Root Stem Transition

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Seminar presentation on

Root- stem transition, i.e. transition from root to stem

Presented by
Pallav Dutta
Roll no -MSBOT002
• Root and stem form continuous structure called axis( i.e. root-stem axis) which lies in a
straight line. As anatomy of root and stem differs , there is naturally a region where root
and stem meet and where the parts of one organ merge into those of another. In this
region the transition from radial to conjoint type of bundles takes place. This region is
known as transition region .The region is also known as hypocotyl region. In this
region the metaxylem rotates by 180o around its longitudinal axis whereas
protoxylem does not take part in rotation. The change of position involving inversion
and twisting of xylem strands from exarch to endarch type is referred to as vascular
transition and the part of the axis where these changes occur is called transition region.
• Eames and MacDaneal (1947) described four types of root stem transition in plants.
TYPES

• TYPE A – In this ,each xylem strand of the radial vascular bundle of the root forks by radial division . The
equally forked two branches of xylem swing laterally, one to the right and one to the left by 180o in their
upward course and join phloem strands on the inner side. As a result the radial bundles with exarch xylem in the
root become collateral with endarch xylem in the stem. In this type the number of primary bundles in the
stem is equal to that of phloem strands present in the root. Example – Mirabilis, Fumaria etc.
• Type B-In this type, both strands of the xylem as well as of the phloem fork equally by radial division, the
forked branches of the xylem swing laterally by 180° in their upward course and join up with the phloem
strands; as a result the xylem strands become inverted. The phloem branches remain in the same position. Thus
the number of vascular bundles in the stem becomes double of that in the root. This type of vascular
transition is very common in species of Cucurbita (Cucurbitaceae), Phaseolus (Leguminosae) etc.
• Type C-In third type, xylem strands do not fork but run upwards to the stem swinging laterally by 180° and the
phloem strands in the meantime divide, the divided branches swing laterally and ultimately join up with the
xylem strands on the outer side. Hence the number of vascular bundles in this case remains the same as
that of the phloem strands in the root. This is found in species of Lathyrus (Leguminosae), Phoenix
(Palmae), Medicago (Leguminosae),etc.
• Type D-The fourth type is rare, found in species of Anemarrhena (Liliaceae) and in few other
monocotyledonous plants. Here half of the xylem strands fork and swing, the other half do not fork but become
inverted; divided or forked xylem strands swing laterally by 180° and join up with the unforked xylem strands
which in the meantime become inverted. The phloem strands do not fork, but simply unite in pairs with the
triple xylem strands. Here the number of vascular bundles in stem is half that of the phloem in the root.

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