka. Journal of National and Smallholder Dairy Farmer Participation for
Science Council of Sri Lanka, 25, 25-28. Sustainable Dairy Development Lessons Learned,
Buvanendran, V. (1969). Comparative performance of F1 and F2 generation Sri Lanka. Available at: http://www.fao.org/
cows in crossbreeding experiments at Karagoda -Uyangoda; Quoted by smallholder Dairy FAO milk report .pdf
Mahadevan, P. (1970) Genetic methods of improving cattle production in Sebastian, L., Mudgal., V.D. and Nair, P.G. (1970).
Ceylon. Comparative efficiency of milk production in
Buvanendran, V. (1971). Breeding for milk and meat in the mid and hill Sahiwal cattle and Murrah buffaloes. Journal of
country of Ceylon. Journal of the National Agriculture Society of Ceylon, Animal Science, 30: 253-256
6: 82-89. Subasinghe, D.H.A, Horadagoda, N.U.,
Buvanendran, V. and Mahadevan, P. (1975). Crossbreeding for milk in Sri Abeygunawardena, H. and Siriwardena, J.A. de S.
Lanka. World Animal Review (FAO), 15:7-13. eds. (1998). Water Buffalo: Improved utilization
Central Bank of Sri Lanka (CBSL) (2007). Consumer finance and socio- through new technologies; National Science
economic survey of Sri Lanka, Part II Foundation, Colombo, Sri Lanka
Department of Census and Statistics (DCS), Sri Lanka. Livestock Subasinghe, D.H.A., Perera, B.M.A.O. and
population by type and by District1998-2007.Avilableat: Pushpakumara, P.G.A. (2008). The Role of the
http:www.statistics.gov.lk/ agriculture/livestock/livestockpopulation Water Buffalo in Rural Agriculture in Sri Lanka.
subnational. html Chapter in the Book being published by the HARTI,
Ibrahim, M.N.M., Staal, S.J., Daniel, S.L.A. and Thorpe, W. (1999). Colombo (paper has been accepted for publication).
Appraisal of the Sri Lanka Dairy Sector, Vol.1, Synthesis Report; Wanasinghe, D.D. (1985). Mastitis among buffaloes
Department of Animal Science, University of Peradeniya, ILRI and in Sri Lanka; Proceedings of the 1st World Buffalo
Ministry of Livestock Development and Estate Infrastructure, Colombo, Congress, Cairo, Egypt, pp.1331-1333.
Sri Lanka Wijeratna, W.V.S. (1970). Crossbreeding Sinhala
Jalatge, E.F.A. (1982) Production characteristics of buffaloes in Sri Lanka; cattle with Jersey and Friesian in Ceylon. Animal
Sarec Report 3, Stockholm, Sweden; pp. 96-102. Production, 12: 478-488