Papers by David Greenfield
Zenodo (CERN European Organization for Nuclear Research), Jun 6, 2023
A new species of dwarfgoby, Eviota taeiae n. sp. is described from Nuusafee Island, Samoa. The ne... more A new species of dwarfgoby, Eviota taeiae n. sp. is described from Nuusafee Island, Samoa. The new species is a member of the Eviota guttata complex that includes E. guttata, E. teresae, and E. albolineata, and is characterized by a complete cephalic sensory-canal pore pattern (pattern 1); a dorsal/anal-fin formula of 9/8; the first 7-10 pectoral-fin rays unbranched, the remainder branched; the fifth pelvic-fin ray 10-12% of length of the fourth ray; pelvic-fin-ray branches short, broad, and connected by membranes; distinct dark spots present on ventral side of head; the eye with four to six white spokes radiating outwards from pupil and separating red-orange on iris; rectangular blocks on the ventral half of the body taller than wide and separated from each other by narrow yellow bars less than a pupil diameter; and a red-orange blotch behind the eye connected to a red-orange spot at the back of the interorbital area.
Zenodo (CERN European Organization for Nuclear Research), Nov 14, 2022
A new species of dwarfgoby, Eviota fluctiphila, n. sp., is described from West Papua in the weste... more A new species of dwarfgoby, Eviota fluctiphila, n. sp., is described from West Papua in the western Pacific Ocean. It has a complete cephalic sensory-canal pore system (pattern 1), dorsal/anal-fin formula of usually 8/8, some pectoral-fin rays branched, the fifth pelvic-fin ray absent, the first dorsal fin not filamentous, no occipital or dark marks on the nape, the pectoral-fin base peppered with scattered melanophores, the first dorsal fin dark crossed by a pale band, two dark postanal spots above the anal-fin base, a dark internal blotch over the preural centrum on and above the midline, and a pear-shaped male urogenital papilla. Eviota fluctiphila occurs in shallow (0.2-2 m), high-energy reef habitats.
A new species of alpheid-shrimp-associated goby is described from Fiji based on 9 specimens, 17.2... more A new species of alpheid-shrimp-associated goby is described from Fiji based on 9 specimens, 17.2–23.4 mm SL. Diagnostic features include an exceptionally small size at maturity, a uniform black color, possession of vomerine teeth, a rounded caudal fin, 65–81 scales in the longitudinal series, 0–15 predorsal scales, body depth at pelvic-fin origin 4.4–5.2 in SL, snout length 4.4–8.4 in HL, and caudal-peduncle depth 3.3–3.8 in HL.<br>
FIGURE 8. Fourth pelvic-fin ray. A. Eviota fallax, paratype, WAM P. 33039.009, 16.2 mm. B. Eviota... more FIGURE 8. Fourth pelvic-fin ray. A. Eviota fallax, paratype, WAM P. 33039.009, 16.2 mm. B. Eviota melasma, USNM 227318, 17.5 mm. Photographs by D. W. Greenfield.
Data collection pre-workshop and editing of species accounts post-workshop was led by the Marine ... more Data collection pre-workshop and editing of species accounts post-workshop was led by the Marine Biodiversity Unit and particular acknowledgement and thanks go to Gina Ralph for her involvement, support and guidance. We especially thank Jack Buchanan, Jessica Deal, Claire Gorman, Christi Linardich and Emilie Stump for their tireless efforts, as well as numerous undergraduate interns who contributed to the process. We greatly appreciate the efforts of Caroline Pollock and Max Fancourt to review the new bony fish assessments. The species distribution maps were digitized through the combined efforts of all of the experts mentioned above, along with Mike Harvey, Christi Linardich, Jack Buchanan and Gina Ralph of the Marine Biodiversity Unit, who we are thankful to for preparing, editing and finalizing the species distribution maps. We also thank two peerreviewers for their helpful comments. Expert participants at the assessment workshops, March 2015, Fiji. v Whale Shark (Rhincodon typus), assessed as Endangered. © B.
FIGURE 3. Perulibatrachus aquilonarius, CAS 231007, front view of head. Photograph by K.K. Bineesh.
FIGURE 5. Perulibatrachus elminensis, 21cm TL, Ivory Coast. Photograph by A. Oddgeir.
FIGURE 2. Perulibatrachus aquilonarius, CAS 231007, dorsal view. Photograph by K.K. Bineesh.
FIGURE 6. Perulibatrachus rossignoli, 20 cm TL, Ghana. Photograph by A. Oddgeir.
Introduction Methods Descriptions Material Examined Sampling Effort Habitat Categories Data Analy... more Introduction Methods Descriptions Material Examined Sampling Effort Habitat Categories Data Analysis Species Accounts Labrisomidae Labrisotnus (1) L. albigenys (2) L. bucciferus (3) L. gobio (4) L. guppyi (5) L. haitiensis (6) L. kalisherae (7) L. nigricinctus (8) L. nuchipinnis Malacoctenus (9) M. aurolineatus (10) M. boehlkei (11) M. delalandei (12) M. erdmani (13) M. gilli 28 (14) M. macropus 28 (15) M. triangulates
FIGURE 4. Eviota melasma, photograph of live fish from Lizard Island, Australia. Photograph by A.... more FIGURE 4. Eviota melasma, photograph of live fish from Lizard Island, Australia. Photograph by A. González-Cabello.
FIGURE 7. Plot of eye diameter as percentage of HL versus SL for Eviota karaspila from Fiji and E... more FIGURE 7. Plot of eye diameter as percentage of HL versus SL for Eviota karaspila from Fiji and E. melasma from Australia, Indonesia, and Papua New Guinea.
FIGURE 5. Eviota melasma, photograph of live fish from Lizard Island, Australia. Photograph by A.... more FIGURE 5. Eviota melasma, photograph of live fish from Lizard Island, Australia. Photograph by A. González-Cabello.
FIGURE 1. Eviota karaspila, photograph of live fish from Fiji, Viti Levu, Nananui-i-Cake. Estimat... more FIGURE 1. Eviota karaspila, photograph of live fish from Fiji, Viti Levu, Nananui-i-Cake. Estimated length 22 mm SL. Photograph by J.E. Randall.
FIGURE 2. Eviota pellucida, OMNH 35616, Ishigaki-jima Island, Ryukyu Islands, Japan. Photograph b... more FIGURE 2. Eviota pellucida, OMNH 35616, Ishigaki-jima Island, Ryukyu Islands, Japan. Photograph by T. Suzuki.
Sixteen species of <em>Eviota</em> have been described since the comprehensive key to... more Sixteen species of <em>Eviota</em> have been described since the comprehensive key to the 107 species in the genus by Greenfield & Winterbottom (2016). The additional species are listed and discussed, along with their diagnostic characters and photographs, and where they fit into the original key. Corrections and updates to the 2016 key are included.
A new species of dwarfgoby, Eviota maculosa n. sp., is described from reefs in the Western Pacifi... more A new species of dwarfgoby, Eviota maculosa n. sp., is described from reefs in the Western Pacific Ocean. Specimens from West Papua (Fakfak), Sulawesi, and Pohnpei match the DNA sequences of the type population from Teluk Saleh, Sumbawa, Indonesia. The new species is most similar to E. punctulata, described from Fiji, but can be distinguished by pigmentation patterns on the fins. Phylogenetic analyses using both mitochondrial and nuclear genes confirm that the new species is part of a species complex with E. punctulata, E. tigrina, and two additional undescribed species from Samoa and Tonga. Within this complex, the new species and E. punctulata and E. tigrina share the synapomorphy of having multiple rows of tricuspid teeth in both the upper and lower jaws, which is unique within Eviota and rare in gobiids in general.<br>
Zootaxa, 2016
Two new species of dwarfgobies are described, Eviota asymbasia from the Sulu Sea, Philippine Isla... more Two new species of dwarfgobies are described, Eviota asymbasia from the Sulu Sea, Philippine Islands, south to Java and west to Damar, and E. bipunctata occurring from the Indian Ocean eastward into the western Pacific Ocean. Eviota asymbasia lacks the IT pore of the cephalic sensory-pore system and usually also the POP, has a dorsal/anal-fin formula of 8/8, has some pectoral-fin rays branched and usually 15 rays, two spots on the pectoral-fin base, no dark spot over the preural centrum, first dorsal fin may be filamentous, six ventral postanal spots, and the male genital papilla is not fimbriate. Eviota bipunctata belongs to the group with cephalic sensory-pore system pattern 2 (lacking only the IT pore), some pectoral-fin rays branched, pectoral-fin base with 1 or 2 prominent dark spots, dorsal/anal-fin formula usually 8/8, and no occipital spots.
Zootaxa, 2015
Two new species of Eviota from Yoron Island, the Ryukyu Islands, Japan, are described. Both speci... more Two new species of Eviota from Yoron Island, the Ryukyu Islands, Japan, are described. Both species belong to the cephalic sensory-canal pore system pattern 2 (lacking only the H [IT] pore); have some pectoral-fin rays branched; have a dorsal/anal-fin formula of 9/8; no dark spot over the ural centrum; no prominent distinct dark spots on the pectoral-fin base; no postocular spots; and no strong dark spots on the caudal fin. The species share the most characters with E. afelei, E. bimaculata and E. punctulata, but differ from Eviota afelei and E. punctulata by having two versus three dark marks over the anal-fin base, and from Eviota bimaculata by lacking the two dark, prominent occipital spots present in that species. Both species differ from all other described species of Eviota in fresh coloration. Eviota flavipinnata has bright golden-yellow dorsal fins and an orange anal fin, and a fifth pelvic-fin ray that is 12% the length of the fourth ray. Eviota rubrimaculata has clear dorsal fins with red spots, large red spots on the body, and lacks the fifth pelvic-fin ray.
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Papers by David Greenfield