Distribution and Host Plants of Leaf-Cutter Bees in Arizona

Item Type text; Book

Authors Butler, George D.

Publisher College of Agriculture, University of Arizona (Tucson, AZ)

Rights Copyright © Arizona Board of Regents. The University of Arizona.

Download date 11/05/2020 21:27:36

Link to Item http://hdl.handle.net/10150/602161

TECHNICAL BULLETIN 167 MAY 1965

Distribution and Host Plants of

LEAF - CUTTER BEES IN ARIZONA

Agricultural Experiment Station

The University of Arizona
Tucson, Arizona
 THE DISTRIBUTION AND HOST PLANTS OF

LEAF- CUTTER BEES IN ARIZONA

GEORGE D. BUTLER, JR.

Leaf -cutter bees, members of the genus Megachile, family Megachilidae, are
useful pollinators of alfalfa and other crops but may cause damage to ornamental
plants by cutting circular and oval pieces from the leaves. These pieces are used
as linings and caps for cells which are constructed in a variety of locations. Some
species nest in the soil, others nest in tunnels of rotting wood, while still others
use borings or cavities in wood or in plant stems. One group of Megachile species,
the subgenus Chelostomoides, has recently been transferred to the genus Chali-
codoma (Michener, 1962) . The species of this genus do not cut leaves but use
resin to construct their cells.
The adults of both Megachile and Chalicodoma are non -metallic, usually
robust, black bees, with the pollen- collecting scopa on the venter of the abdomen.
The front tarsi in the males are often much dilated and modified. Identifications
of all of the specimens discussed in the following paper were made by T. B. Mitchell,
whose invaluable assistance is gratefully acknowledged.
All of the 60 species of Megachile and 15 species of Chalicodoma at present
known from the State are listed below, in the taxonomic sequence followed by
Michener (1951) . The distribution in other areas of the United States was ob-
tained from Michener (1951) and from Mitchell (1962) . The Arizona records,
with the number of separate collections following the first mention of the State
in parenthesis, are based on specimens in collections at the University of Arizona,
the University of California at Berkeley, Davis, and Riverside, and the University
of Kansas. Distribution within the State is based on a division as follows: Mohave
County, northwestern; Coconino County, northcentral; Navajo and Apache Coun-
ties north of Snowflake and St. Johns, northeastern; southern Navajo and Apache
Counties, eastern Gila County, Graham and Greenlee Counties, north of the Gila
Mts., eastern; southern Graham and Greenlee Counties, the Sta. Catalina Mts. of
Pima County, and Cochise County, southeastern; Santa Cruz and Pima Counties,
southcentral; Final, Gila, Maricopa, and Yavapai Counties, central; and Yuma
County, southwestern.
Individual localities within the State have been grouped on a geographical
basis, so that nearby localities are near each other in the list. Several abbreviations
have been used in recording the localities. "Ft. Valley Expt. For. HQ" stands for
the headquarters area of the Fort Valley Experimental Forest, approximately 7
miles northwest of Flagstaff. "Hk Hwy. mi.-" indicates a mile post on the Hitch-
cock Highway on the south slope of the Santa Catalina Mts. Milepost 0, at the
bottom, is at 2800 ft. elevation. The road climbs almost steadily for 20 miles.
Elevations in even thousands and the approximate location they are reached on the
road are as follows: 4000 ft. (mi. 6) , 5000 ft. (mi. 8) , 6000 ft. (mi. 12) , 7000
ft. (mi. 16), 8000 ft. (mi. 20 -25). The end of the road about 26 miles from
the base, is in the upper part of Sabino Creek, at the mouth of Marshall Gulch, at
about 7500 ft. elevation. There is a change in vegetation from desert to Douglas
fir first, partly dependent on the elevation, but partly on the slope and other factors.
Whitaker and Niering (1964) provide an excellent summary of the vegetation of
the south slope of this range. "S.W. Res. Sta." stands for the Southwestern Research
Station of the American Museum of Natural History, near Portal. Some of the
early collectors indicated only the mountain range as the locality so this has been
abbreviated as "Mts." following the name of the mountains. Where there are no
specific localities on a mountain range, a general collecting area has sometimes
been used such as "SE" for the southeastern portion, or the altitude such as "7500'".
Seasonal distribution is expressed in terms of the number of collections
made of the stated months. These numbers in parenthesis do not refer to the
number of specimens collected but to separate collections made at different
Iocalities and on different dates. The date records are followed by a listing of the
flower records, nesting records, and references to papers on the biology of the
species.

Genus MI2GACHIL] Latreille

Subgenus Litomegachile Mitchell
Megachile (Litomegachile) brevis brevis Say
"This is one of the most widely occuring species of bees in North America,
ranging from the Atlantic to the Pacific, both in the United States and in southern
Canada, throughout the season," according to Mitchell (1962) Arizona (28)
: :

mainly northcentral and eastern.

YAVAPAI: 25 mi. S.W. Prescott, Granite Dells, Chino Val. COCONINO: Oak
Creek Cn., 15 mi. N. Sedona, 24 mi. S. Flagstaff. Flagstaff, E. Flagstaff, San Fran-
cisco Mts. NAVAJO: Carrizo, 4 mi. N. White River. APACHE: White Mts. PIMA:
Organ Pipe Cactus Nat. Mon. April (3) , May (1) , June (14) , July (6) , August
(3), and September (1) .
FLOWER RECORDS: Gaillardia, Marrubiurn, Melilotus, Phacelia, and Senecio.
Mitchell (1962) gives 85 flower records for this species.
BIOLOGY: Bohart (1958) , Donahue (1954) , Franklin (1951) , Hixon
(1955), Linsley (1946) , Michener (1953 ) , Parker (1946), Pengelly (1958) ,
Rau (1922 -1934) , Rockwood (1951), and Stevens (1949) .
Megachile (Litomegachile) brevis onobrychidis Cockrell
Known from Nebraska and Texas, west to Idaho, British Columbia, California,
and Arizona (9) widely scattered at single locations in the southwest, south-
:

central, southeastern valleys, and northeast.

YUMA: Yuma. PIMA: Tucson. COCONINO: 0ak Creek Cn. NAVAJO: Painted
Desert. APACHE: Chinle. GREENLEE: Duncan. April (2) , May (1) , June (5) ,
and July (1) .
FLOWER RECORDS: Cercidium and Prosopis. Linsley (1946) collected, this
species from alfalfa. NESTING MATERIAL: Grape leaves.
Megachile (Litomegachile) coquilletti Cockerell
Distributed from Idaho, Utah, west to British Columbia and southern Calif-
ornia; Arizona (10) widely scattered in northcentral, central, southcentral,
:

eastern, and in the southeastern valleys.

COCONINO: Grand Canyon, Grand Canyon mile 52 (Nankoweap), Oak
Creek Cn. NAVAJO: White River. PINAL: 8 mi. S. Toltec. PIMA: Tucson.
GRAHAM: Thatcher. Cochise: 5 mi. S. E. Apache. May (1) , June (5) , July (3) ,
and August (1) .
FLOWER RECORDS: Baccharis and Cercidium. Additional records from

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Mitchell (1935) include Cleome and Eriogonum, and from Linsley (1946) alfalfa.

Megachile (Litomegachile) gentilis Cresson

Known from Texas, New Mexico, Idaho, Oregon, Nevada, California, Hawaii,
and Arizona (137) southwest, central, southcentral, and the southeastern valleys.
:

YUMA: Yuma. MARICOPA: Litchfield Park, Phoenix, Mesa, Tempe, Chandler.

PINAL: Maricopa, 11 mi. S. W. Eloy, Sacaton, Superior, Red Rock, 2 mi. N.
Florence, 3 mi. W. Oracle, 2 mi. E. Oracle. PIMA: Sta. Catalina Mts.: Sabino Cn.,
Hk. Hy. mi. 3, 4, 5, Molino Basin, "Mts. "; Tanque Verde; Redington; Tucson;
10 mi. N. Tucson; Marana; Avra Val.; 20 mi. W. Tucson; 35 mi. S. W. Tucson;
Baboquivari Mts.: Brown Cn., "Mts."; Sahuarita; Continental; Sta. Rita Mts.:
Madera Cn., "Mts. ". SANTA CRUZ: Arivaca, Sycamore Cn. nr. Ruby, Amado, Har-
shaw, 6 mi. S. E. Patagonia, 10 mi. S. W. Patagonia, Patagonia, Lochiel. COCHISE:
Huachuca Mts.: Hicksville, Ramsey Cn., "N. E."; Douglas; Elfrida; Pearce; 13
mi. S. W. Willcox; 6 Ini. N. Kansas Settlement; 30 mi. E. Kansas Settlement;
Bowie; 1 mi. S. W. Portal; 5 mi. W. Portal; S. W. Res. Sta.; Douglas. GRAHAM:
Bonita, Safford, Pinaleño Mts.: 3500- 4500'; Rio Aravaipa 2500'. NAVAJO:
Carrizo Cr., White River. COCONINO: Oak Creek Cn. YAVAPAI: 5 mi. S. Jerome.
April (15) , May (19) June (35) , July (27) , August (26) , September (9) , and
October (5) .
FLOWER RECORDS: Acacia, Amorpha, Asclepias, Aster, Baccharis, Cercid-
ium, Condalia, Croton, Dalea, Echinocactus, Eriogonum, Haplopappus, Helianthus,
Heterotheca, Larrea, Lepidium, Lippia, Lotus, Marrubium, Medicago, Opuntia,
Phacelia, Prosopis, Sapindus, Senecio, Vauquelinia, Verbesìna, Wislizenia, alfalfa
and birdsfoot trefoil.
NESTING RECORDS: Cutting leaves of Amaranthus, Boerhaavia, Fraxinus,
and. grape -holly. Nests found in an ear of corn and in a hole in a peach. Bechtel
(1958) reported nests in stems of Sambucus.
PARASITES: Coelioxys novomexicana Cockerell ( Bechtel 1958) .

Megachile (Litomegachile) lippiae Cockerell ( = texana lippiae Cockerell)

Known from Montana, Nebraska, Texas, west to Utah, California, and
Arizona (110) widely distributed in northcentral, northeast, east, southeastern
:

valleys, southcentral, central and southwest.

COCONINO: Grand Canyon, N. Flagstaff, Ft. Valley Expt. For. Hq., E. Flag-
staff. NAVAJO: Joseph City, 4 mi. N. White River, Ft. Apache. APACHE: Chinle
GRAHAM: Pinaleño Mts.: 5000- 6000'; Rio Aravaipa. YAVAPAI: 6 mi. S. W.
Mingus Mt. GILA: Payson. YUMA: Yuma, Roll. MARICOPA: Phoenix. PINAL:
Superior. PIMA: Sta. Catalina Mts.: Sabino Cn., Hk. Hy. mi. 4, 5, 15, Mud Spring;
Tucson; Tucson, Pantano Wash; Continental; Sta. Rita. Mts.: Madera Cn.; Three
Points (Robles Jn.) ; Baboquivari Mts.: Brown Cn.; Quinlan Mts.: nr. Kitt
Peak; Coyote Mts. SANTA CRUZ: 10 mi. S. W. Patagonia, Patagonia, Canelo.
CoCHISE: 20 mi. E. Benson; 30 mi. E. Pearce; Dos Cabezas, Portal; 5 mi. W. Portal;
S. W. Res. Sta.; Cochise; Tombstone; Elfrida; 6 mi. E. Douglas; Douglas; 5 mi. S. E.
Apache; Huachuca Mts.: Ramsey Cn., 6 mi. W. Montezuma Pass. March (1) ,
April (12) , May (27) , June (19) , July (19) , August (20) , and September (8) .
FLOWER RECORDS: Acacia, Agave, Amorpha, Asclepias, Baccharis, Cercid-
ium, Cirsium, Curcurbita, Erigeron, Haplopappus, Helenium, Helianthus, Hoff -
manseggia, Larrea, Lotus, Marrubium, Medicago, Melilotus, Mortonia, Nama,
Nolina, Opuntia, Petalostemum, Prosopis, Sapindus, Sphaeralcea, Verbesina, Wis-
lizenia, and birdsfoot trefoil. Mitchell (1935) also listed Hyptis, white aster, white
hollyhock and privet. NESTING MATERIAL: Cutting rose leaf.
Megachile (Litomegachile) mendica Cresson
"This is a widely distributed species in the United States, occurring from

-3-
coast to coast," according to Mitchell (1962) ; Arizona (15) scattered in north-
:

west, southcentral and southeastern.

MOHAVE: Grand Canyon mi. 116.5 (Elves Chasm) . PIMA: Sta. Catalina Mts.:
Bear Wallow; Quinlan Mts.: Kitt Peak. GRAHAM: Fort Grant, Rio Aravaipa,
Pinalefio Mts. June (4) , July (2) , and August (2) .
FLOWER RECORDS: Marrubium, Melilotus, Phacelia, and Robinia. Mitchell
(1962) lists 91 plant genera. NESTING RECORD: Reared from rose cane
(Krombein 1958) . BIOLOGY: Rau (1922) .
Megachile (Litomegachile) mendica snowi Mitchell (= snowi Mitchell)
Known from Colorado, New Mexico, and Arizona (57) northcentral, eastern,
:

southeastern mountains, and southcentral. COCONINO: Sedona; 10 mi. N. Sedona;

15 mi. N. Sedona; Oak Creek Cn.: 5600', 6000'; 25 mi. S. Flagstaff; Flagstaff;
E. Flagstaff; 7 mi. E. Ash Fork. NAVAJO: 4 mi. N. Whiteriver, Pinetop. YAVAPAI:
Granite Dells. PIMA: Sta. Catalina Mts.: Bear Wallow, Hk. Hy. mi. 19, 22, 23, 26,
Mud Springs; Tucson; Sta. Rita Mts.: Madera Cn. 6000'. GRAHAM: Pinaleño Mts.:
Wet Cn., 6000', 7500'. COCHISE: Chiricahua Mts.: 6500', S. W. Res. Sta.: Portal;
5 mi. W. Portal; Elfrida; Huachuca Mts.: Ramsey Cn. SANTA CRUZ: Patagonia.
May (4) , June (13) , July (23) , August (14) , and September (3) .
FLOWER RECORDS: Asclepias, Ceanothus, Chilopsis, Cleome, Eriogonum,
Geranium, Helenium, Heliopsis, Heterotheca, Lupinus, Marrubium, Melilotus,
Robinia, Rudbeckia, Verbesina, Viguiera, and yellow composite. Mitchell (1935 )
listed Gilia and privet.
Megachile (Litomegachile) texana Cresson ( = texana cleomis Cockerell)
"This is another widely distributed species, ranging from the Atlantic to the
Pacific in the United States and southern Canada", according to Mitchell (1962) ;
Arizona (82) : distributed throughout all areas except the deserts of the central
portion of the state.
MOHAVE: 23 mi. E. Kingman, Hualapai Mt. Pk. 5300'. YAVAPAI: 4 mi. E.
Jerome, Granite Dells, Camp Wood, Prescott. COCONINO: Grand Canyon, Cam-
eron, San Francisco Mts., at base Humphreys Pk. 9500', Flagstaff, Ash Fork, Oak
Creek Cn., Sedona. NAvAJo: Carrizo Cr., Whiteriver. APACHE: White Mts.,
18 mi. W. Springerville. GRAHAM: Pinaleño Mts.: Graham Mt.; Fort Grant. GILA:
San Carlos, 38 mi. N. E. Globe, Salt R. Cn. COCHISE: Chiricahua Mts.: Cave Cr.,
5 mi. W. Portal, S. W. Res. Sta., "Mts. "; San Bernardino Ranch; Douglas; 6 mi.
E. Douglas; 5 mi. E. Lowell; Huachuca Mts.; Ramsey Cn., "Mts. "; Tombstone.
PIMA: Sta. Catalina Mts.: Sabino Cn., Hk. Hy. mi. 11, "Mts. "; Tucson; Sta. Rita
Mts.: Miller Cn., "Mts. ". YUMA: Bill Williams R. June (26) , July (25) , August
(20) , and September (5) .
FLOWER RECORDS: Asclepias, Amorpha, Berberis, Cirsium, Euphorbia,
Franseria, Haplopappus, Larrea, Lotus, Marrubium, Melilotus, Nolina, Opuntia,
Pentstemon, Phacelia, Rhamnus, Sapindus, and Solidago. Additional host records
are given by Mitchell (1962) Acerates, Afzelia, Aster, Baptisia, Blephilia, Cro-
:

talaria, Cryptantha, Dianthera, Elephantopus, Eupatorium, Galactia, Helianthus,

Helenium, Liatris, Medicago, Melilotus, Nepeta, Oxydendrum, Petalostemon
( =Kuhnistera), Phaseolus, Psorlea, Pycnanthemum, Pyrrhopappus, Rhus, Rubus,
Serinia, Silphium, Stachys, Strophostyles, Tephrosia, Trifolium, Verbena, and Vicia.
NESTING RECORD: In soil (Krombein 1958) . BIOLOGY: Franklin (1951) ,
Krombein (1953), Pengelly (1958) , and Rau (1922) as M. generosa Cresson.
Subgenus Cressoniella Mitchell
Megachile (Cressoniella) zapoteca Cresson
Known from Arizona (18) southeastern mountains. GRAHAM: Fort Grant.
:

COCHISE: Chiricahua Mts.: Cave Cr. 5500', S. W. Res. Sta., 7000-8000', Rustler

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Pk.; Huachuca Mts.: Ramsey Cn., Reef, Palmerlee. PIMA: Sta. Catalina Mts.; Sta.
Rita Mts.: Madera Cn. June (2) , July (6) , August (3) , and September (2) .
FLOWER RECORDS: Asclepias, Helenium, Helianthus, Melilotus, Monarda, and
sweet pea.

Subgenus Megachile Latreille

Megachile (Megachile) centuncularis (Linnaeus)
Mitchell (1962) gives the distribution of this species as "a holarctic species,
its range extending from the Pacific to the Atlantic in Canada and the northern
half of the United States." Arizona (1) southeastern. PIMA: Sta. Catalina
:

Mts.: Hk. Hy. mi. 5. April (1) . FLOWER RECORDS: Pendlera. Mitchell (1962 )
gives 13 other genera, including alfalfa. BIOLOGY: Medler (1959), Michel-
bacher and Hurd (1954) , and Pengelly (1958) .
Megachile (Megachile) montivaga Cresson
Distributed from Nova Scotia to North Carolina, west to British Columbia,
California, and Mexico; Arizona (37) northcentral, central, southcentral, and
:

southeastern valleys and mountains.

CocoNINO: Schultes Pass 10 mi. N. Flagstaff, San Francisco Mts., 6 mi. W.
Flagstaff, Flagstaff, 7 mi. S. Flagstaff, Navajo Mt. 6500', House Rock Val. YAVAPAI:
Weaver Mts. YUMA: Bill Williams Fork, Roll. P1NAL: Superior, 2 mi. E. Oracle,
Eloy. PIMA: Tucson; Sta. Catalina Mts.: Molino Basin, "Mts. ". SANTA CRUZ:
7.5 mi. W. Arivaca, Arivaca, 6 mi. E. Nogales, 2 mi. W. Sonoita, Canelo. COCHISE:
Huachuca Mts.; Fry; Willcox; Chiricahua Mts.: Portal, 5 mi. W. Portal, S. W. Res.
Sta., 8000', 8700', Rustler Pk.; Douglas. April (3) , May (4) , June (5) , July
(13) , August (7) , and September (3) .
FLOWER RECORDS: Aster, Canada thistle, Chilopsis, Cirsium (purple
thistle), Helenium, Marrubium, Phacelia, and Senecio. Other hosts listed by Mitchell
(1962) include: Argemone, Asclepias, Blephilia, Campanula, Centaurea, Cleome,
Coreopsis, Dianthera, Echinacea (= Brauneria), Geranium, Gutierrezia, Hedonia,
Helianthus, Lactuca, Medicago. Monarda, Oenothera, Opuntia, Parthenium, Pent -
stemon, Rosa, Rudbeckia, Scrophularia, Silphium, Verbena, Verbesina, and Veronia.
NESTING RECORD: Cutting flower petals of hollyhock. BIOLOGY: Rau
(1934) and Stevens (1949).
Megachile (Megachile) relativa Cresson
Mitchell (1962) lists the distribution as "Mackenzie to Newfoundland, south
to California, Illinois, Tennessee and Georgia "; Arizona (15) : northcentral, north-
eastern, and southeastern mountains.
COCONINO: San Francisco Mts., at base of Humphrey's Peak 9500', Flagstaff,
Oak Creek Cn. APACHE: White Mts. COCHISE: Chiricahua Mts. June (6), July
(3) , and August (6) .
FLOWER RECORDS: Amorpha, Houstonia, Iris, Mertensia, Phacelia, and
Senecio. In addition, Mitchell (1962) lists: Aster, Baptisia, Brassica, Chrysan-
themum, Epilobium, Geranium, Melilotus, Physalis, Ranunculus, Rhodore, Rosa,
Rubus, Rudbeckia, Solidago, Trifolium, and 7,izia. BIOLOGY: Medler and Koerber
(1958).

Subgenus Eutricharaea Thomson

Megachile ( Eutricharaea) concinna Smith
Mitchell (1952) reported that "although concinna was described from the
West Indies, it belongs to an Old World group and probably was introduced
from Africa during the early part of the nineteenth century. It appeared in
the United States after World War II, and is now distributed from Florida and

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Alabama to Pennsylvania and Ohio in the East, and established as far west as
California "; Arizona (57) southwestern, central, and southcentral.
:

YUMA: Martinez Lake, Yuma, Roll. MARICOPA: Glendale, Sunnyslope, Mesa,

Chandler, Gilbert, 6 mi. E. Mesa, Laveen. PINAL: Eloy. PIMA: Marana, Tucson.
SANTA CRUZ: Patagonia. COCHISE: Bowie. April (6) , May (9) , June (11) ,
July (8) , August (11) , September (6) , October (5) , and November (1) .
FLOWER RECORDS: Acacia, Aster, Baccharis, Heliotropium, Lepidium, Lippia,
Lotus, Medicago, Prosopis, Senecio, Tamarix, bushy Vernonia, berseem clover,
citrus flowers, and sweet potato. Mitchell (1962) gives the additional records
of Asclepias, Bidens, Melilotus, and Polygonum. NESTING MATERIALS: Bou-
gainvillea, Medicago, and periwinkle. BIOLOGY: Butler and Wargo (1963) .
It may be of historical interest to note that the first collection record of this
species in Arizona was made by the author on July 19, 1953 on alfalfa at
Tucson. Five collections were made in 1954 and nine in 1955.

Subgenus Delomegachile Viereck

Megachile ( Delomegachile) frigida frigida Smith (= monardarum Cockrell)
Known from Alaska to Newfoundland, south to Pennsylvania, Michigan,
Nebraska, and in the mountains to New Mexico, central California, and Arizona
(31) northcentral, eastern, and at high elevations in southeastern.
:

CocoNINO: San Francisco Mts., 8 mi. N. Flagstaff, 6 mi. W. Flagstaff,

Flagstaff, Oak Creek Cn., Sedona. APACHE: McNary, White Mts., GRAHAM:
Pinaleño Mts.: 8000', 8500', Treasure Pk. 8900', Hospital Flat 9000'. COCHISE:
Chiricahua Mts. May (2) June (14) , July (4) , August (6) , and September (3)
, .

FLOWER RECORDS: Iris, Marrubium, Medicago, Pentstemon, Phacelia,

Polemonium, and Robinia. Mitchell (1962) adds: Apocynum, Astragalus, Cam-
panula, Epilobiurn, Malvastrum. Melilotus, Monarda, Rosa, Trifolium, Verbena,
and Vicia. NESTING RECORD: In decaying poplar (Krombein 1958) .

BIOLOGY: Hobbs (1957, Hobbs et al. 1961), Pengelly (1958), and Stephen
(1955 and 1956).
Megachile (Delomegachile) gemula Cresson
Distributed from Mackenzie to Nova Scotia, south to Georgia, California and
Arizona (1) southeastern. PIMA: Sta. Catalina Mts. FLOWER RECORDS:
:

Mitchell (1962) lists 19 genera.

Megachile ( Delomegachile) gernula cressonii Dalla Torre
Known from Washington, Nevada and Arizona (1) southeastern. PIMA:
:

Sta. Catalina Mts. June (1) . The males of gemula cressonii cannot be separated
from those of gernula.
Megachile ( Delomegachile) melanophaea melanophaea Smith
Distributed from Mackenzie to Newfoundland, south to Pennsylvania, Colo-
rado, California, and Arizona (19) eastern and southeastern at high elevations.
:

COCONINO: Oak Creek Cn. APACHE: 5 mi. S. McNary, Greer, 5 mi. W. Alpine,
White Mts. GRAHAM: Pinaleño Mts.: Hospital Flat, Treasure Pk. COCHISE: Chiri-
cahua Mts.: 8000', "Mts. ". PIMA: Sta. Catalina Mts.: mile 22, 23, 24, 26, 9150',
Bear Wallow, "Mts. ". June (12) , July (4) , and August (1) .
FLOWER RECORDS: Nolina, Robinia, and Sisymbriurn. Mitchell (1962)
lists: Agastache, Apocynum, Astragalus, Azalea, Campanula, Cypripedium, Epilo-
bium, Helianthus, Medicago, Phacelia, Psoralea, Ranunculus, Raphanus. Rhodora,
Rosa, Rubus, Rudbeckia, Symphoricarpos, Taraxacum, and Vicia. BIOLOGY:
Pengelly (1958). PARASITE: Stevens (1949) reported rearing a specimen of
Coelioxys ru fitarsis Smith.

6-
Megachile (Delomegachile) melanophaea rohweri Cockerell ( =rohweri Cockerell)
Known from Colorado, Utah, and Arizona (18) northcentral, eastern, and
:

southeastern at high elevations. CocoNiNo: San Francisco Mts., Oak Creek Cn.
APACHE: 5 mi. S. McNary, White Mts. GRAHAM: Pinaleño Mts.: Treasure
Pk. 8900', Hospital Flat, 9000', Soldier Cr. 9400'. COCHISE: Chiricahua Mts. PIMA:
Sta. Catalina Mts.: Hk. Hy. mi. 22, 23, 24, 25, 26, 27. May (2), June (10),
July (2) , and August (3) .
FLOWER RECORDS: Cirsium, Erysimum, Lupinus, Nolina, Robinia, Sisym-
brium, and Trifolium. NESTING MATERIAL: Symphoricarpos leaves.

Subgenus Phaenosarns Mitchell

Megachile (Phaenosarus) augustini Cockerell (= subfortis Mitchell)
Known from Colorado, New Mexico, Utah, and Arizona (33) Eastern, :

central, southcentral and southeastern valleys and mountains. APACHE: 32 mi.

and 15 mi. W. Springerville. NAVAJO: Joseph City. CocoNINO: Sedona, Oak Creek
Cn. YAVAPAI: Kirkland Jn., 9.2 mi. W. and 3 mi. W. Prescott, Prescott. GILA:
Payson. MARICOPA: Phoenix. PIMA: Sta. Catalina Mts.: Sabino Cn.; Sta. Rita Mts.:
Bog Spr.; Baboquivari Mts.: Schaeffer Cn. and above. SANTA CRUZ: Atascosa
Mts. COCHISE: Skeleton Cn. 6 mi. S. E. Apache; Huachuca Mts.: Hicksville,
Palmerlee, 4 mi. E. Montezuma Pass; Chiricahua Mts.: S. W. Res. Sta., 2.5 mi.
N. E. Portal, 5 mi. W. Portal, Cave Cr., 4500- 6000', 7000 -8000'; Douglas. GRAHAM:
Pinaleño Mts.: 5000 -6000', 7500'. August (16) , September (15) , and October (2) .
FLOWER RECORDS: Baileya, Helianthus, Heterotheca, Verbesina, Viguiera,
and Zexmenia.
Subgenus Derotropis Mitchell
Megachile (Derotropis) xerophila Cockerell
Known from the deserts of southern California and Arizona (8) southwest
:

and southcentral. YUMA: Dateland, 10 mi. E. Aztec. MARICOPA: Sentinel, W.

Theba. PIMA: Tucson. SANTA CRUZ: 6 mi. N. Patagonia. March (1) , April (5) ,
and May (1) . FLOWER RECORDS: Baileya and Geraea. Mitchell (1936a)
lists Chaenactis and Helianthus.
Megachile (Derotropis) yumensis Mitchell
Known only from Arizona (1) central. MARICOPA: Wickenburg. April.
:

Subgenus Xeromegachile Mitchell

Megachile (Xeromegachile) bohartii Mitchell
Known from California, Mexico, and Arizona ( 12 ) : central and south -
-

central. GILA: Gila R. nr. Christmas. PIMA: Tucson; Tucson, Campbell Ave. Ext.;
Sta. Catalina Mts.: Sabino Cn., 6000'. April (5) , and May (7) FLOWER .

RECORDS: Cercidium and Prosopis.

Megachile (Xeromegachile) casadae Cockerell
Known from South Dakota, Nebraska, Oklahoma, Texas, Wyoming, Colo-
rado, New Mexico, Utah, and Arizona (6) central and eastern. YAVAPAI: 4 mi.
:

S. Jerome. COCONINO: Oak Creek Cn. NAVAJO: Carrizo. GRAHAM: Pinaleño

Mts.: Marijilda Cn. PINAL: Oracle. April (2) , May (2) , June (1), and July (1) .
FLOWER RECORDS: Opuntia; Mitchell (1936b) lists Helianthus and
Populus.
Megachile (Xeromegachile) fucata Mitchell
Known from Colorado, New Mexico, Utah, California deserts, and Arizona
(3) southwestern and southcentral. YUMA: Wellton. PIMA: 44 mi. W. Sells,
:

Tucson. April (2) . Mitchell (1936b) lists Larrea.

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Megachile (Xeromegachile) histrata Mitchell
Known from Arizona (7) central, southcentral, and southeastern valleys.
:

PINAL: 32 mi. E. Gila Bend. PIMA: Tucson; Sta. Catalina Mts.: Sabino Cn.,
Hk. Hy. mi. 0, "Mts. ". COCHISE: 24-25 mi. E. Douglas. March (1), April (5),
May (1) . FLOWER RECORDS: Baileya, Cercidium, Larrea, and Pram pis.
Megachile (Xeromegachile) instita Mitchell
Known from New Mexico and Arizona (5) central and southcentral. PINAL:
:

Ray. PIMA: Tucson; Sta. Catalina Mts.: Upper Sabino Cn.; Sta. Rita Mts.: Madera
Cn. April (2) May (3) . FLOWER RECORDS Acacia, Cercidium, Cirsium,
:

and Prosopis.
Megachile (Xeromegachile) inyoensis Mitchell
Known from California and Arizona (2) southcentral. PIMA: Tucson Campbell
:

Ave. Ext.; Sta. Catalina Mts.: Sabino Cn. May (2) . FLOWER RECORDS:
Encelia.
Megachile (Xeromegachile) legalis Cresson (y = couleena Mitchell)
Known from Washington, Oregon, Colorado, Nevada, eastern California, and
Arizona (1) northcentral. CocoNINO: Flagstaff. June. FLOWER RECORD:
:

Senecio. Mitchell (1944) lists Cirsium and Iris.

Megachile (Xeromegachile) manifesta Cresson
Known from Alberta, Montana, North Dakota to New Mexico, southern
California, and Arizona (18) : Northcentral, eastern, southcentral, and south-
eastern lowlands.
COCONINO: 4 mi. N. Flagstaff, 7 mi. S. Flagstaff, 15 mi. and 10 mi. N.
Sedona, Oak Creek Cn., Sedona. GILA: E. Verde R. 4500'. APACHE: Springer -
ville. PIMA: Sta. Rita Mts. SANTA CRUZ: W. slope Patagonia Mts. COCHISE:
Huachuca Mts.: Ramsey Çn.; Chiricahua Mts.: S. W. Res. Sta., 1 mi. S. W. Portal,
"Mts. "; Douglas; Skeleton Cn. 6 mi. S. E. Apache. June (1) , July (0) , August
(4), September (13).
FLOWER RECORDS: Aster, Grindelia, Haplopappus, Helianthus, Viguiera,
and yellow composite. Mitchell (1936b) lists Cleome and Gutierrezia.
Megachile (Xeromegachile) maurata Mitchell (probably a of M. (Derotropis)
subanograe Mitchell)
Known from Wyoming, eastern California, and Arizona (1) southwestern.
:

PIMA: Organ Pipe Cactus Nat. Mon. April. Mitchell (1944) lists Sphaeralcea.

Megachile (Xeromegachile) mojavensis Mitchell

Known from southern and eastern _California deserts and Arizona (9) central,
:

eastern, southcentral, and southeastern at the lower elevations. PINAL: Superior.

PIMA: 10 mi. N. Tucson; Tucson, Campbell Ave. Ext.; Tucson, Pima Cn.; Sta. Cata-
lina Mts.: Sabino Cn., Hk. Hy. mi. 2; Quinlan Mts.: entr. to Kitt Pk. COCHISE:
25 mi. N. E. Douglas. APACHE: 11 mi. E. McNary. April (4), May (4), June (1).
FLOWER RECORDS: Aster, Chaenactis, Encelia, and Phacelia.
Megachile (Xeromegachile) mucorosa Cockerell (9 = nebraskana Mitchell)
Known from Nebraska, Kansas, Colorado, Texas, and Arizona (1) south-
eastern. COCHISE: Pearce. July. FLOWER RECORDS: Helianthus. Mitchell
(1936b) lists Eustoma and bluebell.
Megachile (Xeromegachile) palmensis Mitchell
Known from southern California and Arizona (4) southern part of central.
:

PINAL: N. W. Stanfield, Superior, Oracle. March (1) , April (2) , May (1) .
FLOWER RECORDS: Encelia and Cirsium. Mitchell (1944) lists Sphaeralcea.

-8-
Megachile (Xeromegachile) parallela reta Mitchell
Known only from Arizona (1) southcentral. PIMA: Tucson. May. FLOWER
:

RECORDS: Helianthus; Mitchell (1937a) lists Ericameria.

Megachile (Xeromegachile) pseudonigra Mitchell
Known from Oregon, California, and Arizona (1) : southeastern. PIMA: Sta.
Catalina Mts. June.
Megachile (Xeromegachile) redlandica Mitchell
Known only from Arizona (2) southcentral. PIMA: Tucson. April (1) , May
:

(1) FLOWER RECORDS: Eschscholtzia and Prosopis. Mitchell (1936b) lists

.

Encelia.
Megachile (Xeromegachile) soledadensis Cockerell
Known from Texas, New Mexico, southern California and Arizona (21) :

southwestern, central, southcentral, and southeastern valleys. YUMA: 30 mi. N.

Yuma. MARICOPA: 6 mi. W. Wickenburg. GILA: Payson PINAL: Picacho Pass.
PIMA: Pepper Sauce Cn.; Tucson, Campbell Ave. Ext. SANTA CRUZ: Sonoita, 25
mi. E. and 19 mi. E. Sonoita. COCHISE: Dos Cabezas, Cochise, Chiricahua Mts.,
S. W. Res. Sta., Portal, 1 mi. S. W. Portal, Skeleton Cn., Douglas, 6 mi. E. Douglas.
May (1) , August (11) , September (9) .
FLOWER RECORDS: Baccharis, Baileya, Bebbia, Chrysothamnus, Encelia,
Haplopappus, Heterotheca, Hymenothrix, and Viguiera. Mitchell (1944) lists
Verbesina.
Megachile (Xeromegachile) stoddardensis Mitchell
Known from Arizona (1) southeastern. COCHISE: nr. Paradise. April.
:

Megachile (Xeromegachile) subnigra angelica Mitchell

Known from Oregon, western portions of California, and Arizona (14) :

eastern, southeastern at lower elevations, southcentral, central, and southwestern.

APACHE: Alpine, 11 mi. E. McNary. NAVAJO: Forestdale. GRAHAM: Pinaleño
Mts.: Marijilda Cn. PINAL: Superior, 5 mi. S. Ray. PIMA: Sta. Catalina Mts.:
Sabino Cn., Hk. Hy. mi. 2, Molino Basin; 10 mi. and 24 mi. N. Tucson; Tucson;
Quinlan Mts.: Kitt Peak. YUMA: Wellton. April (8), May (2) , June (3) .

FLOWER RECORDS: Aster, Encelia, Haplopappus, and yellow composite.

Mitchell (1936b) records Bebbia.
Megachile (Xeromegachile) subparallela Mitchell
Known only from Arizona (9) southcentral and southeastern in the valleys.
:

SANTA CRUZ: 10 mi. E. Ndgales, W. slope Patagonia Mts. COCHISE: 5 mi. W.

Portal, S. W. Res. Sta., Skeleton Cn. 6 mi. E. Apache. August (2) , September (7) .
FLOWER RECORDS: Eriogonum, Haplopappus, and Heterotheca. NESTING
RECORD: Nesting in the ground under Brayulinea densa using whole leaves of
Desmodium for its cells.
Megachile (Xeromegachile) wheeleri Mitchell
Known from Saskatchewan, North Dakota, Colorado to British Columbia,
and the mountains of California; Arizona (1). COCHISE: Skeleton Cn. 6 mi. S. E.
Apache. September.

Subgenus Argyropile Mitchell

Megachile (Argyropile) nebraskana Mitchell
Known from Nebraska, Kansas, Texas, Wyoming, Colorado, New Mexico,
Utah, and Arizona (1) : southeastern valley. COCHISE: San Simon. July. FLOWER
RECORDS: Mitchell (1937a) lists Helianthus, Monarda and Ratibida.

--9-
Megachile (Argyropile) parallela parallela Smith (= parallela facunda Cresson)
Known from North Dakota, Indiana, North Carolina, Florida, and British
Columbia to Mexico; Arizona (98) southwestern, central, southcentral and south-
:

eastern at the lower elevations. YUMA: Yuma, Gila Val., Roll. MARICOPA:
Marinette, Glendale, W. Phoenix, 6 mi. N. and 12 mi. N. Phoenix, Phoenix,
Tempe, Chandler. YAVAPAI: Prescott. PINAL: Casa Grande, Eloy, Picacho, 5 mi.
S. Ray. PIMA: Tucson; Sta. Catalina Mts.: Pepper Sauce Cn., Hk. Hy. mi. 5.5,
"Mts. "; 12 mi. S. Tucson; Sta. Rita Rng. Res.; Sta. Rita Mts.: mi. 0 -2 Mt.
Wrightson Trail, Bogs Springs, Madera Cn.; Baboquivari Mts. SANTA CRUZ: Ruby,
2 mi. N. Sonoita, Sonoita, 10 mi. E. Sonoita, Canelo, 4 mi. E. and 9 mi. E.
Lochiel. COCHISE: 7 mi. W. Benson; Huachuca Mts.: 4 mi. E. Montezuma Pass,
"S. E. ", mouth of Carr Cn., Hicksville, Ramsey Cn.; 7 mi. E. Tombstone; Pearce;
Elfrida; 12 mi. and 10 mi. W. Bisbee; 10 mi. W. Douglas; Douglas; 6 mi. E.
Douglas; 8 mi. N.E. Douglas; 2 mi. S. Chiricahua; Skeleton Cn. 6 mi. S. E. Apache;
5 mi. S. Apache; Apache; 5 mi. W. Portal; 1 mi. S. W. Portal; Portal; 6 mi. S. E.
San Simon; San Simon. GRAHAM: Pinaleño Mts.: 3500- 4500', 5000', "Mts ".
YAVAPAI: 15 mi. S. Sedona. COCONINO: Oak Creek Cn. April (3) , May (8) ,
June (9) , July (14) , August (27) , September (24) , October (12) , and
November (1) .
FLOWER RECORDS: Aster, Baccharis, Encelia, Haplopappus, Helianthus,
Heterotheca, Senecio, Verbesina, Viguiera, Xanthocephalum, and Zexmenia. Addi-
tional records from Mitchell (1962) include: Boltonia; Ceanothus, Cephalanthus,
Cirsium, Coreopsis, Gaillardia, Gilia, Helianthus, Heliopsil, Hypericum, Medi-
cago, Melilotus, Petalostemum, Phaseolus, Ratibida (- Lepachys), Rudbeckia,
Silphium, and Verbena. Kearney (1923) observed the bees on Gossypium (cotton) .
NESTING MATERIAL: Fischer (1951) reported these bees using leafcuttings
of Trifolium and Spiraea; nesting in the ground.
BIOLOGY: Fischer (1951) .

Megachile ( Argyropile) rossi Mitchell

Known only from Arizona (16) central, southcentral, and southeastern
:

valleys and mountains. COCONINO: 10 mi. N. Sedona, Sedona. PINAL: Superior.

PIMA: Tucson; Tucson, Cambell Ave. Ext.; Tucson, Pima Cn.; Sta. Catalina Mts.:
Sabino Cn., Hk. Hy. mi. 5, "Mts. "; Sta. Rita Mts. SANTA CRUZ: Canelo. COCHISE:
Chiricahua Mts.: S. W. Res. Sta., 5000'; 6 mi. E. Douglas. May (2) , June (1) ,
July (0) , August (2) , September (6) , October (4) , and November (1) .
FLOWER RECORDS: Aster, Bidens, Cercidium, Encelia, Verbesina, Viguiera,
and yellow composite.

Megachile (Argyropile) sabinensis Mitchell (= pararubi Mitchell)

Known from Texas and Arizona (22) southwestern, central, eastern, south -
:

central, and the southeastern valleys. YUMA: 30 mi. W. Yuma. PINAL: Superior.
PIMA: Sta. Catalina Mts.: Sabino Cn. COCHISE: 4 mi. N and 12 mi. N. Ft.
Huachuca; Turner; 10 mi. W. Bisbee, 18 mi. E., 10 mi. E., 6 mi. E. Douglas; 6 mi.
S. E. Apache, Skeleton Cn.; Chiricahua Mts.: Texas Cn. 5000 -6000', 1 mi. S. W.
Portal, 1 mi. N. E. and 14 mi. N. E. Portal. GILA: E. end Roosevelt L. NAVAJO:
Shumway. May (1) , June (1) , July (1) August (10) , and September (9) .
,

FLOWER RECORDS: Aster, Baileya, Eriogonum, Haplopappus, and Heli-

anthus.

Megachile (Argyropile) townsendiana Cockerell

Known from North Carolina, Georgia, and Florida, westward to Texas,
Colorado, and Lower California; Arizona (24) southwestern, central, south -
:

central, and southeastern at the lower elevations. YUMA: Roll, Dateland. MARI-

-10-
COPA: Sentinel, Phoenix. PINAL: Eloy, 10 mi. S. Eloy, 10 mi.W. Eloy, Superior.
GILA: Mogollon Rim 6500'. PIMA: 19 mi. S. Tucson, Continental. COCHISE:
Tombstone, Elfrida, Willcox, San Simon, 5 mi. W. Portal, 2.5 mi. N. E. Portal,
Portal, 6 mi. E. Douglas. GREENLEE: Duncan. June (2) , July (7) , August (10) ,
September (3) , October (1) , and November (1) .
FLOWER RECORDS: Baileya, Hoffmanseggia, Medicago, Prosopis, and
Verbesina. Mitchell (1962) lists Actinella, Chrysopsis, Helianthus, Melanthera.
NESTING RECORD: A bee was observed nesting in sandy ground.

Subgenus Xanthosarus Robertson

Megachile (Xanthosarus) armata Smith (incl. comata Cresson)
Known from Arizona (21) mainly northcentral but also eastern and south-
:

eastern mountains. COCONINO: Grand Canyon, San Francisco Mts., Flagstaff,

6 mi. W. Flagstaff, Ft. Valley Expt. For., Mormon Lake 7000', Oak Creek Cn.
YAVAPAI: Prescott. GILA: Mogollon Rim 6500'. APACHE: White Mts. Diamond
Ranch. PIMA: Sta. Catalina Mts.: Mud Spr., Mt. Lemmon 9150'. June (4) , July
(5), August (9), September (1).
FLOWER RECORDS: Asclepias, black -eyed susan, Lotus, Marrubium, Pha-
celia, and Senecio. Mitchell (1936a) lists Gilia.
Megachile (Xanthosarus) cochisiana Mitchell (incl. pallidiana Mitchell)
Known from New Mexico, Nevada, southern and eastern California, and
Arizona (6) extreme southwestern and extreme southeastern. YUMA: Yuma.
:

COCHISE: Douglas, Hereford. May (1) June (4) , and August (1)
, .

FLOWER RECORDS: Cercidium and Medicago. NESTING MATERIAL:

Grape leaves.
Megachile (Xanthosarus) dentitarsus Sladen
Known from Washington, Montana, North Dakota, South Dakota, Wyoming,
Nebraska, Colorado, Utah, New Mexico, and Arizona (4) southwest and central.
:

YUMA: Yuma. COCONINO: Granite Dells. June (3) , and August (1) .

FLOWER RECORDS: Medicago. Mitchell (1936a) lists: Aster, Cleome,

Grindelia, Gutierrezia, Helianthus, Melilotus, Solidago, Trifolium, and Verbena.
NESTING RECORDS: Cutting grape leaves and flying around chinaberry tree.
Megachile (Xanthosarus) perihirta Cockerell
Ranges from British Columbia to Mexico, east to Nebraska and Texas,
according to Mitchell (1962) Arizona (4) northcentral and eastern. COCONINO:
: :

Navajo Mt. 6500', San Francisco Mts. NAVAJO: Whiteriver. APACHE: White
Mts. June (3) and July (1) .

FLOWER RECORDS: Phacelia and Robinia. Mitchell (1962) gives: Aster,

Cirsium, Cleome, Epilobium, Eriogonum, Helianthus, Melilotus, and Trifolium.
BIOLOGY: Linsley (1946) .

Subgenus Pseudocentron Mitchell

Megachile (Pseudocentron) sidalceae Mitchell
Known from Texas, New Mexico, southern California, Mexico and Arizona
(268) ; the most abundant Arizona species: northwest, southwest, central, south -
central, and southeast in the valleys. MOHAVE: 23 mi. E. Kingman, Red Lake.
YUMA: Yuma, Wellton, Roll, Tacna, Mohawk, Bill Williams Fork. MARICOPA: 10
mi. E. Sentinel, Theba, 17 mi. W. Gila Bend, Phoenix, Tempe, 20 mi. E. Mesa,
Chandler. GILA: Salt River Cn. Bridge, E. end Roosevelt L., San Carlos, Winkel-
man, Globe, PINAL: 10 mi. W. Stanfield, Superstition Mts., 4 mi. E. Florence Jn.,
Superior, Florence, 20 mi. N. Oracle Jn., 3 mi. W. Oracle, 2 mi. E. Oracle, mth.
Aravaipa Cn. nr. Mammoth, 10 mi. S. Toltec, 11 mi. E. and 10 mi. S. Eloy, 11

--11-
mi. S. W. and 8 mi. S. W. Eloy, Picacho Peak, Picacho, Red Rock. PIMA: 3 mi. W.
Marana; Marana; Cortaro; Avra Valley; Tucson; 10 mi. E. Tucson, 10 mi. N.
Tucson, 20 mi. W. Tucson, 8 mi. S., 10 mi. S., 13 mi. S., and 18 mi. S. Tucson;
Sta. Catalina Mts.: Sabin Cn., Pepper Sauce Cn., Hk. Hy. mi. 0; Redington;
Tanque Verde; Saguaro Nat. Mon.; Sahuarita; Continental; Sta. Rita Rng. Res.;
Sta. Rita Mts.: Madera Cn.; Tucson Mts.; Robles Jn. (3- Points) ; Quinlan
Mts.: Kitt Peak; Arivaca; 7.5 mi. W. Arivaca; Organ Pipe Cactus Nat. Mon. SANTA
CRUZ: Amado, Sycamore Cn. nr. Ruby, Nogales, 10 mi. S. W. Patagonia, 6 mi.
S. E. Patagonia, 5 mi. W. Patagonia, Patagonia, Gardner Cn., Sonoita, 25 mi.
E. Sonoita, Canelo. COCHISE: Dry Cn.; S. E. Whetstone Mts.; 5 mi. W. Benson;
Benson; 11 mi. E. and 20 mi. E. Benson; St. David; Tombstone; Texas Cn.;
Dragoon; W. side Willcox Playa; Huachuca Mts.: "N. E. end ", Ramsey Cn.,
Pahnerlee, Petersen's Ranch, "S. E. end "; Kansas Settlement; Bonita; Willcox;
Pearce; Elfrida; 6 mi. E. Douglas; Douglas; 5 mi. S. E. Apache; S. W. Res. Sta.;
Bowie; San Simon. GRAHAM: Safford, 15 mi. S. Safford, Indian Hot Springs,
Thatcher. GREENLEE: Duncan. YAVAPAI: 8 mi. N. E. Cottonwood, S. W. Mingus
Mt., 3 mi. W. Sta. Maria R., Sta. Maria R., Congress. APACHE: White Mts.
COCONINO: San Francisco Mts. March (1) , April (10) , May (33), June (71) ,
July (89) , August (49) , September (13) , and October (2) .
FLOWER RECORDS: Acacia, Aloysia, Asclepias, Baccharis, Cephalanthus,
Cercidium, Cleome, Condalia, Croton, Encelia, Eriogonum, Ferocactus (barrel
cactus) , Franseria, Funastrum, Gutierrezia, Haplopappus, Helianthus, Ho ff man -
seggia, Hymenothrix, Kallstroemia, Larrea, Lepidium, Lotus (birdsfoot trefoil),
Medicago, Melilotus, Mimosa, Mortonia, Olneya, Opuntia ( cholla) , Pectis, Pro -
sopis, Psilostrophe, Salix, Salsola, Sapindus, Senecio, Tamarix, Verbesina, and Wis-
lizenia. Mitchell (1937a) lists Isocoma (= Haplopappus).
NESTING MATERIALS: Amaranth/a, Boerhaavia, Fraxinus, alfalfa, china-
berry, cotton, cottonwood, rose, and umbrella tree.
Subgenus Acentron Mitchell
Megachile (Acentron) albitarsis Cresson
"Michigan and Indiana to North Carolina and Florida, west through Texas
to Mexico ", according to Mitchell (1962) ; Arizona (1) central. PINAL: Flor-
:

ence. July (1) FLOWER RECORDS: Mitchell (1962) lists 25 plant genera.
.

Subgenus Sayapis Titus

Megachile (Sayapis) fidelis Cresson
Known from South Dakota, Montana, Idaho, Oregon, California, and Arizona
(37) northcentral and the southeastern mountains. COCONINO: San Francisco
:

Mts., Humphrey's Peak at base 9500', Schultes Pass 8000' 10 mi. N. Flagstaff,
4 mi. N. Flagstaff, Flagstaff, 7 mi. S. Flagstaff, 15 mi. N. Sedona, Oak Creek Cn.
6000', Sedona. PIMA: Sta. Catalina Mts.: Mt. Lemmon, "Mts." COCHISE: Chiri-
cahua Mts.: S. W. Res. Sta., 5 mi. W. Portal, 7600', Onion Saddle, 8500', Rustlers
Pk. GRAHAM: Pinaleño Mts. 9000'. June (1) , July (10) August (19) , Septem-
,

ber (7).
FLOWER RECORDS: Aster, Chrysopsis, Cirsium, Grindelia, Helenium,
Helianthus, Heliopsis, Heterotheca, Monarda, Phacelia, Senecio, and Viguiera.
Mitchell (1937b) lists: Cleome, Cosmos, Daucus, Erigeron, Eriogonum, Gutier-
rezia, Helenium, Rudbeckia, Solidago, and Verbena. NESTING SITE: Mitchell
(1937b) reported this species nesting in a hole in a small log.
Megachile (Sayapis) frugalis frugalis Cresson
"Pennsylvania and New Jersey to Florida, west to California and Mexico ",
according to Mitchell (1962) ; Arizona (8) central, southcentral, and south-
:

western. YUMA: Bill Williams Fork. GILA: 5 mi. N. Seneca (Salt River Cn.) ,

-12-
Globe. PINAL: Superior. PIMA: Tucson, Sta.. Catalina Mts., Sta. Rita Mts. YuMA:
Yuma, Bill Williams Fork. May (2) June (3) , October (1) .
FLOWER RECORDS: Helianthus. Mitchell (1937b) lists Eriogonum, Mar-
rubium, Opuntia and Tephrosia.
Megachile (Sayapis) frugalis pseudo frugalis Mitchell
Known from California and Arizona (91) southwest, southcentral, central,
:

and northcentral. YUMA: Yuma, Yuma Test Center, Palm Cn. MARICOPA: Phoenix,
Bartlett Dam. COCONINO: Grand Canyon mile 72 (Unkar Rapids) and 87.6
( Bright Angel Cr.) , Sedona. YAVAPAI: Granite Dells. GILA: 10 mi. S. Payson,
Salt River Cn., Gila R. 3 mi. S. W. Christmas. PINAL: 4 mi. E. Florence Jn.,
Florence Jn., Superior. PIMA: Sta. Catalina Mts.: Sabino Cn., Hk. Hy. mi. 0, 5,
and 6, "Mts. "; Tucson; San Xavier Mission; Baboquivari Mts.: Brown Cn. SANTA
CRUZ: Sycamore Cn. nr. Ruby, Nogales. COCHISE: Benson, Chiricahua Mts.
April (4) , May (37) , June (22) , July (16) , August (10) , and September (2) .

FLOWER RECORDS: Aster, Bebbia, Cercidium, Chilopsis, Cissus, Cleome,

Hyptis, Prosopis, Sapindus, and Verbesina. Mitchell (1937b) gives additional
records of Croton, Eriogonum, and Lotus.
Megachile (Sayapis) inimica inimica Cresson
Distributed from Florida to Texas; Arizona (10) central and southcentral
:

mountains. COCONINO: Oak Creek Cn. 6000'. YAVAPAI: nr. Jerome 5400'. PIMA:
Sta. Catalina Mts.: Pepper Sauce Cn. COCHISE: Chiricahua Mts.; 5 mi. W. Portal,
S. W. Res. Sta., 6500', Rustler Pk.; Huachuca Mts.: Carr Cn. July (2) , August (5) ,
and September (3) .
FLOWER RECORDS: Helenium, Heliopsis, Heterotheca, Melilotus, and
Sphaeralcea. Additional records given by Mitchell (1962) include: Achyranthes,
Avicennia, Bidens, Borrichia, Coccoloba, Crotalaria, Laciniaria, Ocimum, Salva,
Vitex, and Ximensia. NESTING RECORD: Mitchell (1937b) reported this
species nesting in "worm holes" in mesquite fence posts and trees with leaf -
cutting from Monisia.
Megachile (Sayapis) inimica sayi Cresson
Distributed from "Pennsylvania to Florida, west to California and Mexico ",
according to Mitchell (1962) ; Arizona (64) mainly southcentral and the
:

southeastern mountains. CocoNINO: Oak Creek Cn. 6000'. GILA: Payson, Sierra
Ancha Expt. Sta. GRAHAM: PinaIefio Mts.' 3500 -4500', 5000', 7000', 7300', Wet
Cn., "Mts." COCHISE: 5 mi. W. and 7 mi. W. Benson; Chiricahua Mts.: Portal,
5 mi. W. Portal, 5 mi. N. W. of S. W. Res. Sta., 7000- 8000', 4 mi. W. and 9.3
mi. W. Onion Saddle, Rustler Pk., Cave Creek; Huachuca Mts Ramsey Cn.,
Palmerlee, Carr Cn., Montezuma Pass, 4 mi. E. Montezuma Pass; Douglas. SANTA
CRUZ: W. slope Patagonia Mts., 10 mi. S. W. Patagonia, 9 mi. E. Lochiel, Canelo,
2.5 mi. W. Peña Blanca, 5 mi. W. Peña Blanca, Sycamore Cn. nr. Ruby. PIMA:
Sta. Catalina Mts.; Sta. Rita Mts.: Madera Cn., Bog Spr., Florida Cn., "Mts." PINAL:
Sacaton. July (8) , August (36) , and September (20) .
FLOWER RECORDS: Aster, Haplopappus, Helenium. Helianthus, Hetero-
theca, Kallstroemia, Marrubium, Lotus, Melilotus, Pentstemon, Senecio, Solidago,
Verbesina, Vicia, Viguiera, and Zexmenia. Mitchell (1962) includes: Arctium,
Bidens, Blephilia, Brauneria, Cephalanthus, Chrysopsis, Cleome, Coreopsis, Gilia,
Heliopsis, Lepachys, Liatris, Lythrum, Melilotus, Petalostemum, Pycnanthemum,
Rudbeckia, Silphium, Strophostyles, Teucrium, Verbena, and Vernonia.
Megachile (Sayapis) mellitarsis Cresson
Known from Washington, Oregon, California, Nevada, Colorado, New Mexico,
and Arizona (4) northcentral and central. COCONINO: Flagstaff, San Francisco
:

Mts. PINAL: 5 mi. S. Ray. April (1) and June (3) . FLOWER RECORDS:

-13-
Encelia and Phacelia. Mitchell (1937b) lists Aster and Eriogonum.
Megachile (Sayapis) newberryae Cockerell (incl. howardi Cockerell)
Known from New Mexico, California and Arizona (83) southwestern, cen-
:

tral, southcentral, and the southeastern valleys. YUMA: McAllister Wash nr.
Martinez Lake, Yuma, E. Mohawk, Palm Cn., Quartzite. MARICOPA: 7 mi. W. Gila
Bend, 29 mi. S. Gila Bend, 10.6 mi. E. Gila Bend, Gillespie Dam, Phoenix, Mari -
copa Mts. YAVAPAI: 12 mi. S. Jerome. PINAL: Maricopa, Apache Jn., Florence,
Superior, Ray, 27 mi. N. Oracle Jn., Picacho. GILA: 12 mi. S. Payson, Gila R.
nr. Christmas, 5 mi. N. Seneca 4000'. PIMA: Sta. Catalina Mts.: Sabino Cn.,
Hk. Hy. 3000'; 10 mi. N. Tucson; Tucson; Tucson, Campbell Ave. Ext.; Tucson,
Pima Cn.; Tanque Verde; Redington; Saguaro Nat. Mon.; 5 mi. W. Tucson;
Tucson Mts.; Organ Pipe Cactus Nat. Mon.: Alamo Cn.; Sta. Rita Rng. Res.;
Sahuarita; Continental. SANTA CRUZ: Sycamore Cn. nr. Ruby, Patagonia. COCHISE:
Benson, Douglas, Bowie. April (27) , May (42) , June (9), July (2) , and
August (3) .

FLOWER RECORDS: Acacia, Asclepias, Aster, Cercidium, Encelia, Funa-

strum, Helenium, Olneya, Opuntia, Pittosporum, Prosopis, Senecio, and Verbesina.
NESTING: Cutting leaves of- Celtis; nesting in holes in Prosopis.
Megachile (Sayapis) policaris Say
Distributed from Georgia and Florida, west to California and Mexico; Arizona
(115) northwestern, southwestern, central, eastern, southcentral, and the south-
:

eastern valleys and mountains. CocoNINO: Oak Creek Cn., Sedona. YAVAPAI:
Clemenceau, Chino Val. GILA: Mogollon Rim 6500', 5 mi. N. Seneca 4000',
Sierra Ancha, Globe. NAVAJO: W. Whiteriver. GREENLEE: Duncan. GRAHAM:
Pinaleño Mts.: Mt. Graham. CocHISE: Chiricahua Mts.: 8000', S. W. Res. Sta.,
3.5 mi. S. W. Portal, 5 mi. W. Portal; E. Tombstone; Tombstone; Dragoon Mts.:
Cochise Stronghold; 5 mi. W. Hereford; 8 mi. N. E. Douglas; 10 mi. E. Douglas;
Douglas; Huachuca Mts.: "S. E. ", 4 mi. E. Montezuma Pass, Hicksville, rd. to
Petersen's Ranch, mouth Carr Cn., Ramsey Cn.; 12 mi. N. Ft. Huachuca; 5 mi.
W. Benson; Benson; Dry Cn. S. E. Whetstone Mts. SANTA CRUZ: Canelo, Patagonia
Mts., 6 mi. S. E. Patagonia, Patagonia, Peña Blanca, 5 mi. W. Peña Blanca, Syca-
more Cn. nr. Ruby. PIMA: Palo Alto; Continental; Sta. Rita Mts.: Madera Cn.;
Tucson; 10 mi. W. Tucson; 10 mi. N. Tucson; Sta. Catalina Mts.: Sabino Cn.,
Hk. Hy. mi. 5, Molino Basin, Pepper Sauce Cn., "Mts. "; Baboquivari Mts.: Elkhorn
Ranch on S. slope of N. end; 7.5 mi. W. Arivaca; Organ Pipe Cactus Nat. Mon.
PINAL: Superior, Florence, Red Rock, Casa Grande, 3 mi. W. Oracle, 2 mi. E.
Oracle. MARICOPA: Phoenix, Tempe. YUMA: Yuma, Roll, Bill Williams Fork.
MOHAVE: 20 mi. N. E. Kingman. March (1) , April (7) , May (8) , June (18) ,

July (18) , August (41) , September (15) , October (6) , and November (2) ;
the longest flight period for any of the Arizona Megachile.
FLOWER RECORDS: Acacia, Ascleüias, Aster. Baccharis, Baileya, Chry-
sothamnus, Condalia, Croton, Encelia, Eriogonum, Haplopappus, Helianthus,
Larrea, Melilotus, Opuntia, Petalostemum, Salsola, Senecio, Tamarix, Verbesina,
and Viguiera. Additional records are given by Mitchell (1962) as: Amorpha,
Brauneria, Coreopsis, Erigeron, Oenothera, Salvia, Silphium, Verbena, and Verbesina.
Megachile (Sayapis) pugnata pomonae Cockerell
Known from Nevada, California, and Arizona (1) northcentral. COCONINO:
:

7 mi. S. Flagstaff. September. FLOWER RECORDS: Cirsium; Stevens (1949) lists

the following plants: Arctium, Dicentra, Jerusalem artichoke, purple cone -
flower, yellow Cruciferae.
Megachile (Sayapis) pugnata pugnata Say
Distributed from Quebec and the New England States to the Pacific Coast,
south to Georgia, Texas, and Arizona (9) northcentral and northeastern. Coco-
:

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NINO: Humphrey's Peak, San Francisco Mts., Schultes Pass 10 mi. N. Flagstaff,
Flagstaff, Oak Creek Cn. APACHE: White Mts. June (4) , July (1) , August (1) ,
and September (1) .
FLOWER RECORDS: Marrubium and Phacelia. Mitchell (1962) lists 23
additional genera.

Genus CHALICODOMA Lepeletier

Chalicodoma (Chelostomoides) adelphodonta ( Cockerell) (incl. 9 tucsonensis
(Mitchell) )
Known from Arizona (10) southcentral. PINAL: 3 mi. W. Oracle. PIMA:
:

Continental, Sta. Rita Mts.: Florida Cn., Madera Cn. SANTA CRUZ: Ruby, Syca-
more Cn. nr. Ruby, 5.5 mi. W. Peña Blanca, 7 mi. S. Duquesne. COCHISE: 2 rni.
N. Cochise. June (1) , August (8) and September (1)
, .

FLOWER RECORDS: Allionia and Kallstroemia.

Chalicodoma (Chelostomoides) angelarum (Cockerell)
Known from Washington, Oregon, California, Nevada, and Arizona (5) :

northcentral, southeastern. CocoNiNO: San Francisco Mts. GRAHAM: Pinaleño

Mts.: 5000 -6000'. APACHE: White Mts. COCHISE: Huachuca Mts., S. W. Res.
Sta. June (2) , July (1) , and August (2) .
FLOWER RECORDS: Franseria, Melilotus, and Phacelia. Mitchell (1937c)
lists: Aster, Cordylanthus, Encelia, Eriogonum, Lotus, Lupinus; Penstemon, Rud-
beckia, Verbena, and Vitex. PARASITES: Linsley (1944) reported Sapyga
fulvicornis Cresson (- Sapyga angustata Cresson) .
Chalicodoma (Chelostomoides) hrowni (Mitchell) (incl. 9 felipiana (Mitchell)
Known from southern California and Arizona (16) MOHAVE: 23 mi. E.
:

Kingman. YUMA: Yuma, Palm Cn. MARICOPA: Gillespie Dam. PINAL: 13 mi.
S. E. Oracle 4500'. GILA: 5 mi. N. Seneca 4000'. Salt River Cn., 18 mi. S. E.
Globe. COCONINO: Sedona, Oak Creek Cn. NAVAJO: Carrizo Cr. PINAL: 3 mi.
W. Oracle. PIMA: Sta. Rita Mts. COCHISE: Hauchuca Mts.: Ramsey Cn.; Douglas.
May (3) , June (11) , and July (2) .

FLOWER RECORDS: Asclepias, Canotia, Opuntia, Prosopis, and Sapindus.

Timberlake (1957) gives Frankenia.
Chalicodoma (Chelostomoides) chilopsidis (Cockerell) (incl. longula (Fox) and
pratti (Cockerell) )
Known from Texas, New Mexico, California, Mexico, and Arizona (125) ,

the most abundant Arizona Chalicodoma: western, central, southcentral, and the
southeastern valleys. YUMA: McAllister Wash nr. Martinez L., Yuma, 19 mi. E.
Yuma, Wellton, Bill Williams Fork. MARICOPA: 10 mi. S. Gila Bend, 6.3 mi. E.
and 10.6 mi. E. Gila Bend, Buckeye, Maricopa Mts., Glendale, Phoenix, Tempe,
Chandler. GILA: Ray, mth. Aravaipa Cr. nr. Mammoth, 10 mi. S. Payson, 12 mi.
S. Payson, Gila R. nr. Christmas. PINAL: Superior. Apache Jn., Florence, Coolidge,
27 mi. N. Oracle Jn., Picacho, 3 mi. W. Oracle, Oracle. PIMA: 3 mi. W. Marana:
Cortaro; 10 mi. N. Tucson; Tucson, Campbell Ave. Ext.; Tucson; Tucson, "A"
Mt.; Tanque Verde; Redington; Sta. Catalina Mts.: Sabino Cn., Hk. Hy. mi. 0,
2, 4, 6; Organ Pipe Nat. Mon.; Sahuarita; Continental; Sta. Rita Rng. Res.; Sta.
Rita Mts..; 15 mi. S. Robles Jn. (Three Points); Papago Ind. Res. SANTA CRUZ:
19 mi. N. W. Nogales, Nogales, Patagonia. COCHISE: Kansas Settlement. GRAHAM:
Safford. March (7) . April (28) , May (57) . June (30) , July (2) , and August (1)
.

FLOWER RECORDS: Acacia, Cercidium, Chilopsis. Cirsium, Dalea, Encelia,

Helianthus, Heliotropium, Ho ff manseggia, Larrea, Marrubium, Medicago, Mimosa,

-15-
Olneya, Prosopis, Sapindus, Senecio, Vauquelinia, and Verbesina. Mitchell (1956)
lists Melilotus. NESTING: "Plugging a nail hole with a wax ", "nesting in nail
hole in door frame ", and "flying around trunk of a dead mesquite".
Chalicodoma (Chelostomoides) davidsoni (Cockerell)
Known from California and Arizona (5) central, southcentral, and south-
:

eastern. MARICOPA: 12 mi. N. Phoenix, Chandler. PIMA: Sta. Catalina Mts.: N.

slope Mt. Lemmon; Sta. Rita Mts.: Lower Madera Cn. CocHISE: Portal. June (2) ,
July (2) , and August (1) .

FLOWER RECORDS: Senecio. Mitchell (1937c) gives Dicentra and Krom-

bein (1958) gives Pentstemon.
Chalicodoma (Chelostomoides) dischorina (Cockerell)
Known from California, Baja California, and Arizona (50) central and
:

southcentral. YUMA: Yuma, 11 mi. E. Aztec. MARICOPA: Gila Bend, E. Gila

Bend. PINAL: Superior, mth. Aravaipa Cn. nr. Mammoth, Ray, Gila R. Cn. nr.
Christmas, Coolidge, Picacho, N. Oracle Jn., Oracle. PIMA: 10 mi. N. Tucson;
Tucson, Campbell Ave. Ext.; Tanque Verde; Sta. Catalina Mts.: Sabino Cn., Hk.
Hy. mi. 0, 4, 3000'; Sta. Rita Rng. Res.; Suhuarita; Continental; 15 mi. S. Robles
Jn. (Three Points) ; W. Papago Ind. Res.; 13 mi. N. Ajo; Organ Pipe Cactus
Nat. Mon. April (28) May (19) , and June (3)
, .

FLOWER RECORDS: Acacia, Baileya, Cercidium, Coursetia, Encelia, Larrea,

Lepidium, Prosopis, and Senecio. Krombein (1958) lists Medicago and Melilotus.
Chalicodoma (Chelostomoides) exilis exilis (Cresson)
Known from Texas, New Mexico, and Arizona (8) northcentral, south-
:

central, and southeastern. COCONINO: Oak Creek Cn. PIMA: Sta. Catalina Mts.:
Sabino Cn. SANTA CRUZ: Nogales, Patagonia. COCHISE: Huachuca Mts.: Ramsey
Cn.; Chiricahua Mts.; S. W. Res. Sta., Cave Creek, Rustler Pk. July (4), August
(3) , and September (1) .
FLOWER RECORDS: Dalea and Melilotus. Mitchell (1937c) lists Baptisia,
Coreopsis, Lespedeza, Poly gala, and Tephrosia.
Chalicodoma (Chelostomoides) exilis parexilis (Mitchell)
Known from Maryland and Indiana to Florida, and west to Arizona (24) :

southcentral and the southeastern mountains. CocoNINO: Sedona, Oak Creek

Cn. NAVAJO: Whiteriver. GRAHAM: Pinalefio Mts.: 6000', 7000', 9000'. Co-
CHISE: Huachuca Mts.: Ramsey Cn.; Chiricahua Mts.; S. W. Res. Sta.; 5 mi. S.
Apache. SANTA CRUZ: 10 mi. S. Patagonia, 6 mi. S. E. Patagonia, Harshaw.
PIMA: Sta. Rita Mts.; Sta. Catalina Mts.: Molino Basin, Hk. Hy. mi. 19. June (3) ,
July (9) , August (10) , and September (2) .
FLOWER RECORDS: Baccharis, Ceanothus, Chilopsis, Condalia, Crotalaria,
Melilotus, Mentha, Monarda. and Vicia. Mitchell (1962) lists: Asclepias, Bap -
tisia, Erigeron, Hydrolea, Ilex, Oenothera, Phaseolus, Rhus, Tephrosia, Trifolium,
and Vaccinium.
Chalicodoma (Chelostomoides) lobatifrons (Cockerell)
Southern California, Mexico, Arizona (12) generally southcentral. PIMA:
:

Tucson Mts.; Tucson; Tucson, Campbell Ave. Ext.; Sta. Catalina Mts.: Upper Sabino;
Continental. MARICOPA: Bartlett Dam, Sentinel. MOHAVE: Grand Canyon mi.
179.2 (Lava Falls) May (7) , June (1) , July (3) and August (1) .
.

FLOWER RECORDS: Cercidium, Prosopis, and Wislizenia. Mitchell (1937c)

lists Eriogonum and Olneya.
Chalicodoma (Chelostomoides) manni (Mitchell)
Known from Arizona (1) . CocHISE: Huachuca Mts.: Ramsey Cn.

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Chalicodoma (Chelostomoides) occidentalis (Fox)
Known from Texas, New Mexico, California, Mexico, and Arizona (48) :
southwestern, central, southcentral, and the southeastern valleys. YUMA: Yuma.
MARICOPA: 12 mi. N. and 6 mi. N. Phoenix, Tempe, Chandler. GILA: 10 mi. S.
Payson. PINAL: Superior, Olberg. PIMA: Tucson; Sta. Catalina Mts.: Sabino Cn.,
Molino Basin; Continental; Sta. Rita Mts.: Madera Cn. SANTA CRUZ: Nogales,
Sycamore Cn. nr. Ruby. CocHISE: Benson, Tombstone, W. Dos Cabezas. GRAHAM:
Pinaleño Mts.: 5000 -6000'; Stockton Pass; Safford; Solomon; 7 mi. W. Calva
Jn. COCONINO: Sedona. April (3) , May (13) , June (14) , July (12) , August
(6) , September (3) , October (2) , and November (1) .
FLOWER RECORDS: Caleopsis, Cercidium, Chilopsis, Cleome, Helianthus,
Larrea, Lotus (birds -foot trefoil), Marrubium, Medicago, Prosopis, Ruellia,
Senecio, Verbesina, Vernonia, (bushy), and pepper tree. Mitchell (1937) adds
Phacelia and (1936) Nolina and Tamarix. NESTING SITES: "Using a hole in
a brick wall", in dead wood of a pepper tree ", and in a trap nest in wood with
3/16" diameter hole. Mitchell (1956) reported a nest in an adobe wall.
Chalicodoma (Chelostomoides) odontostoma (Cockerell) (incl. duplexa (Mitch-
ell) )
Known from California, Mexico, and Arizona (54) mainly southwestern,
:

central, and southcentral. YUMA: Yuma, 19 mi. S. Quartszite, Roll, E. Mohawk,

10 mi. E. Aztec. MARICOPA: 12 mi. W. Theba, 29 Mi. E. Gila Bend, Gillespie
Dam. PINAL: Superior, Florence, Coolidge, Picacho. GILA: 12 mi. S. Payson,
mth. Aravaipa Cn. nr. Mammoth. PIMA: Cortaro; Tucson Mts.; N. Tucson;
Tucson; Tanque Verde; Sta. Catalina Mts.: Sabino Cn., Hk. Hy. mi. 0; Sahuarita;
Continental; Sta. Rita Rng. Res.; Sta. Rita Mts.; W. side Papago Res.; Organ
Pipe Cactus Nat. Mon. SANTA CRUZ: 12 mi. S. Tubac, Sycamore Cn. nr. Ruby,
Nogales, Patagonia. CocHISE: Tombstone, Douglas. April (25) , May (19) , and
June (10).
FLOWER RECORDS: Acacia, Asclepias, Baileya, Cercidium, Helianthus,
Heliotropium, Lepidium, Medicago, Prosopis, Senecio, and Verbesina. Mitchell
(1956) lists Melilotus.
Chalicodoma (Chelostomoides) spinotulata (Mitchell)
Known from New Mexico, southern California, and Arizona (22) north -
:

central, eastern, and southcentral. COCONINO: Oak Creek Cn., Sedona. YAVAPAI:
5 mi. S. and 15 mi. S. Sedona. NAVAJo: Carrizo Cr., Ft. Apache. GILA: Salt River
Cn., 5 mi. N. Seneca 4000'. PINAL: 3 mi. W. Oracle. PIMA: 10 mi. N. Tucson;
Tucson; San Xavier Mission; 18 mi. S. Tucson; Sta. Rita Mts.: Madera Cn.; Sta.
Catalina Mts.: Sabino Cn., Upper Sabino Cn., Hk. Hy. mi. 5, 26. April (1) , May
(3) , June (9), July (4) , August (2) , September (2) , and October (1) .
FLOWER RECORDS: Allionia, Asclepias, Cercidium, cane cactus, Eriodictyon,
Geranium, Marrubium, Senecio, and Sapindus. Mitchell (1937c) adds Cordy-
lanthus, Eriogonum, Lotus, Phacelia, and (1956) adds Cryptantha and Robinia.
Chalicodoma (Chelostomoides) subexilis (Cockerell)
Known from Nebraska, Colorado, Utah, Nevada, New Mexico, California,
and Arizona (55) northcentral and the southeastern mountains. COCONINO:
:

Schultes Pass 8000' 10 mi. N. Flagstaff; San Francisco Mts., Ash Fork, 6 mi. W.
Flagstaff, Lake Mary 7 mi. S. E. Flagstaff, Oak Creek Cn., Sedona. YAVAPAI:
Prescott. YUMA: Bill Williams Fork. PIMA: Sta. Catalina Mts.: Hk. Hy. mi. 10,
19, 23, 26, Bear Wallow, Mt. Lemmon, "Mts. "; Sta. Rita Mts. COCHISE: Huachuca
Mts.: Ramsey Cn., 6500', "top "; Chiricahua Mts.: Turkey Cr. 6000', 7000', 7800',
8000', Onion Saddle, Rustlers Park, Fly's Peak 8500 -9700', "Mts. ", Cave Creek
Cn., S. W. Res. Sta. GRAHAM: Pinaleño Mts. 5500', 7000', 8500'. APACHE: White

-17-
Mts. April (1) , June (7) , July (23) August (20) , September (4) , and October
,

(1).
FLOWER RECORDS: Amorpha, Geranium, Lotus, Marrubium, Melilotus,
Monarda, Phacelia, Phaseolus, Salvia, Trifolium, Verbesina, and Vicia. Mitchell
(1937c) lists Helianthus, Opuntia, and Psoralea, and (1956) lists sweet pea and
red clover. PARASITES: Hicks (1927) records Coelioxys gilensis Cockerell and
Hurd (1958) gives Dioxys pomonae pomonae Cockerell.
Chalicodoma (Chelostomoides) subspinotulata (Mitchell)
Known from Arizona (2) southcentral. PIMA: Coyote Mts. 3500',
: Sta.
Catalina Mts. May (1) and August (1) .

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:

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