SHILAP Revista de Lepidopterología
Sociedad Hispano-Luso-Americana de Lepidopterología
[email protected]
ISSN (Versión impresa): 0300-5267
ESPAÑA
2007
M. F. V. Corley / E. Marabuto / P. Pires
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL (INSECTA: LEPIDOPTERA)
SHILAP Revista de Lepidopterología, septiembre, año/vol. 35, número 139
Sociedad Hispano-Luso-Americana de Lepidopterología
Madrid, España
pp. 321-334
Red de Revistas Científicas de América Latina y el Caribe, España y Portugal
Universidad Autónoma del Estado de México
http://redalyc.uaemex.mx
SHILAP Nº 139
24/9/07
19:08
Página 321
SHILAP Revta. lepid., 35 (139), septiembre 2007: 321-334
CODEN: SRLPEF
ISSN:0300-5267
New Lepidoptera for the fauna of Portugal
(Insecta: Lepidoptera)
M. F. V. Corley, E. Marabuto & P. Pires
Abstract
75 species are added to the Portuguese Lepidoptera fauna and two species deleted, mainly as a result of recent
fieldwork by the authors.
KEY WORDS: Insecta, Lepidoptera, distribution, Portugal.
Lepidoptera novo para a fauna de Portugal
(Insecta: Lepidoptera)
Resumo
Como resultado do trabalho de campo desenvolvido recentemente pelos autores, são adicionadas 75 espécies
de Lepidoptera para a fauna de Portugal e duas são retiradas da lista nacional.
PALAVRAS CHAVE: Insecta, Lepidoptera, distribuição geográfica, Portugal.
Nuevos Lepidoptera para la fauna de Portugal
(Insecta: Lepidoptera)
Resumen
Con el trabajo de campo efectuado recientemente por los autores del presente trabajo se añaden 75 especies de
Lepidoptera a la fauna de Portugal y se suprimen dos especies de la lista nacional.
PALABRAS CLAVE: Insecta, Lepidoptera, distribución geográfica, Portugal.
Introduction
Recent field work in Portugal has resulted in the accumulation of a substantial number of records
of Lepidoptera species previously unrecorded or only doubtfully recorded from Portugal. 75 species are
listed below, of which 16 are also new to the Iberian Peninsula. Two species are deleted from the Portuguese fauna.
Material and Methods
Most species were captured at mercury vapour light over a white sheet. Where this was not the
case, the means of capture is given. Specimens are retained in the collections of the original collectors,
unless otherwise stated.
The order and nomenclature of families and species follows VIVES MORENO (1994) with some
modifications reflecting more recent works. The nomenclature of plant names follows TUTIN et al.
(1964-1980).
For most species in the list that follows a reference is given to a work in which the genitalia are
321
SHILAP Nº 139
24/9/07
19:08
Página 322
M. F. V. CORLEY, E. MARABUTO & P. PIRES
figured. In many cases the same work is useful for the whole of a family, so the reference is given after
the family heading. This has not been considered necessary for most Macrolepidoptera for which good
illustrations of the imagines are available in several books. Two species in this list have been identified
from pupal exuviae, using PATOGKA & TURGÁNI (2005).
Following each species newly recorded for Portugal, a summary of the previously known distribution is given. The expression “Middle latitudes of Europe” is frequently used here, referring to a broad
band with imprecise borders extending approximately from southern France or northern Spain and England or Ireland in the west, across Europe bordered on the north by the Baltic Sea and in the south by a
line crossing Italy, Croatia, Bosnia and Serbia, finally following the Romanian-Bulgarian border to the
Black Sea. Many species that essentially belong to this zone extend beyond its northern boundary into
northern England or Scotland and into southern parts of Fennoscandia. A considerable number of species
belonging to this zone have been found to extend into northern Portugal, either in the mountains bordering Galicia in Spain, or in the coastal regions of Minho and Douro Litoral. In Spain these species may
have been recorded in the Cantabrian mountains and sometimes in the foothills of the Pyrenees; probably they also occur in Galicia, but the Galician Microlepidoptera in particular are very little studied.
The account for most species ends with an indication of the known larval food-plant. In nearly all
cases this is based on information from outside Portugal, and for this reason is placed in square brackets [ ]. Although the foodplant in Portugal is most likely to be the same as elsewhere in Europe, or at
least a closely related species, this cannot be assumed without verification. Some of the new Portuguese
records are based on larval or pupal findings, in which case outside information on foodplant is not given, unless it adds to the Portuguese data.
Abbreviations and symbols
coll.
conf.
det.
ZMUP
*
**
collection
confirmed by
determined by
Natural History Museum of the Oporto Science Faculty
New for Portugal
New for the Iberian Peninsula
Provinces:
ALG
AAL
BAL
BA
BB
BL
DL
E
M
TM
Algarve
Alto Alentejo
Baixo Alentejo
Beira Alta
Beira Baixa
Beira Litoral
Douro Litoral
Estremadura
Minho
Trás-os-Montes
Localities with UTM 10 km square and altitude: (District in brackets)
Abreiro station (Vila Flor)
Alto dos Moinhos (Lisboa)
Amieiro (Alijó)
Apúlia (Esposende)
Barragem da Serra Serrada, Serra de Montesinho (Bragança)
Braçais (Marvão)
Brunheda (Carrazeda de Ansiães)
322 SHILAP Revta. lepid., 35 (139), septiembre 2007
PF47
MC88
PF37
NF19
PG84
PD45
PF37
170 m
90 m
150 m
10 m
1300 m
600 m
160 m
SHILAP Nº 139
24/9/07
19:08
Página 323
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
Cabo de São Vicente (Vila do Bispo)
Casmilo, Serra de Janeanes (Condeixa-a-Nova)
Ceira (Coimbra)
Contim, Santalha (Vinhais)
Covão do Boi, Serra da Estrela (Manteigas)
Covões, Vale do Zêzere, Serra da Estrela (Manteigas)
Fiolhal (Carrazeda de Ansiães)
Freixo de Numão (Vila Nova de Foz Côa)
Fresulfe (Vinhais)
Gondesende (Bragança)
Horta, Serra da Lousã (Lousã)
Ilha da Culatra (Faro)
Lama Grande, Serra de Montesinho (Bragança)
Madriz (Soure)
Minhota (Marvão)
Moimenta (Vinhais)
Monte Barata (Castelo Branco)
Monte da Lage (Serpa)
Pinhão (Alijó)
Poço do Inferno, Serra da Estrela (Manteigas)
Ponte de Castrelos (Bragança)
Ponte de Parâmio (Vinhais)
Póvoa e Meadas (Castelo de Vide)
Ribeira da Azenha (Fundão)
Ribeiradio (Oliveira de Frades)
Salgueirais (Celorico da Beira)
Santa Clara (Coimbra)
São Fiel (Castelo Branco)
São Lourenço (Carrazeda de Ansiães)
São Romão (São Brás de Alportel)
Seia
Serra da Nogueira (Bragança)
Setúbal
Soutelo (Bragança)
Valinhas, Monte Córdova (Santo Tirso)
Vesúvio (Vila Nova de Foz Côa)
Vila Franca (Viana do Castelo)
Vilamoura (Loulé)
Vilarinho, Espinhosela (Bragança)
Vilarinho das Azenhas (Vila Flor)
NA09
NE43
NE54
PG54
PE16
PE26
PF36
PF44
PG74
PG73
NE63
PA09
PG84
NE34
PD36
PG64
PD49
PC30
PF26
PE27
PG73
PG74
PD27
PE35
NF50
PE39
NE45
PE23
PF37
NB91
PE17
PG72
NC06
PG83
NF47
PF45
NG21
NB70
PG74
PF58
75 m
300 m
100 m
600 m
1900 m
1100 m
130 m
450 m
650 m
790 m
660 m
4m
1390 m
6m
650 m
760 - 890 m
230 m
200 m
100 m
1200 m
640 m
770 m
360 m
490 m
340 m
940 m
80 m
480 m
160 m
250 m
500 m
1320 m
40 m
1000 m
280 m
120 m
5m
10 m
790 m
190 m
List of families and species
NEPTICULIDAE (see JOHANSSON et al., 1990, and LASTUVKA & LASTUVKA, 1997)
* Simplimorpha promissa (Staudinger, 1871)
TM: Vacated mines on Pistacia terebinthus, by R. Tua at Vilarinho das Azenhas, 25-VII-2006,
Corley, det. van Nieukerken. Southern Europe and the warmer parts of Central Europe. [Leaf-miner in
Cotinus, Rhus and Pistacia spp.].
* Stigmella speciosa (Frey, 1857)
TM: Vacated mines on Acer monspessulanum, by R. Tua at Abreiro station, 22-VII-2006, Corley,
conf. van Nieukerken. Middle latitudes of Europe and the south-east. [Leaf miner in several species of
Acer, particularly A. pseudoplatanus].
SHILAP Revta. lepid., 35 (139), septiembre 2007 323
SHILAP Nº 139
24/9/07
19:08
Página 324
M. F. V. CORLEY, E. MARABUTO & P. PIRES
**Ectoedemia occultella (Linnaeus, 1767)
TM: Mines on Betula pubescens, 1 km west of Barragem da Serra Serrada, Montesinho, 26-IX2005, Corley, conf. van Nieukerken. Holarctic species, present in all Europe except the south.
TISCHERIIDAE (see MEDVEDEV, 1989 and PUPLESIS & DISKUS, 2003)
* Tischeria decidua Wocke, 1876
M: Flying in late afternoon in woodland near R. Lima, north of Vila Franca, 15-V-2006, Corley.
Middle latitudes of Europe. [Larva mines leaves of various Quercus spp. and Castanea].
HELIOZELIDAE (see HEATH, 1976, for leafmines)
**Antispila treitschkiella (Fischer von Röslerstamm, 1843)
TM: Vacated mines on Cornus sanguinea, Fresulfe, 25-IX-2005, Corley, conf. van Nieukerken.
Middle latitudes of Europe and the south-east.
ADELIDAE (see KÜPPERS, 1980)
* Nemophora degeerella (Linnaeus, 1758)
TM: Flying in morning sunshine, Moimenta, 4-VI-2005, J. M. Grosso-Silva, in coll. Corley, det.
Corley. All Europe except the far north and far south.
[The larva is reported to feed on Anemone, Stellaria, Rumex and Vaccinium, but also on leaf-litter].
TINEIDAE
Tinea pellionella Linnaeus, 1758 (see ROBINSON, 1979)
BB: At light outside house, Ribeiro da Azenha, 5-IX-2006 P. Sousa, genitalia preparation in coll.
Corley, det. Corley. Old records (BA: Seia, 11-VI-1880 Eaton (STAINTON, 1881); BB: São Fiel,
Mendes (MENDES, 1905); E: Setúbal, Vieilledent (VIEILLEDENT, 1905) were treated as unconfirmed by ROBINSON (1979). This species was also listed for Portugal without data by PETERSEN &
GAEDIKE (1992) and VIVES MORENO (1994). Cosmopolitan. [The larva feeds mainly on dry dead
animal matter such as wool, fur and feathers].
* Tinea flavofimbriella (Chrétien, 1925) (see PETERSEN, 1957 and AGENJO, 1962)
TM: By R. Tua at São Lourenço, 20-VII-2006, Corley (2 females). In the absence of males, it is
not possible to be certain of its generic placement, but it is very unlikely that it is correctly placed in
Tinea. Food source unknown.
DOUGLASIIDAE (see GAEDIKE, 1974)
*Tinagma ocnerostomella (Stainton, 1850)
DL: Casmilo, Serra de Janeanes, 13-V-2006, Corley and Pires. Middle latitudes of Europe and the
south-east. This species has recently been added to the Spanish fauna (GAEDIKE, 2007). [The larva
feeds in the stems of Echium].
GRACILLARIIDAE (see MEDVEDEV, 1981)
**Caloptilia cuculipennella (Hübner, 1796)
TM: Pupal exuviae on Fraxinus angustifolia, by R. Tuela east of Moimenta, 3-VI-2005, Corley.
Middle latitudes of Europe, reaching northwards into Fennoscandia. [Larva on Ligustrum and Fraxinus
spp.].
324 SHILAP Revta. lepid., 35 (139), septiembre 2007
SHILAP Nº 139
24/9/07
19:08
Página 325
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
* Caloptilia elongella (Linnaeus, 1761)
M: Near R. Lima, north of Vila Franca, 15-V-2006, Corley. Middle and northern latitudes of Europe, also in Greece. [Larva on Alnus].
**Caloptilia fidella (Reutti, 1853)
TM: Larvae on Humulus lupulus, by R. Tua at Brunheda, 21-VII-2006, Corley. Central Europe
from France to Russia and Ukraine.
* Phyllonorycter harrisella (Linnaeus, 1761)
DL: Valinhas, Monte Córdova, 16-V-2006 and 7-IX-2006, Corley. Middle latitudes of Europe, rare
in the south-east. [Leaf miner on deciduous Quercus spp.].
**Phyllonorycter monspessulanella (Fuchs, 1897)
TM: Mines on Acer monspessulanum, by R. Tua at Abreiro station, 22-VII-2006, Corley (det.
from pupal exuviae). France, Germany and Italy.
PLUTELLIDAE (see GAEDIKE, 1970)
* Acrolepiopsis marcidella (Curtis, 1850)
BL: Casmilo, Serra de Janeanes, 13-V-2006, Corley & Pires. Western Europe from south England
to Spain and Italy, Croatia, Sardinia. [Larva on Ruscus].
LYONETIIDAE
**Leucoptera aceris (Fuchs, 1903) (see MEY, 1994)
TM: Vacated mines on Acer monspessulanum, Freixo de Numão, 22-IX-2005, Corley; one female
from occupied mine on Acer monspessulanum, by R. Tua at Abreiro Station, 22-VII-2006, Corley. Central Europe. [The larva also mines Acer campestre].
OECOPHORIDAE (see EMMET & LANGMAID, 2002a; TOKÁR et al, 2004)
* Borkhausenia minutella (Linnaeus, 1758)
TM: By R. Tuela, east of Moimenta, 2-VI-2005, Corley. Known from nearly all the countries of
Europe. [The larva feeds on dead vegetable matter].
DEPRESSARIIDAE (see EMMET & LANGMAID, 2002a)
* Agonopterix ocellana (Fabricius, 1775)
DL: Valinhas, Monte Córdova, 16-V-2006, Corley. Middle latitudes of Europe and much of the
north. [Larva on Salix spp].
**Agonopterix arenella ([Denis & Schiffermüller], 1775)
TM: Ponte de Parâmio, 3-VI-2005, Corley; larva on Centaurea sp., by R. Assureira at Contim,
Santalha, 5-VI-2005, Corley. Middle latitudes of Europe and southern Fennoscandia; patchily distributed in south Europe with records from Greece and Sardinia. [The larva feeds on various Asteraceae,
including Cirsium, Carduus, Serratula and Arctium].
* Agonopterix liturosa (Haworth, 1811)
TM: By R. Tuela, east of Moimenta, 25-IX-2005, Corley. Central, western and northern Europe to
central Fennoscandia. [Larva feeds on Hypericum spp.].
SHILAP Revta. lepid., 35 (139), septiembre 2007 325
SHILAP Nº 139
24/9/07
19:08
Página 326
M. F. V. CORLEY, E. MARABUTO & P. PIRES
COLEOPHORIDAE
**Coleophora gryphipennella (Hübner, 1796) (see EMMET, 1996)
TM: Gondesende, 5-VI-2005, Corley. Middle latitudes of Europe and southern Fennoscandia.
[Larva on Rosa].
* Coleophora internitens Baldizzone & Wolf, 1999 (see BALDIZZONE & WOLF, 1999)
ALG: At actinic light, Ilha da Culatra, 14-VIII-2005 J. P. Cardoso, det. Corley. A number of specimens in poor condition, not retained. Genitalia preparation in coll. Corley. Spain, Tunisia. Foodplant
unknown.
** Coleophora saponariella Heeger, 1848 (see RAZOWSKI, 1990)
TM: Empty larval cases on Saponaria officinalis by R. Tua at Vilarinho das Azenhas, 25-VII2006, Corley. Central Europe from France to Poland and Romania; Sweden.
ELACHISTIDAE (see TRAUGOTT-OLSEN & SCHMIDT NIELSEN, 1977)
**Elachista atricomella Stainton, 1849
BL: Ceira, 26-V-2006, Pires, det. Corley. Middle latitudes of Europe, north to southern Norway
and Sweden. [Leaf miner in Dactylis].
* Elachista chrysodesmella Zeller, 1850
M: Near R. Lima, north of Vila Franca, 15-V-2006, Corley. Patchily distributed in the middle latitudes of Europe from France to Russia and Albania; Spain; Sweden (Gotland). Absent from Britain.
[Leaf miner in Brachypodium and various other grasses].
**Elachista cinereopunctella (Haworth, 1828)
BL: Santa Clara, Coimbra, 12-V-2006, Corley and Pires. Middle latitudes of Europe; Greece;
Sweden (Gotland). [Leaf miner in Carex and various grasses].
AGONOXENIDAE (see KOSTER & SINEV, 2003)
**Chrysoclista splendida Karsholt, 1997
BA: Resting on Salix by day, Salgueirais, 13-VI-2005, Ferreira, det. Koster. The single male specimen has genitalia matching those of C. splendida, but there are slight differences in forewing pattern
from Central European material. Central Europe from France to Poland and Bulgaria; one record from
Sweden. [Larva presumed to feed under bark of Salix spp.].
COSMOPTERIGIDAE (see KOSTER & SINEV, 2003).
*Pyroderces wolschrijni Koster & Sinev, 2003
E: Alto dos Moinhos, Lisboa, 30-X-2006, Marabuto, det. Corley. South Spain, Malta and Morocco. [Foodplant unknown].
CHRYSOPELEIIDAE (see KOSTER & SINEV, 2003).
**Sorhagenia janiszewskae Riedl, 1962
TM: By R. Tua at Amieiro, 23-VII-2006, Corley. Middle latitudes of Europe and southern
Fennoscandia, but not known from Italy. [Larva in young twigs of Frangula and Rhamnus spp.].
326 SHILAP Revta. lepid., 35 (139), septiembre 2007
SHILAP Nº 139
24/9/07
19:08
Página 327
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
GELECHIIDAE (see ELSNER et al., 1999 and EMMET & LANGMAID, 2003b, except for Neofriseria baungaardiella and Scrobipalpa hyoscyamella).
* Metzneria neuropterella (Zeller, 1839)
TM: Lama Grande, Montesinho, 4-VI-2005, Corley. Nearly all the countries of Europe, absent
from the Mediterranean Islands. [Larva in seed head of Cirsium acaule or Centaurea nigra].
**Monochroa hornigi (Staudinger, 1883)
TM: By R. Tua at Brunheda, 21-VII-2006, Corley; BAL: Monte da Lage, 7-X-2006, Marabuto,
det. Corley. Middle latitudes of Europe and southern Fennoscandia, but poorly recorded and not known
from some countries within this range. [Larva in stems of Persicaria lapathifolia].
* Eulamprotes unicolorella (Duponchel, 1843)
TM: At car headlights, Gondesende, 5-VI-2005, Corley. Nearly all the countries of Europe, except
Greece, Albania and the Mediterranean Islands. Foodplant unknown.
* Bryotropha desertella (Douglas, 1850)
TM: Moimenta, 3-VI-2005, Corley. Almost all Europe. [Larva on various mosses].
* Altenia scriptella (Hübner, 1796)
TM: By R. Tua at São Lourenço, 20-VII-2006, Corley. Middle latitudes and south Europe. [Larva
on Acer campestre, A. pseudoplatanus and A. platanoides. In Portugal presumably on A. monspessulanum].
* Gelechia sabinellus (Zeller, 1839)
TM: By R. Tua at Brunheda, 21-VII-2006, Corley. Most of Europe north to southern Fennoscandia. [Larva on Juniperus communis and J. sabina. In Portugal presumably on J. oxycedrus].
* Neofriseria baungaardiella Huemer & Karsholt, 1999 (see HUEMER & KARSHOLT, 1999)
TM: By R. Tua at São Lourenço, 20-VII-2006, Corley. Greece and Spain. Foodplant unknown.
* Scrobipalpa hyoscyamella (Stainton, 1869) (see POVOLNX, 2002)
TM: By R. Tua at Vilarinho das Azenhas, 25-VII-2006, Corley; larvae in spun leaves and among
fruits of Solanum dulcamara, 12-IX-2006, Corley. This is a new food-plant for this species, previously only recorded on Hyoscyamus. Solanum dulcamara is also the food-plant of the closely related
Scrobipalpa costella (Humphreys & Westwood, 1845), which has not been recorded from Portugal.
Distribution uncertain, owing to confusion with S. costella, but definitely recorded from France and
Spain.
* Syncopacma suecicella (Wolff, 1958)
TM: Larva on Genista florida, 1 km west of Barragem da Serra Serrada, Montesinho, 4-VI-2005,
Corley. Western Europe from England (Cornwall) and south Sweden to Spain and Sardinia, eastwards
to Slovakia; Greece. [Larva is recorded from Genista pilosa elsewhere.].
TORTRICIDAE (see RAZOWSKI, 2002 and RAZOWSKI, 2003)
* Phtheochroa syrtana (Ragonot, 1888)
ALG: At actinic light, Ilha da Culatra, 14-VIII-2005, J. P. Cardoso, det. Corley. A number of specimens in poor condition, not retained. Genitalia preparation in coll. Corley. Spain, Algeria and Tunisia.
Foodplant unknown.
SHILAP Revta. lepid., 35 (139), septiembre 2007 327
SHILAP Nº 139
24/9/07
19:08
Página 328
M. F. V. CORLEY, E. MARABUTO & P. PIRES
* Aethes tesserana ([Denis & Schiffermüller], 1775)
M: Apúlia, 17-V-2006, Corley. Recorded from almost all the countries of Europe. [Larva in roots
and root-stock of various Asteraceae, particularly Picris].
* Cochylis dubitana (Hübner, 1799)
TM: R. Tuela, east of Moimenta, 2-VI-2005, Corley. Europe except the south-east. [Larva
polyphagous on Asteraceae, feeding on flowers and seeds].
* Ptycholoma lecheana (Linnaeus, 1758)
TM: Ponte de Castrelos, 26-V-2006, Marabuto. Almost all the countries of Europe, but absent
from most of the Mediterranean islands. [Polyphagous on leaves of trees and shrubs].
* Isotrias hybridana (Hübner, 1817)
TM: Ponte de Parâmio, 3-VI-2005, Corley, Marabuto and Pires, in coll. Corley. Central and South
Europe. [Larva on Crataegus, Acer and Quercus].
* Rhopobota naevana (Hübner, 1817)
M: Apúlia, 17-V-2006, Corley. Europe except the south-east. [Polyphagous on buds, flowers,
fruits and leaves of trees and shrubs].
Pelochrista mollitana (Zeller, 1847)
BL: Ceira, 26-V-2006, Pires, det. Corley. This species was listed for Portugal in RAZOWSKI
(1996), but we have not traced the record on which this was based. Scattered through much of middle
and south Europe but absent from a number of countries; south Sweden. Foodplant unknown.
* Rhyacionia pinicolana (Doubleday, 1849)
BL: Hortas, Serra da Lousã, 20-VI-2001, Pires, det. Corley. Most of Europe, absent from the
Mediterranean islands and parts of the south-east. [Larva mining shoots of Pinus].
**Grapholita fissana (Frölich, 1828)
TM: Netted in late morning sunshine, Vilarinho, Espinhosela, 6-VI-2005, Corley. South-east, east
and central Europe, extending westwards to France and northwards to south Sweden. [Larva in seedpods of Vicia cracca].
* Pammene argyrana (Hübner, 1799)
BL: Santa Clara, Coimbra, 12-V-2006, Corley and Pires; DL: Valinhas, Monte Córdova, 16-V2006, Corley. Europe except the northernmost parts, the south-east and the Mediterranean islands. [Larva lives as an inquiline in hymenopterous galls on Quercus robur].
* Pammene suspectana (Lienig & Zeller, 1846)
TM: By R. Tua at Brunheda, 21-VII-2006, Corley. Central and north-west Europe, southwards to north
Spain. [Larva under bark of Fraxinus excelsior].
* Dichrorampha plumbana (Scopoli, 1763)
TM: Netted in late afternoon sunshine, 1 km west of Barragem da Serra Serrada, Montesinho, 4VI-2005, Corley. Almost all the countries of Europe, but absent from most Mediterranean islands. [Larva in roots of Leucanthemum vulgare, Achillea millefolium and Tanacetum].
* Dichrorampha petiverella (Linnaeus, 1758)
TM: Netted at dusk, Ponte de Parâmio, 3-VI-2005, Pires, det. Corley, in coll. Corley. Almost all
328 SHILAP Revta. lepid., 35 (139), septiembre 2007
SHILAP Nº 139
24/9/07
19:08
Página 329
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
Europe except some south-eastern countries; absent from some Mediterranean islands. [Larva in roots
of Leucanthemum vulgare, Achillea millefolium, A. ptarmica and Tanacetum].
HETEROGYNIDAE (see ZILLI & RACHELI, 1989)
Heterogynis paradoxa Rambur, 1837
TM: Soutelo, cocoon, 6-VI-2005, Corley. Previously recorded from Portugal (TM: Vesúvio, IV1941, Silva Cruz (SILVA CRUZ & GONÇALVES, 1955)). The Silva Cruz collection (ZMUP) contains
one cocoon of H. penella (det. T. Monteiro as H. paradoxa) and 10 males, 8 of them from Vesúvio, IV1941. According to ZILLI & RACHELI (1989) the only reliable way to separate H. paradoxa and H.
penella is by the cocoons. These males are therefore indeterminable, and the Silva Cruz record of H.
paradoxa must be considered doubtful. Spain. [Larva on Cytisus].
PYRALIDAE
*Actenia rungsi Leraut, 2000 (see LERAUT, 2000 and SLAMKA, 2006)
ALG: Cabo de São Vicente, 8-IX-1991, Corley. Recorded as A. honestalis (Treitschke, 1829) in
PASSOS DE CARVALHO & CORLEY (1995). Spain, south France, north-west Africa. Foodplant unknown.
Delete Actenia honestalis (Treitschke, 1828) from Portuguese list.
Ten specimens in coll. Corley from various localities have been examined and found to belong to
A. rungsi (see above). Many other records exist, but are likely to belong to A. rungsi also. As there is no
evidence for the presence of A. honestalis in Portugal, this species should be deleted from the Portuguese fauna.
*Eccopisa effractella Zeller, 1848 (see ROESLER, 1973)
BL: Madriz, 23.vii.2003, P. Pires, det. Corley. Middle and south Europe. [Larva on leaves of
Malus, Prunus and Corylus].
*Assara conicolella (Constant, 1854) (see ROESLER, 1973)
BL: Ceira, Coimbra, 1.vi.2006, P. Pires, det. Corley. Spain, France and Corsica; Tunisia. [Larva in
cones of Pinus halepensis].
* Scoparia pyralella ([Denis & Schiffermüller], 1775) (see GOATER et al, 2005)
TM: Netted at dusk, Ponte de Parâmio, 3-VI-2005, Pires and Corley, in coll. Corley; netted in afternoon sunshine, 1 km west of Barragem da Serra Serrada, 4-VI-2005, Corley. S. pyralella klimeschi
Leraut, 1981 was recorded from Algarve (PASSOS DE CARVALHO & CORLEY, 1995) based on a
misidentification of S. staudingeralis (see CORLEY et al., 2000). All Europe. [Larva reported to feed
on decaying vegetable matter and on roots of Senecio jacobaea].
* Xanthocrambus delicatellus (Zeller, 1863) (see BLESZYNSKI, 1965)
ALG: São Romão, 12-IX-2001, Corley; TM: by R. Tua at São Lourenço, 20-VII-2006, Corley.
France, Spain, Sardinia and Sicily. [Foodplant unknown].
Delete Xanthocrambus caducellus (Müller-Rutz, 1909) from Portuguese list.
Recorded originally from TM: Pinhão, IX-1960, Silva Cruz (MONTEIRO, 1982a, 1982b). Monteiro examined three females, the above and two from Algarve (Odeleite, IX-1975 and São Romão, IX1976) (MONTEIRO & PASSOS DE CARVALHO, 1984). He found BLESZYNSKI’s (1965) genitalia
figures difficult to interpret, remarking that the Portuguese specimens appeared to be closest to the
North African X. watsoni Bleszynski, 1960, but settling for X. caducellus as the most probable match.
SHILAP Revta. lepid., 35 (139), septiembre 2007 329
SHILAP Nº 139
24/9/07
19:08
Página 330
M. F. V. CORLEY, E. MARABUTO & P. PIRES
Two further Algarve records were added by CORLEY et al., (2000), but Corley also found interpretation of the female genitalia problematic, and accepted Monteiro’s identification. Subsequently males
have been found in Algarve and in the Alto Douro region which are unquestionably X. delicatellus.
Monteiro’s material has not been re-examined, but it is assumed that this is also X. delicatellus.
* Opsibotys fuscalis ([Denis & Schiffermüller], 1775) (see MEDVEDEV, 1986 and SLAMKA, 1995)
TM: Netted at dusk, Ponte de Parâmio, 3-VI-2005, Corley. Almost all Europe, except Greece and
most Mediterranean islands. [Larva on flowers and seeds of Rhinanthus, Melampyrum, Pedicularis and
Solidago].
* Sitochroa verticalis (Linnaeus, 1758) (see MEDVEDEV, 1986 and SLAMKA, 1995)
TM: Lama Grande, Serra de Montesinho, 4-VI-2005, Corley; Ponte de Castrelos, 26-V-2006,
Marabuto. All Europe except some Mediterranean islands. [Larva apparently polyphagous, reported
feeding on Cirsium, Teucrium, Cytisus, Atriplex, Rumex and Diplotaxis].
PTEROPHORIDAE (see GIELIS, 1996)
*Hellinsia carphodactyla (Hübner, 1813)
BB: Monte Barata, 1-VII-2005, Pires, det. Corley. Middle latitudes of Europe, in the east extending north to Estonia and south to Albania. Not in Fennoscandia. [Larva on Inula, Buphthalmum and
Carlina, the spring generation in the shoots and the summer generation in the flower buds and seedheads].
**Stenoptilia gratiolae Gibeaux & Nel, 1990
TM: Resting on and flying over Gratiola officinalis, on right bank of R. Tua, opposite Brunheda,
21-VII-2006, Corley. Central and eastern Europe, extending to Belgium and Norway in the north-west.
[Larva on the flowers and seed-heads of Gratiola].
* Cnaemidophorus rhododactylus ([Denis & Schiffermüller], 1775)
TM: Larvae on Rosa, by R. Tuela east of Moimenta, 3-VI-2005, Corley. This species was
listed for Portugal in GIELIS (2003), but no record has been traced. Almost all Europe except the
far north.
* Oxyptilus parvidactyla (Haworth, 1811)
TM: Netted at dusk, Ponte de Parâmio, 3-VI-2005, Corley. Almost all Europe except the far north,
some Mediterranean islands and some countries in the south-east. [Larva on leaves of Pilosella].
GEOMETRIDAE
* Abraxas sylvata (Scopoli, 1763)
BA: On wall below street light, Ribeiradio, 17-VIII-2001, Marabuto. Almost all Europe, absent
from northernmost parts, some countries in the south-east and the Mediterranean islands. In the Iberian
peninsula known only from the foothills of the Pyrenees. [Larva on Ulmus].
* Crocallis albarracina Wehrli, 1940 (see DANTART et al, 1993)
AAL: Minhota, Serra de São Mamede, 18-IX-1995, Corley; Braçais, 28-IX-1998, Corley; BA:
Covão do Boi, 4-IX-2002, Corley. These specimens were originally identified as C. elinguaria (Linnaeus, 1758), but were re-examined at the suggestion of Victor Redondo. Two of the specimens on
which the records of C. elinguaria given by Marabuto et al. (2004) were based have been re-examined
and also belong here: BA: Covões, Vale do Zêzere, 22-VIII-2003, Marabuto; Poço do Inferno, 23-VIII330 SHILAP Revta. lepid., 35 (139), septiembre 2007
SHILAP Nº 139
24/9/07
19:08
Página 331
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
2004, Marabuto. The presence of C. elinguaria in Portugal now requires confirmation. Spain. [The larva has been found on Cupressus, but there must be other foodplants].
* Aethalura punctulata ([Denis & Schiffermüller], 1775)
TM: Ponte de Castrelos, 26-V-2006, Marabuto. Middle latitudes of Europe, absent from the
Mediterranean islands and some countries in the south-east. [Larva on Betula and Alnus].
* Idaea nigrolineata (Chrétien, 1911) (see HAUSMANN, 2004)
TM: By R. Tua at Fiolhal, 26-VII-2006, Corley. Spain (south-east and north-east), Morocco to
Tunisia. [Larva has been bred from Viscum cruciatum, but probably polyphagous on withered leaves,
like its congeners].
* Dysstroma truncata (Hufnagel, 1767)
M: Near R. Lima, north of Vila Franca, 15-V-2006, Corley; Apúlia, 17-V-2006, Corley. Almost all
Europe, but not recorded from Hungary, Albania and most Mediterranean islands. [Larva
polyphagous].
* Eupithecia weissi Prout, 1938 (see MIRONOV, 2003)
BL: Casmilo, Serra de Janeanes, 8-IX-2006, Corley, Marabuto and Pires, in coll. Corley. France,
Spain, Morocco and Algeria. [Larva on Chamaemelum in Morocco; foodplant unknown in Europe].
LASIOCAMPIDAE
* Eriogaster rimicola ([Denis & Schiffermüller], 1775)
AAL: Larva on Quercus pyrenaica, Póvoa e Meadas, 29-IV-2006, C. Franquinho, det. Marabuto.
Central and south Europe.
LYMANTRIIDAE
* Orgyia recens (Hübner, 1819)
TM: Larva on Quercus pyrenaica, Serra da Nogueira, 20-VI-2004, Marabuto (not reared); larva,
Gondesende, 5-VI-2005, Corley (not reared). Middle latitudes of Europe, southern parts of Fennoscandia. [Larva polyphagous].
NOCTUIDAE
* Pseudozarba bipartita (Herrich-Schäffer, 1850)
ALG: At light in restaurant, collected from table, Vilamoura, 1-VIII-2004, Marabuto. South Europe.
* Mesogona acetosellae ([Denis & Schiffermüller], 1775)
TM: By R. Tuela below Fresulfe, 23-IX-2005, Corley. Middle and south Europe, not reaching
northern coasts and absent from Mediterranean islands. [Larva on Quercus].
* Apamea sordens (Hufnagel, 1766)
TM: Moimenta, 4-VI-2005, Corley. Almost all Europe, except northernmost parts and some
Mediterranean islands. [Larva on Dactylis and other grasses].
* Mythimna languida (Walker, 1858) (see HACKER et al., 2002)
ALG: Vilamoura, 3-X-2004, Marabuto. Most of Africa and the south-western parts of Asia. In Europe recorded from southernmost parts, particularly Spain, Greece and some Mediterranean islands. AlSHILAP Revta. lepid., 35 (139), septiembre 2007 331
SHILAP Nº 139
24/9/07
19:08
Página 332
M. F. V. CORLEY, E. MARABUTO & P. PIRES
though many records are probably of migrants, there are indications that it may be established in parts
of Spain. [Foodplant unknown].
Acknowledgements
We are most grateful to João Pedro Cardoso, Sónia Ferreira, Carlos Franquinho, José Manuel
Grosso-Silva and Pedro Sousa for allowing us to examine material collected by them, and to publish
the resulting records. M. F. V. C. thanks Prof. Paulo Célio Alves of CIBIO for his interest and support,
Dr. Luzia Sousa for permission to examine the ZMUP collections, Victor Redondo for the recommendation to re-examine Crocallis elinguaria, Sjaak Koster for the identification of Chrysoclista splendida,
Dr. Erik J. van Nieukerken for identification of leafmines, the director of Parque Natural de Montesinho for accommodation and Sónia Ferreira, José Manuel Grosso-Silva and Pedro Sousa for assistance
and companionship in the field. E. M. thanks Dr. Dinis Cortes for company in the field at Ponte de Castrelos.
BIBLIOGRAPHY
AGENJO, R., 1962.– Resultados científicos de una Pensión de Estudios en el “Museum National d’Histoire Naturelle”, de París, con la descripción de un género y otra especie nuevos de lepidópteros españoles, dedicados
al Excmo. Sr. D. Jesús Rubio y García-Mina, Ministro de Educación Nacional.– Eos, 38: 147-189.
BALDIZZONE, G. & WOLF, H. W. van der, 1999.– New and little-known species of Coleophoridae from Spain
(Lepidoptera: Coleophoridae).– SHILAP Revta. lepid., 27(105): 19-37.
BLESZYNSKI, S., 1965.– Crambidae. In H. G. AMSEL, F. GREGOR & H. REISSER.– Microlepidoptera
Palaearctica, 1: 552 pp. Verlag Georg Fromme & Co. Wien.
CORLEY, M. F. V., GARDINER, A. J., CLEERE, N. & WALLIS, P. D., 2000.– Further additions to the Lepidoptera
of Algarve, Portugal (Insecta: Lepidoptera).– SHILAP Revta. lepid. 28(111): 245-319.
DANTART, J., DOMÍNGUEZ, M. & BAIXERAS, J., 1993.– Contribution to the knowledge of the genus Crocallis
Treitschke, 1825: Iberian species.– Nota lepid., 15: 195-216.
ELSNER, G., HUEMER, P. & TOKÁR, Z., 1999.– Die Palpenmotten (Lepidoptera, Gelechiidae) Mitteleuropas:
208 pp. Bratislava.
EMMET, A. M., (Ed.) 1996.– The Moths and Butterflies of Great Britain and Ireland, 3: 452 pp. Harley Books,
Colchester.
EMMET, A. M. & LANGMAID, J. R., 2002a.– The Moths and Butterflies of Great Britain and Ireland, 4(i): 326
pp. Harley Books, Colchester.
EMMET, A. M. & LANGMAID, J. R., 2002b.– The Moths and Butterflies of Great Britain and Ireland, 4(ii): 277
pp. Harley Books, Colchester.
GAEDIKE, R., 1970.– Revision der paläarktischen Acrolepiidae (Lepidoptera).–Ent. Abhandl., 38: 1-54.
GAEDIKE, R., 1974.– Revision der paläarktischen Douglasiidae (Lepidoptera).– Acta faun. ent. Mus. natn. Pragae,
15: 79-102.
GAEDIKE, R., 2007.– New and poorly known Lepidoptera from the West Palaearctic (Tineidae, Acrolepiidae, Douglasiidae, Epermeniidae).– Nota lepid., 29(3/4): 159-176.
GIELIS, C., 1996.– Pterophoridae.– In P. HUEMER, O. KARSHOLT & L. LYNEBORG (eds.).– Microlepidoptera
of Europe, 1: 222 pp. Apollo Books, Stenstrup.
GIELIS, C., 2003.– Pterophoroidea & Alucitoidea – In: World Catalogue of Insects, 4: 1-198. Apollo Books, Stenstrup.
GOATER, B., NUSS, M. & SPEIDEL, W., 2005.– Pyraloidea I. (Crambidae: Acentropinae, Evergestinae, Heliothelinae, Schoenobiinae, Scopariinae).– In P. HUEMER & O. KARSHOLT. (eds.).– Microlepidoptera of Europe,
4: 304 pp. Apollo Books, Stenstrup.
HACKER, H., RONKAY, L. & HREBLAY, M., 2002..– Hadeninae I.– Noctuidae Europaeae, 4: 419 pp. Entomological Press, Sorø.
HAUSMANN, A., 2004.– Sterrhinae.– In A. HAUSMANN (ed.).– The Geometrid Moths of Europe, 2: 600 pp.
Apollo Books, Stenstrup.
HEATH, J., (Ed.) 1976.– The Moths and Butterflies of Great Britain and Ireland, 1: 343 pp. The Curwen Press, London and Blackwell Scientific Publications, Oxford.
332 SHILAP Revta. lepid., 35 (139), septiembre 2007
SHILAP Nº 139
24/9/07
19:08
Página 333
NEW LEPIDOPTERA FOR THE FAUNA OF PORTUGAL
HUEMER, P., 1988.– A taxonomic revision of Caryocolum (Lepidoptera: Gelechiidae).– Bull. Br. Mus. nat. Hist.
57: 339-571.
HUEMER, P. & KARSHOLT, O., 1999.– Gelechiidae I (Gelechiinae: Teleiodini, Gelechiini).– In P. HUEMER, O.
KARSHOLT & L. LYNEBORG. (eds.).– Microlepidoptera of Europe, 3: 356 pp. Apollo Books, Stenstrup.
JOHANSSON, R., NIELSEN, E. S., NIEUKERKEN, E. J. van, & GUSTAFSSON, B., 1990.– The Nepticulidae and
Opostegidae (Lepidoptera) of North West Europe.– Fauna ent. scand., 23: 1-739.
KOSTER, J. C. & SINEV, S. Yu., 2003.– Momphidae, Batrachedridae, Stathmopodidae, Agonoxenidae, Cosmopterigidae, Chrysopeleiidae.– In P. HUEMER, O. KARSHOLT & L. LYNEBORG. (eds.).– Microlepidoptera of
Europe, 5: 387 pp. Apollo Books, Stenstrup.
KÜPPERS, P. V.,1980.–.– Untersuchungen zur Taxonomie und Phylogenie der Westpaläarktischen Adelinae (Lepidoptera: Adelidae).– Wiss. Beit. Karlsruhe, 7: 1-495.
LASTUVKA, A. & LASTUVKA, Z., 1997.– Nepticulidae Mitteleuropas: Ein illustrierter Begleiter: 230 pp. Konvoj, Brno.
LERAUT, P., 2006.– Contribution à l’étude du genre Actenia Guenée (Lepidoptera, Pyralidae. Pyralinae).– Revue fr.
Ent. (N. S.), 22 (4): 239-244.
MARABUTO, E., PIRES, P. & CARDOSO, J. P., 2004.– Contribuição para o conhecimento dos Macrolepidópteros
do Parque Natural da Serra da Estrela, Portugal (Lepidoptera).– Boln. S.E.A., 34: 171-175.
MEDVEDEV, G. S., (Ed.) 1981.– Opredelitel Nasekomich Evropeiskoi Chasti SSSR, 4, part 2. Lepidoptera: 786 pp.
Leningrad.
MEDVEDEV, G. S., (Ed.) 1986.– Opredelitel Nasekomich Evropeiskoi Chasti SSSR,. 4, part 3. Lepidoptera: 503
pp. Leningrad.
MEDVEDEV, G. S., (Ed.) 1989.– Keys to the Insects of the European Part of the USSR. 4, part 1. Lepidoptera: 991
pp. E. J. Brill, Leiden.
MENDES, C., 1905.– Lepidópteros de Portugal. II. Lepidópteros da região de S. Fiel (Beira Baixa).– Brotéria, Lisboa, 4: 166-177.
MEY, W., 1994.– Taxonomische Bearbeitung der westpaläarktischen Arten der Gattung Leucoptera Hübner, [1825].
s. l. (Insecta, Lepidoptera).– Dt. ent. Z. (N. S.), 41: 173-234.
MIRONOV, V., 2003.– Larentiinae II (Perizomini and Eupitheciini).– In A. HAUSMANN (ed.).– The Geometrid
Moths of Europe, 4: 463 pp. Apollo Books, Stenstrup.
MONTEIRO, T., 1982a.– Espécies portuguesas da Subfamilia Crambinae (Lep. Crambidae.).– Anais Fac. Ciênc.
Porto, 63: 167-206.
MONTEIRO, T., 1982b.– Crambinae novos para Portugal (Pyralidae, Lepidoptera).–– Bolm Soc. port. Ent., 7 (Supl.
A): 91-103.
MONTEIRO, T. & PASSOS DE CARVALHO, J., 1984.– Lepidópteros do Algarve.– Anais Fac. Ciênc. Porto, 64:
95-219.
PASSOS DE CARVALHO, J. & CORLEY, M. F. V., 1995.– Additions to the Lepidoptera of Algarve, Portugal.–
SHILAP Revta. lepid., 23(91): 191-230.
PATOGKA, J. & TURGÁNI, M., 2005.– Lepidoptera Pupae. Central European Species, 1: 542 pp; 2: 321 pp. Apollo Books, Stenstrup.
PETERSEN, G., 1957.– Die Genitalien der paläarktischen Tineiden (Lepidoptera, Tineidae).– Beit. ent., 7: 55-176,
338-379, 557-595.
PETERSEN, G. & GAEDIKE, R., 1992.– Artenbestand und geographische Verbreitung der Tineiden der Iberischen
Halbinsel.– SHILAP Revta. lepid., 20(80): 325-353.
POVOLNX, D., 2002.– Iconographia tribus Gnorimoschemini Regionis Palaearcticae: 110 pp., 87 pl. Slamka,
Bratislava.
PUPLESIS, R. & DISKUS, A., 2003.– The Nepticuloidea and Tischerioidea (Lepidoptera) - a global review, with
strategic regional revisions: 512 pp. Lutute, Kaunas.
RAZOWSKI, J., 1990.– Motyle (Lepidoptera) Polski. Coleophoridae.– Monografie Fauny Polski, 16: 1-270.
RAZOWSKI, J. 1996.– Tortricidae.– In O. KARSHOLT & J. RAZOWSKI (eds.).– The Lepidoptera of Europe: 130157. Apollo Books, Stenstrup.
RAZOWSKI, J., 2002.– Tortricidae of Europe. Tortricinae and Chlidanotinae, 1: 247 pp. Slamka, Bratislava.
RAZOWSKI, J., 2003.– Tortricidae of Europe. Olethreutinae, 2: 301 pp. Slamka, Bratislava.
ROBINSON, G. S., 1979.– Clothes moths of the Tinea pellionella complex: a revision of the world’s species.– Bull.
Br. Mus. nat. Hist. (Ent.), 38: 57-128.
SHILAP Revta. lepid., 35 (139), septiembre 2007 333
SHILAP Nº 139
24/9/07
19:08
Página 334
M. F. V. CORLEY, E. MARABUTO & P. PIRES
ROESLER, R.–U., 1973.– Phycitinae. 1 Teilband. Trifine Acrobasiina.– In H. G. AMSEL, F. GREGOR & H.
REISSER.– Microlepidoptera Palaearctica, 4: 752 pp. Verlag Georg Fromme & Co., Wien.
SILVA CRUZ, M. A. & GONÇALVES, T., 1955.– Notas Lepidopterólogicas. III. Contribuição Sistematica para o
Inventário dos Heteroceros Portugueses.– Mems Estud. Mus. zool. Univ. Coimbra, 230: 1-14.
SLAMKA, F., 1995.– Die Zünslerfalter (Pyraloidea) Mitteleuropas: 112 pp. Slamka, Bratislava.
SLAMKA, F., 2006.– Pyraloidea of Europe, 1: 138 pp. Slamka, Bratislava.
STAINTON, H. T., 1881.– Notes on the entomology of Portugal. V. Lepidoptera (continued). Microlepidoptera
(Tineina) collected by the Rev. A. E. Eaton in 1880.– Entomologist’s mon. Mag., 17: 246-249.
TOKÁR, Z., LVOVSKY, A. & HUEMER, P., 2005.– Die Oecophoridae s. l. (Lepidoptera) Mitteleuropas: 120 pp.
Slamka, Bratislava.
TRAUGOTT-OLSEN, E. & NIELSEN, E. S., 1977.– The Elachistidae (Lepidoptera) of Fennoscandia and Denmark.– Fauna ent. scand., 6: 1-299.
TUTIN, T. G., HEYWOOD, V. H., BURGES, N. A., MOORE, D. M., VALENTINE, D. H., WALTERS, S. M. &
WEBB, D. A., 1964-1980.– Flora Europaea, 1: xxxii + 464 pp., 2: xxvii + 455 pp., 3: xxix + 370 pp., 4: xxix
+ 505 pp., 5: xxxvi + 452 pp. Cambridge University Press, Cambridge.
VIEILLEDENT, P. 1905.– Lepidópteros da Região de Setúbal.– Brotéria, Lisboa, 4: 185-206.
VIVES MORENO, A., 1994.– Catálogo sistemático y sinonímico de los lepidópteros de la Península Ibérica y
Baleares (Insecta: Lepidoptera) (Segunda Parte): 775 pp. Ministerio de Agricultura, Pesca y Alimentación,
Madrid.
ZILLI, A. & RACHELI, T., 1989.– Revisional notes on Spanish Heterogynis Rambur, 1837 (Lepidoptera, Heterogynidae).– Entomologist’s Gaz., 40: 125-152.
M. F. V. C.
Pucketty Farm Cottage
Faringdon
GB-Oxfordshire SN7 8JP
GRAN BRETAÑA/ GREAT BRITAIN
E-mail:
[email protected]
y / and
CIBIO, Centro de Investigação em
Biodiversidade e Recursos Genéticos
Universidade do Porto
Campus Agrário de Vairão
P-4485-661 Varião
PORTUGAL / PORTUGAL
E. M.
Rua Maestro Frederico de Freitas, 5-7º dto
P-1500-399 Lisboa
PORTUGAL / PORTUGAL
E-mail:
[email protected]
P. P.
Rua das Lagoas, 175
P-3030-867 Ceira
PORTUGAL / PORTUGAL
E-mail:
[email protected]
(Recibido para publicación / Received for publication 2-VII-2007)
(Revisado y aceptado / Revised and accepted 15-VIII-2007)
334 SHILAP Revta. lepid., 35 (139), septiembre 2007