Additional Notes Handout 2
Additional Notes Handout 2
Additional Notes Handout 2
(i) One oogonium produces one ovum and three polar bodies.
(ii) Polar bodies have small amount of cytoplasm. It helps to retain sufficient amount of
cytoplasm in the ovum which is essential for the development of early embryo. Formation of
polar bodies maintains half number of chromosomes in the ovum.
(iii) During meiosis first crossing over takes place which brings about variation.
(iv) Oogenesis occurs in various organisms. Therefore, it supports the evidence of basic
relationship of the organisms.
Structure of Ovum
The ova are developed from the primitive germ cells which are imbedded in the substance of the
ovaries. Each primitive germ cell gives rise, by repeated divisions, to a number of smaller cells
termed ogonia, from which the ova or primary ocytes are developed.
Human ova are extremely minute, measuring about 0.2 mm. in diameter, and are enclosed within
the egg follicles of the ovaries; as a rule each follicle contains a single ovum, but sometimes two
or more are present. By the enlargement and subsequent rupture of a follicle at the surface of the
ovary, an ovum is liberated and conveyed by the uterine tube to the cavity of the uterus.
Ovum refers to secondary oocyte stage of oogenesis, where the second maturation division is yet
to occur. In this condition the matured ovum is released from ovary and enters into uterus in
mammals for fertilization. In mammals including man the ovum is discharged from the Graafian
follicle (ovulation) with one polar body.
Each ovum is a rounded and non-motile structure approx. (0.15 mm in man). In appearance and
structure the ovum differs little from an ordinary cell, but distinctive names have been applied to
its several parts
(1) the cytoplasm of the ordinary animal cell frequently termed the formative yolk
(2) the nutritive yolk or deutoplasm, which consists of numerous rounded granules of fatty and
albuminoid substances imbedded in the cytoplasm. In the mammalian ovum the nutritive yolk is
extremely small in amount, and is of service in nourishing the embryo in the early stages of its
development only
Cortex - The peripheral layer of ooplasm and contains many microvilli and cortical granules.
Microvilli are tubular outrushing of plasmalemma functions to transport of substances into and
out of egg cytoplasm. The cortical granules though not present in mammalian eggs are
mucopolysaccharides covered over by Golgi membranes.
The germinal vesicle or nucleus is a large spherical body which at first occupies a nearly central
position, but becomes eccentric as the growth of the ovum proceeds. The nucleus is enclosed by
a delicate nuclear membrane, and contains in its interior a well-defined nucleolus or germinal
spot.
zona pellucida or zona striata another primary membrane covering ovum. It is a thick
membrane, which persists for some time after fertilization has occurred, and may serve for
protection during the earlier stages of segmentation. Between the two membranes is a narrow
space known as perivitelline space.
Corona radiata
Formed when several layer of epithelial cells from the follicle, adhere to the outer surface of
zona pellucida and are arranged radially, during discharge of ovum from the graafian follicle,
forming corona radiate. The cells are radially arranged around the zona, those of the innermost
layer being columnar in shape.