BEETLES (COLEOPTERA) RECORDED FROM GOSFORTH PARK NATURE RESERVE

RUSSELL, A (1927). Note of an occurrence of niccolite and ullmannite at the Settlingstones Mine, Fourstones, Northumberland. Mineralogical Magazine 21, 383-387. TINDLE, A G (2008). Minerals of Britain and Ireland. Terra Publishing, Harpenden. WALKER, D R G (1988). Refgersite, NiSO4.H2 O: a new British occurrence from Warwickshire. Journal of the Russell Society 2, 29-30. YOUNG, B, STYLES, M T and BERRIDGE, N G (1985). Niccolite-magnetite mineralization from Upper Teesdale, North Pennines. Mineralogical Magazine 49, 555-559. YOUNG, B and NANCARROW, P H A (1988). Millerite from the Cumbrian Coalfield. Journal of the Russell Society 2, 5-7. BEETLES (COLEOPTERA) RECORDED FROM GOSFORTH PARK NATURE RESERVE Martin Luff School of Biology, Newcastle University 7 Southcote Orchard, Totnes, Devon, TQ9 5PA [email protected] SUMMARY Coleoptera were collected from the Gosforth Park Nature Reserve in 2008, and to a lesser extent in 2009, using pitfall traps, an aerial interception trap, beating and sweeping of vegetation and by hand searching. A total of 221 species was recorded, including eighteen regionally notable, nine nationally notable and six new to Northumberland. Pitfall traps caught 2,577 Carabidae from thirty-nine species. Most were found in all trapping sites, except those typical of wetter habitats which were only caught near the lake. The results are compared with other records from the area and their significance is discussed. INTRODUCTION The Gosforth Park Nature Reserve is situated north of Newcastle upon Tyne just outside the urbanised zone and immediately to the south-east of Newcastle race course. The bulk of the Reserve lies in NGR NZ2570, with the south and eastern sections in NZ2569, NZ2669 and NZ2670 (see Fig. 1). The total area of ca eighty ha contains mixed 19th century deciduous woodland, 20th century conifer plantation, alder and willow carr, reed beds and a lake. The only other extensive list of Coleoptera from Gosforth Park is that of Jessop and Hammond (1993). They found ninety-four species using a large flight interception trap. The catch and diversity was, however, less than that from Lockhaugh Bank Wood in the lower Derwent valley, Co. Durham. A list of Coleoptera from Castle Eden Dene, Co. Durham (Luff et al., 1974; Luff and Sheppard, 1980) contained 450 species, but that included coastal cliffs and grassland as well as woodland. The Endangered (RDB1) fungus beetle Triplax scutellaris Carpentier, first recorded from Gibside by R S Bagnall in 1905, was found in Gosforth Park in 1979 (Luff and Walker, 1981). The Invertebrate Site Register (ISR) data (Ball, 1986; 1987) now on the National Biodiversity Network (NBN), (http://data.nbn.org.uk/), lists twenty-three beetle species from Gosforth Park, including T. scutellaris. There are also records of aquatic Coleoptera from Gosforth Park in the local atlas by Eyre et al. (1985). The present study aims to build on the earlier work of Jessop and Hammond (1993) towards a more complete inventory of the terrestrial beetles of the Reserve. 114 115 very wet conditions in 2008, especially in the latter part of the summer, the traps at P2 were mostly flooded out by 20 August and were abandoned by the end of the season. On 9 June, an aerial interception trap was installed at a height of about 7 m in an old oak tree near to Lake Lodge (A on Fig. 1). The trap comprised two interlocking clear Perspex sheets, 60 cm tall and 40 cm wide, above a square metal funnel leading to a collecting bottle containing saturated salt solution. This was emptied weekly until 10 September. On occasions when the traps were not being serviced, general collecting was carried out by sweeping herbaceous vegetation, beating the lower branches of trees, sieving leaf and reed litter, searching under bark, in fungal fruiting bodies and in rotting wood. Species statuses are taken from the Review of Scarce and Threatened Coleoptera by Hyman and Parsons (1992; 1994) (Nationally Notable Species) and the Red Data Book for Northumberland (NRDB) Kerslake (1998) (Regionally Notable species). Nomenclature and the taxonomic order in the list follow Duff (2008). RESULTS A total of 221 species was recorded. The breakdown of species statuses is as follows (including a few statuses unknown but ascribed by the author): Common 168; Local 44; Nationally Notable 9. Eighteen species are Regionally Notable, of which six are new to Northumberland and three new to north-east England. Thus 27/221 (12.2% of the total species) are either Regionally or Nationally Notable. The full list is appended, but this section provides further information on the ground beetles and details of all those species that are either Nationally Notable, or Regionally Notable (NR) as listed in the Red Data Book for Northumberland. Species new to the region or to the county, which need to be added to the NR list, are listed as ‘NR+’. Fig. 1. Map of Gosforth Park Reserve, showing locations of pitfall trap sites ‘P1-P6’ and interception trap, ‘A’. Carabidae (ground beetles) METHODS The Reserve was visited approximately weekly from 8 April to 2 October 2008 and once (29 May) in 2009. On 22 April 2008, sets of five pitfall traps, each comprising a plastic cup, 8.5 cm diameter and 10 cm deep, part filled with saturated salt solution, were installed at six sites, P1-P6 (see Fig. 1): P1: in oak wood near entrance P2: at edge of reed litter south of lakeside path P3: among nettles in carr north of lake P4: in conifer plantation P5: in beech woodland near golf course P6: in grassy clearing with young birch trees among conifers. The traps were emptied at 2-3 week intervals. Sites P2-P4 were all very damp; because of the 116 Of the thirty-nine species recorded all but three were trapped in pitfall traps. Table 1 gives the numbers of each species caught at each site, ranked according to total catch and omitting species of which fewer than five individuals were trapped. The four most abundant species, Pterostichus madidus, Nebria brevicollis, P. cristatus and P. nigrita each had their maximum catch in a different pitfall set, but all occurred at least singly in all sets of traps. Only P. nigrita, a known inhabitant of wet habitats, was almost restricted to the wettest sites P3 and P2. Other species that tended to favour the wetter sites, such as Loricera pilicornis, Clivina fossor and Bembidion mannerheimi were caught in relatively low numbers. Pterostichus melanarius, a species more typical of open grassland and arable fields, was found in all pitfall sites except, surprisingly, P5 adjacent to the race course. This species may be an immigrant from nearby arable fields. 117 Table 1 Numbers of Carabidae caught in pitfall traps at each site, ranked in order of total catch. Species with fewer than five individuals are omitted. The highest catch of each species is emboldened. Species Pterostichus madidus Nebria brevicollis Pterostichus cristatus Pterostichus nigrita Patrobus atrorufus Platynus assimilis Pterostichus niger Pterostichus melanarius Loricera pilicornis Pterostichus strenuus Carabus violaceus Calathus rotundicollis Notiophilus biguttatus Cychrus caraboides Clivina fossor Bembidion mannerheimi Trechus secalis Agonum fuliginosum Leistus terminatus Total 1 60 33 184 1 0 0 50 9 1 24 0 13 5 1 0 0 0 0 0 381 2 30 3 8 18 0 1 4 4 5 1 1 0 0 0 0 0 0 5 2 82 Pitfall trap Site 3 4 87 70 343 210 136 49 111 3 109 21 84 27 15 3 16 4 15 14 5 3 4 2 1 3 8 2 1 2 7 1 4 3 5 0 0 0 3 0 923 448 5 342 27 57 1 0 0 3 0 2 2 9 2 1 5 0 1 0 0 0 452 6 Total 109 698 64 680 31 465 1 135 0 130 0 112 13 88 14 47 8 45 7 42 18 34 1 20 3 19 1 10 0 8 0 8 1 6 0 5 0 5 271 2557 Notable species Details of the Nationally or Regionally Notable species are given below. All dates are in 2008 unless stated otherwise. Agonum thoreyi Dejean, 1828 (Fig. 5) NR, Local Found occasionally in P3 and in reed litter. This marsh species is widespread and common in the south but more local further north, occurring as far north as southern Scotland. Recorded from Gosforth Park in the ISR. Ophonus rufibarbis F., 1792 (Fig. 6) NR, Common One in P5, 25 July. This phytophagous species is common in southern England north to Yorkshire, but much more local further north. Hydraenidae Hydraena britteni Joy, 1907 (Fig. 7) NR, Common One in litter, 30 July. A water beetle associated with decomposing plant material in slowly running water. Two recent records in NRDB but not recorded from Gosforth Park in the national water beetle recording scheme. Ptiliidae Ptinella cavelli (Broun, 1893) NR, Naturalised Few found under bark of fallen beech, 10 September. A tiny woodland species, introduced from New Zealand and first recorded in Britain in 1975. Leiodidae Agathidium varians Beck, 1817 (Fig. 8) NR, Local One in aerial trap, 17 July. Common over most of England, becoming more local northwards. Catops coracinus Kellner, 1846 (Fig. 9) NR, Local One in pitfalls P5, 25 July. A local beetle found in decaying fungus and carrion. Silphidae Dendroxena quadrimaculata (Scopoli, 1772) (Fig. 10) Notable One larva was found in P1 on 3 July, but adults were not found despite extensive searching.This conspicuous species is an active predator of geometrid caterpillars in oak foliage. Restricted to old woodland. Very local in northern England with only one other recent Northumberland record from Hyon’s Wood, 1994 (Luff et al., 1996). There is an old record from Gibside, Co. Durham (Bold, 1872). Carabidae Leistus rufomarginatus (Duftschmid, 1812) (Fig. 2) NR, Local, naturalised One larva in P5, 20 May, one adult in P4, 25 July. Larva and adult in oak litter, 29 May 2009. A relatively recent immigrant to Britain, originally in the south-east, now found in woodlands as far north as central Scotland (Luff, 1998). Leistus spinibarbis (F., 1775) (Fig. 3) NR, Local One found under bark of fallen beech, 12 August. A woodland species near its northern limit with few Northumberland records (Luff et al., 1996). Pterostichus cristatus (Dufour, 1820) (Fig. 4) Notable Many in all pitfalls throughout the season. This species is almost restricted to Cumbria, Durham, Northumberland and southern Scotland. Possibly accidentally introduced via docks on the Tyne in the 19 th century, the distribution is still centred on the River Tyne (Luff, 1998). Staphylinidae Phloeonomus pusillus (Gravenhorst, 1806) (Fig. 11) NR, Local One in Polyporus on birch. A local species usually found under conifer bark. Aleochara ruficornis Gravenhorst, 1802 Notable. One in pitfalls P6, 20 August. A local species, possibly associated with moles’ runs and nests, which is distributed widely in the north-east (Luff et al., 1996). Leptusa norvegica Strand, 1941 Notable One under bark of fallen beech logs, 20th August. A sub-cortical species previously confused with L. fumida (Erichson) (Fig. 12). Probably the first record for north-east England. Gyrophaena strictula Erichson, 1839 Notable Several on fungal fruiting bodies on fallen logs, 12 August. Another minute but fungivorous species, associated with Oak Mazegill Daedalea quercina (L.). Recorded from Lockhaugh Bank Wood by Jessop and Hammond (1993), but not from Gosforth Park. Oligota apicata Erichson, 1837 Notable Several in Polyporus on birch logs on 9 June and 30 July. This minute fungus-living species is 118 119 probably predatory on Ciidae eggs and larvae. First recorded from the north-east by Jessop and Hammond (1993) from Lockhaugh Bank Wood, but not from Gosforth Park, so this is a new record for Northumberland. Bisnius subuliformis (Gravenhorst, 1802) (Fig. 13) NR+, Local (Formerly known as Philonthus fuscus). One in aerial trap in oak tree 24 July. This small rove beetle is found in litter in birds’ nests. Possibly predatory on fly or flea larvae. Mainly in southern England becoming rarer northwards. First record for north-east England. Endomychidae Endomychus coccineus (L., 1758) (Fig. 14) NR Local One under bark of fallen beech tree, 20 August. A brightly-coloured ‘false-ladybird’ beetle. The only other recent Northumberland record is from Morralee Wood, 1986 (National Biodiversity Network). Coccinellidae Halyzia sedecimguttata (L., 1758) (Fig. 15) NR Local The Orange Ladybird One by beating oaks, 20 May. This ladybird has recently extended its range throughout northern England. Latridiidae Dienerella elongata (Curtis, 1830) (Fig. 16) NR+ Local Several in Polyporus on birch stump, 20 August. A minute, elongated beetle found on moulds and fungi. Apparently the first published record for Northumberland, although the author also has two examples from Heddon-on-the-Wall collected in 1991. This species was previously confused with D. clathrata (Mannerheim) (Krell et al., 2005); the map given for D. elongata by these authors shows no confirmed records between Yorkshire and northern Scotland. Chrysomelidae Chrysomela aenea L., 1758 (Fig. 21) NR Local The Alder Leaf Beetle Several on alders east of the lake, 24 June. This species has been recorded from scattered Northumberland sites. Pyrrhalta viburni (Paykull, 1799) (Fig. 22) NR+ Local The Guilder Rose Beetle. Several larvae on Viburnum in warden’s garden, 9 June; adults reared from captured larvae, 15 July. Sometimes abundant on the same plants in other years (P Drummond, pers. comm.). A known pest of Viburnum in gardens in southern England, but with only two Northumberland localities (Cox, 2007). Attelabidae Attelabus nitens (Scopoli, 1763) (Fig. 23) Notable Several beaten from oaks, 29 May 2009. A leaf-rolling weevil of oaks, only known in the county from Gosforth Park (ISR). Erirhinidae Grypus equiseti (F., 1775) (Fig. 24) Notable One in pitfalls P2, 9 June. A weevil associated with Equisetum; it is widespread in the region (Luff et al., 1966) and does not merit its Notable status. Curculionidae Curculio villosus F., 1781 (Fig. 25) Notable One in oak leaf litter, 29 April, one by beating young oak by south path, 20 May. A nut weevil found in galls on Quercus. One previous recent Northumberland record from Ned’s Whin, 1995 (NRDB). DISCUSSION Tetratomidae Tetratoma fungorum F., 1790 (Fig. 17) NR Local. In Polyporus on birch logs, 20 August. A fungus beetle with few recent records from Northumberland, where it is near the northern limit of its range. Salpingidae Vincenzellus ruficollis (Panzer, 1794) (Fig. 18) NR Local. Two under bark of fallen beech, 20 August. Local in Northumberland. Cerambycidae Pachytodes cerambyciformis (Schrank, 1761) (Fig. 19) NR Local On umbelliferae in warden’s garden, 3 July. The larvae of this species breed underground in decaying stumps. There are six previous Northumberland records. Phymatodes testaceus (L., 1758) (Fig. 20) NR+ Local The Tanbark Beetle One found flying in warden’s garden, 29 July. This brown or brown/metallic species breeds in oaks. There is a single pre-1970 record from Co. Durham (Twinn and Harding, 1999), and the present record is not only a new record for Northumberland but also the furthest north that the species has been found in Britain. 120 It is clear even from this partial survey, mostly in a single, very wet year, that the Reserve has an extensive beetle fauna with several species of local note. The beetle fauna of the site found in the present work still represent a very incomplete picture of its total beetle diversity. A further species, Sinodendron cylindricum (L.) (Lucanidae) was reported to the author by the warden; there is also a photograph of Leptura quadrifasciata L. (Cerambycidae) in the members’ hut. Jessop and Hammond (1993) listed ninety-three species of beetle from Gosforth Park, but only thirty-five of these were found in the present work. Luff and Walker (1981) recorded two further species of Triplax (Erotylidae). The national water beetle recording scheme list from the Park (G N Foster, pers. comm.) includes twenty-two species, of which only seven were recorded in this work. The resulting recorded beetle fauna of the Park is outlined in Table 2; the total of more than 310 species is probably the most extensive list from any locality in Northumberland. However there are still many gaps in our knowledge of the Reserve’s beetle fauna. The list of ground-active Carabidae and Staphylinidae is probably fairly complete, due to the extensive pitfall trap sampling in the present work. But there may still be many species of plant-living beetles unrecorded, as evidenced by the fact that the single site visit made in 2009, collecting mainly by beating oak and other trees, added three extra species among the fourteen found. The widespread two-spot ladybird Adalia bipunctata (L.) (Coccinellidae) has not yet been formally recorded, and it is likely that the immigrant harlequin ladybird, Harmonia axyridis (Pallas) may be found there in the not too distant future as it has been recorded in Northumberland and southern Scotland since 2008 (see http://www.harlequin-survey.org). 121 The Gosforth Park beetle list still contains fewer species than the 450 from Castle Eden Dene (Luff et al., 1974; Luff and Sheppard, 1980) or the 447 from Gibside (Jessop and Eyre, 1992). Also in County Durham, flight interception trapping recorded 204 species from Lockhaugh Bank Wood (Jessop and Hammond, 1993) and 243 from Phragmites reed beds near Billingham (Jessop and Hammond, 1997). It is hoped that further recording by other coleopterists may add further to our knowledge of the undoubted extensive beetle fauna of Gosforth Park. Table 2 Numbers of species of beetle recorded from Gosforth Park. Source of data Number of species Total Additional 221 221 2 2 2 2 23 15 93 58 22 14 This paper Other records (see text) Luff and Walker (1981) Ball (1986; 1987) Hammond and Jessop (1993) National water beetle recording scheme EYRE, M D, BALL, S G and FOSTER, G N (1985). An atlas of the water beetles of Northumberland and Durham. Hancock Museum, Special Publication 1. HYMAN, P S (revised PARSONS, M S) (1992). A review of the scarce and threatened Coleoptera of Great Britain. Part 1. U.K. Nature Conservation 3. HYMAN, P S (revised PARSONS, M S) (1994). A review of the scarce and threatened Coleoptera of Great Britain. Part 2. U.K. Nature Conservation 12. JESSOP, L and EYRE, M D (1992). The beetles of Gibside: analysis of a lost fauna. Vasculum 77, 43-62. Cumulative 221 223 225 240 298 312 ACKNOWLEDGMENTS I am indebted to the Natural History Society of Northumbria and to Natural England for this opportunity to collect in the Reserve and SSSI. In particular thanks are due to the warden, Paul Drummond, for his help, enthusiasm and for tree climbing to install the interception trap. I am also grateful to Mr Colin Johnson and Dr Colin Welch for identification of Ptiliidae + some Cryptophagidae and some Staphylinidae respectively. Data from the national water beetle recording scheme were kindly made available by Professor Garth Foster. For permission to use colour pictures of selected notable species I must thank Dr Ortwin Bleich, Germany (Figs. 2-4) and Professor Lech Borowiec, Poland (Figs. 5-25). JESSOP, L and HAMMOND, P M (1993). Quantitative sampling of Coleoptera in northeast woodlands using flight interception traps. Transactions of the Natural History Society of Northumbria 56, 41-60. JESSOP, L and HAMMOND, P M (1997). Assessment of Coleoptera assemblages in Phragmites reed beds using large area flight interception traps. Vasculum 82, 33-60. KERSLAKE, L (editor) (1998). Red Data Book for Northumberland. Transactions of the Natural History Society of Northumbria 58, 39-322. KRELL, F-T, JOHNSON, C, BOOTH, R and MENDEL, H (2005). The British Dienerella separanda (Reitter) is D. clathrata (Mannerheim): with a compilation of British records of D. clathrata and D. elongata (Curtis) (Latridiidae). The Coleopterist 14, 117-123. LUFF, M L (1998). Provisional atlas of the ground beetles (Coleoptera, Carabidae) of Britain. Biological Records Centre, Huntingdon. LUFF, M L, EYRE, M D and JESSOP, L (1996). Records of new and local Coleoptera in northeast England. Entomologist’s Gazette 47, 257-265. LUFF, M L and SHEPPARD, D A (1980). The Coleoptera of Castle Eden Dene: a supplement. Vasculum 65, 53-60. REFERENCES BALL, S G (1986). Invertebrate Site Register. Review of the invertebrate sites in England. Tyne and Wear. Nature Conservancy Council Report 70. LUFF, M L, SELMAN, B J and FOSTER, G N (1974). The Coleoptera of Castle Eden Dene. Transactions of the Natural History Society of Northumbria 41, 179-193. BALL, S G (1987). Invertebrate Site Register. Regional review of rare and notable invertebrates in north-east England. Nature Conservancy Council Report 72. LUFF. M L and WALKER, M (1981). Triplax scutellaris Charp. and other interesting Coleoptera recently found in Northumberland. Entomologist’s Monthly Magazine 117, 62. BOLD, T J (1872). A catalogue of the insects of Northumberland and Durham (Revision of Coleoptera). Natural History Transactions of Northumberland and Durham 4, 1-117. TWINN, P F G and HARDING, P T (1999). Provisional atlas of the longhorn beetles (Coleoptera, Cerambycidae) of Britain. Biological Records Centre, Huntingdon. COX, M L (2007). Atlas of the seed and leaf beetles of Britain and Ireland. Pisces Publications, Newbury. DUFF, A G (2008). Checklist of beetles of the British Isles. Duff, Wells. 122 123 APPENDIX Complete species list Those species treated in more detail above are indicated by ‘*’. Dytiscidae Carnivorous water beetles Hydroporus memnonius Nicolai, 1822 Ilybius fuliginosus (F., 1792) Helophoridae Helophorus flavipes (F., 1792) Helophorus grandis Illiger, 1798 Carabidae Ground beetles Cychrus caraboides (L., 1758) Carabus violaceus L., 1758 Leistus fulvibarbis Dejean, 1826 Leistus rufescens (F., 1775) *Leistus rufomarginatus (Duftschmid, 1812) NR *Leistus spinibarbis (F., 1775) NR Nebria brevicollis (F., 1792) Notiophilus biguttatus (F., 1779) Loricera pilicornis (F., 1775) Clivina fossor (L., 1758) Patrobus atrorufus (Stroem, 1768) Trechus secalis (Paykull, 1790) Asaphidion curtum (Heyden, 1870) Ocys harpaloides Serville, 1821. Bembidion lampros (Herbst, 1784) Bembidion biguttatum (F., 1779) Bembidion mannerheimii Sahlberg, 1827 *Pterostichus cristatus (Dufour, 1820) Notable Pterostichus madidus (F., 1775) Pterostichus melanarius (Illiger, 1798) Pterostichus niger (Schaller, 1783) Pterostichus nigrita agg. (comprises P. nigrita (Paykull, 1790) and P. rhaeticus Heer, 1837/8) Pterostichus strenuus (Panzer, 1796) Calathus fuscipes (Goeze, 1777) Calathus rotundicollis Dejean, 1828 Platynus assimilis (Paykull, 1790) Agonum fuliginosum (Panzer, 1809) *Agonum thoreyi Dejean, 1828 NR Curtonotus aulicus (Panzer, 1796) Amara communis (Panzer, 1797) Amara familiaris (Duftschmid, 1812) Amara plebeja (Gyllenhal, 1810) Harpalus rufipes (Degeer, 1774) *Ophonus rufibarbis (F., 1792) NR Trichocellus placidus (Gyllenhal, 1827) Bradycellus verbasci (Duftschmid, 1812) Badister bullatus (Schrank, 1798) Paradromius linearis (Olivier, 1795) Dromius quadrimaculatus (L., 1758) Hydrophilidae Water scavenger beetles Cercyon melanocephalus (L., 1758) Megasternum concinnum (Marsham, 1802) Anacaena globulus (Paykull, 1798) Histeridae Predaceous pill beetles Margarinotus striola (Sahlberg, 1819) Hydraenidae *Hydraena britteni Joy, 1907 NR Ptiliidae Feather wing beetles. *Ptinella cavelli (Broun, 1893) Acrotrichis intermedia (Gillmeister, 1845) Leiodidae *Agathidium varians Beck, 1817 NR Nargus velox (Spence, 1813) Nargus wilkinii (Spence, 1813) * Catops coracinus Kellner, 1846 NR Catops nigricans (Spence, 1813) Catops tristis (Panzer, 1793) Silphidae Carrion and burying beetles. Nicrophorus vespilloides Herbst, 1783 *Dendroxena quadrimaculata (Scopoli, 1772) Notable Silpha atrata L., 1758 Staphylinidae Rove beetles. Anthobium unicolor (Marsham, 1802) Olophrum piceum (Gyllenhal, 1810) Lesteva longoelytrata (Goeze, 1777) Omalium italicum Bernhauer, 1902 Omalium rivulare (Paykull, 1789) *Phloeonomus pusillus (Gravenhorst, 1806) NR Megarthrus bellevoyei (Saulcy, 1862) Megarthrus depressus (Paykull, 1789) Proteinus brachypterus (F., 1792) Lordithon thoracicus (F., 1777) Tachyporus chrysomelinus (L., 1758) 124 Tachyporus hypnorum (F., 1775) Tachinus humeralis Gravenhorst, 1802 Tachinus laticollis Gravenhorst, 1802 Tachinus marginellus (F., 1781) Tachinus pallipes (Gravenhorst, 1806) Tachinus proximus Kraatz, 1855 Tachinus rufipes (Linnaeus, 1758) *Aleochara ruficornis Gravenhorst, 1802 Notable Autalia longicornis Scheerpeltz, 1947 Amischa analis (Gravenhorst, 1802) Geostiba circellaris (Gravenhorst, 1806) Mocyta fungi (Gravenhorst, 1806) Philhygra melanocera (Thomson, 1856) Atheta castanoptera (Mannerheim, 1830) Atheta setigera (Sharp, 1869) *Leptusa norvegica Strand, 1941 Notable *Gyrophaena strictula Erichson, 1839 Notable *Oligota apicata (Erichson, 1837) Notable Oxypoda procerula Mannerheim, 1830 Stenus bimaculatus Gyllenhal, 1810 Stenus fulvicornis Stephens, 1833 Stenus impressus Germar, 1824 Stenus juno (Paykull, 1789) Stenus picipes Stephens, 1830 Stenus pusillus Stephens, 1833 Lathrobium brunnipes (F., 1792) Lathrobium fulvipenne (Gravenhorst, 1806) Othius subuliformis (Stephens, 1833) Othius punctulatus (Goeze, 1777) Bisnius fimetarius (Gravenhorst, 1802) *Bisnius subuliformis (Gravenhorst, 1802) (formerly Philonthus fuscus). NR+ Gabrius splendidulus (Gravenhorst, 1802) Philonthus carbonarius (Gravenhorst, 1802) Philonthus cognatus Stephens, 1832 Philonthus decorus (Gravenhorst, 1802) Philonthus marginatus (Müller, 1764) Quedius curtipennis Bernhauer, 1908 Quedius fuliginosus (Gravenhorst, 1802) Quedius fumatus (Stephens, 1833) Quedius lateralis (Gravenhorst, 1802) Quedius maurorufus (Gravenhorst, 1806) Quedius molochinus (Gravenhorst, 1806) Quedius scintillans (Gravenhorst, 1806) Ocypus brunnipes (F., 1781) Ocypus olens (Mueller, 1764) The Devil’s coach horse Tasgius melanarius (Heer, 1839) Ontholestes tessellatus (Geoffroy, 1785) Scarabaeidae Dung beetles and chafers. Aphodius contaminatus (Herbst, 1783) Aphodius rufipes (L., 1758) Clambidae Clambus punctulum (Beck, 1817) Scirtidae Microcara testacea (L., 1767) Cyphon coarctatus Paykull, 1799 Cyphon ochraceus Stephens, 1830 Elateridae Click beetles. Hypnoidus riparius (F., 1792) Denticollis linearis (L., 1758) Athous haemorrhoidalis (F., 1801) Agriotes pallidulus (Illiger, 1807) Dalopius marginatus (L., 1758) Melanotus villosus (Geoffroy in Fourcroy, 1785) Cantharidae Soldier beetles. Podabrus alpinus (Paykull, 1798) Cantharis cryptica Ashe, 1947 Cantharis nigricans (Müller, 1776) Cantharis pellucida F., 1792 Rhagonycha fulva (Scopoli, 1763) Rhagonycha lignosa (Müller, 1764) Rhagonycha limbata Thomson, 1864 Rhagonycha testacea (L., 1758) Malthinus flaveolus (Herbst, 1786) Malthodes marginatus (Latreille, 1806) Dermestidae Hide beetles. Anthrenus fuscus Olivier, 1789 Anobiidae Wood borers. Ptilinus pectinicornis (L., 1758) Kateretidae Brachypterus glaber (Newman, 1834) Kateretes rufilabris (Latreille, 1807) Nitidulidae Pollen and sap beetles. Meligethes aeneus (F., 1775) Epuraea aestiva (L., 1758) Epuraea melanocephala (Marsham, 1802) Epuraea pallescens (Stephens, 1835) 125 Scraptiidae Tumbling flower beetles. Anaspis maculata (Geoffroy in Fourcroy, 1785) Anaspis regimbarti Schilsky, 1895 Anaspis rufilabris (Gyllenhal, 1827) Monotomidae Rhizophagus dispar (Paykull, 1800) Cryptophagidae Silken fungus beetles. Telmatophilus typhae (Fallen, 1802) Cryptophagus dentatus (Herbst, 1793) Cryptophagus denticulatus Heer, 1841 Cryptophagus scanicus (L., 1758) Atomaria mesomela (Herbst, 1792) Atomaria nitidula (Marsham, 1802) Atomaria rubella Heer, 1841 Cerambycidae Longhorn beetles. Grammoptera ruficornis (F., 1781) *Pachytodes cerambyciformis (Schrank, 1761) NR *Phymatodes testaceus (L., 1758) NR+ Chrysomelidae Leaf beetles. Cryptocephalus labiatus (Linnaeus, 1761) *Chrysomela aenea L., 1758 NR Galerucella lineola (F., 1781) *Pyrrhalta viburni (Paykull, 1799) NR+ Phyllotreta flexuosa (Illiger, 1794) Aphthona euphorbiae (Schrank, 1781) Crepidodera fulvicornis (F., 1792) Psylliodes affinis (Paykull, 1799) Byturidae Raspberry beetles. Byturus tomentosus (Degeer, 1774) Cerylonidae Cerylon ferrugineum Stephens, 1830 Endomychidae *Endomychus coccineus (L., 1758) NR Coccinellidae Ladybird beetles. Scymnus nigrinus Kugelann, 1794 Exochomus quadripustulatus (L., 1758) Anisosticta novemdecimpunctata (L., 1758) Propylea quattuordecimpunctata (L., 1758) *Halyzia sedecimguttata (L., 1758) NR Anatis ocellata (L., 1758) Adalia decempunctata (L., 1758) Fig. 2. Leistus rufomarginatus (Duftschmid) Length 9mm Fig. 3. Leistus spinibarbis (F.) Length 9.5mm Fig. 4. Pterostichus cristatus (Dufour) Length 16mm Fig. 5. Agonum thoreyi Dejean 7mm Fig. 6. Ophonus rufibarbis (F.) Length 8mm Fig. 7. Hydraena britteni Joy Length 2mm Rhynchitidae Deporaus betulae (L., 1758) Attelabidae *Attelabus nitens (Scopoli, 1763) Notable Erirhinidae *Grypus equiseti (F., 1775) Notable Notaris acridulus (L., 1758) Latridiidae Mould beetles. Cartodere bifasciata (Reitter, 1877) Cartodere nodifer (Westwood, 1839) Enicmus transversus (Olivier, 1790) *Dienerella elongata (Curtis, 1830) NR+ Cortinicara gibbosa (Herbst, 1793) Curculionidae Weevils Archarius pyrrhoceras Marsham, 1802 *Curculio villosus F., (1781 Notable) Anthonomus pedicularius (L., 1758) Orchestes fagi (L., 1758) Orchestes quercus (L., 1758) Nedyus quadrimaculatus (L., 1758) Parethelcus pollinarius (Forster, 1771) Euophryum confine (Broun, 1881) Strophosoma melanogrammum (Forster, 1771) Barynotus obscurus (F., 1775) Otiorhynchus singularis (L., 1767) Phyllobius argentatus (L., 1758) Barypeithes pellucidus (Boheman, 1834) Sciaphilus asperatus (Bonsdorff, 1785) Anoplus plantaris (Naezen, 1794) Cisidae Fungus beetles. Cis bidentatus (Olivier, 1790) Cis bilamellatus Wood, 1884 Cis fagi Waltl, 1839 Cis nitidus (F., 1792) Tetratomidae *Tetratoma fungorum F., 1790 NR Salpingidae *Vincenzellus ruficollis (Panzer, 1794) NR Salpingus planirostris (F., 1787) Salpingus ruficollis (L., 1761) 126 127 Fig. 8. Agathidium varians Beck Length 2.5mm Fig. 9. Catops coracinus Kellner Length 3mm Fig. 14. Endomychus coccineus (L.) Length 4.6mm Fig. 15. Halyzia sedecimguttata (L.) Length 6mm Fig. 10 Dendroxena quadrimaculata (Scopoli) Length 12mm Fig. 11. Phloeonomus pusillus (Gravenhorst) Length 1.8mm Fig. 16. Dienerella elongata (Curtis) Length 1.5mm Fig. 17. Tetratoma fungorum F. Length 4mm Fig. 12. Leptusa fumida (Erichson) Length 2.3mm Fig. 13. Bisnius subuliformis (Gravenhorst) Length 6mm Fig. 18. Vincenzellus ruficollis (Panzer) Length 4mm Fig. 19. Pachytodes cerambyciformis (Schrank) Length 10mm 128 129