Crocidurinae is a subfamily of small mammals in the shrew family Soricidae, which in turn is part of the order Eulipotyphla. A member of this family is called a crocidurine, or a white-toothed shrew. Crocidurinae is one of three subfamilies in Soricidae, along with the African shrews of Myosoricinae and the red-toothed shrews of Soricinae. They are found in Africa, Europe, and Asia, primarily in forests, savannas, shrublands, and grasslands, though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew, one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[1] to the hero shrew and Thor's hero shrew, at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates, small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 21 species are categorized as endangered species, while the Andaman shrew, Christmas Island shrew, Harenna shrew, Jenkins's shrew, Nicobar shrew, and Wimmer's shrew are categorized as critically endangered.

Gray shrew
Greater white-toothed shrew (Crocidura russula)

The 235 extant species of Crocidurinae are divided into nine genera; 192 of them are in Crocidura, the most species of any mammal genus.[2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]

Conventions

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IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (6 species)
 EN Endangered (21 species)
 VU Vulnerable (13 species)
 NT Near threatened (8 species)
 LC Least concern (128 species)
Other categories
 DD Data deficient (58 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

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The subfamily Crocidurinae consists of nine genera: Crocidura, containing 192 species; Diplomesodon and Feroculus with a single species each; Paracrocidura, with three species; Ruwenzorisorex, with one species; Scutisorex, containing two species; Solisorex, with one species, Suncus, containing nineteen species; and Sylvisorex, containing fifteen species.

Subfamily Crocidurinae

  • Genus Crocidura (white-toothed shrews): 191 species
  • Genus Diplomesodon (piebald shrew): one species
  • Genus Feroculus (Kelaart's long-clawed shrew): one species
  • Genus Paracrocidura (large-headed shrews): three species
  • Genus Ruwenzorisorex (Ruwenzori shrew): one species
  • Genus Scutisorex (hero shrews): two species
  • Genus Solisorex (Pearson's long-clawed shrew): one species
  • Genus Suncus (musk shrews and pygmy shrews): nineteen species
  • Genus Sylvisorex (forest shrews): fifteen species
Crocidurinae[4]  

Crocidurines

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The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]

Genus Crocidura Wagler, 1832 – 192 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
African black shrew


C. nigrofusca
Matschie, 1895
Central and eastern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[7]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [7]

African dusky shrew


C. caliginea
Hollister, 1916
Central Africa
 
Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[8]

Habitat: Forest[9]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [9]

African giant shrew

 

C. olivieri
(Lesson, 1827)

20 subspecies
  • C. o. anchietae
  • C. o. bueae
  • C. o. cara
  • C. o. cinereoaenea
  • C. o. darfurea
  • C. o. giffardi
  • C. o. guineensis
  • C. o. hansruppi
  • C. o. hedenborgiana
  • C. o. kivu
  • C. o. manni
  • C. o. martiensseni
  • C. o. nyansae
  • C. o. occidentalis
  • C. o. odorata
  • C. o. olivieri
  • C. o. spurelli
  • C. o. sururae
  • C. o. toritensis
  • C. o. zuleika
Sub-Saharan Africa
 
Size: 11–14 cm (4–6 in) long, plus 8–10 cm (3–4 in) tail[10]

Habitat: Forest, savanna, and shrubland[11]

Diet: Invertebrates[10]
 LC 


Unknown  [11]

Andaman shrew


C. andamanensis
Miller, 1902
South Andaman Island in India
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[12]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown  [12]

Andaman spiny shrew


C. hispida
Thomas, 1913
South Andaman Island in India
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[13]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [13]

Annamite shrew


C. annamitensis
Jenkins, Lunde, & Moncrieff, 2009
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[14]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [14]

Ansell's shrew


C. ansellorum
Hutterer & Dippenaar, 1987
South-central Africa
 
Size: 5–6 cm (2–2 in) long, plus 4–6 cm (2 in) tail[15]

Habitat: Forest[16]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [16]

Arabian shrew


C. arabica
Hutterer, 1988
Oman and Yemen
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[17]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [17]

Armenian shrew


C. armenica
Gureev, 1963
Armenia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[18]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [18]

Asian gray shrew


C. attenuata
H. Milne-Edwards, 1872
Eastern and southeastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[19]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [19]

Asian lesser white-toothed shrew

 

C. shantungensis
Miller, 1901

Two subspecies
  • C. s. quelpartis
  • C. s. shantungensis
Eastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[20]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [20]

Bailey's shrew


C. baileyi
Osgood, 1936
Ethiopia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[21]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [21]

Bale shrew


C. bottegoides
Hutterer & Yalden, 1990
Ethiopia
 
Size: 4–6 cm (2 in) long, plus 4–5 cm (2 in) tail[22]

Habitat: Grassland and forest[23]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [23]

Banka shrew


C. vosmaeri
Jentink, 1888
Bangka Island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[24]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [24]

Batak shrew


C. batakorum
Hutterer, 2007
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[25]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [25]

Bates's shrew


C. batesi
(Dollman, 1915)
Western Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[26]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [26]

Beccari's shrew


C. beccarii
Dobson, 1886
Sumatra
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[27]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [27]

Beletta shrew


C. yaldeni
Lavrenchenko, Voyta, & Hutterer, 2016
Ethiopia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[28]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [28]

Bicolored musk shrew


C. fuscomurina
(Heuglin, 1865)
Sub-Saharan Africa
 
Size: 4–8 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[29]

Habitat: Savanna, grassland, and desert[30]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [30]

Bicolored shrew

 

C. leucodon
(Hermann, 1780)
Europe and western Asia
 
Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[31]

Habitat: Shrubland and grassland[32]

Diet: Small mammals, frogs, toads, lizards, and invertebrates[31]
 LC 


Unknown  [32]

Black-footed shrew

 

C. nigripes
Miller & Hollister, 1921

Two subspecies
  • C. n. lipara
  • C. n. nigripes
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[33]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [33]

Blackish white-toothed shrew


C. nigricans
Bocage, 1889
Angola
 
Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[34]

Habitat: Unknown[35]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [35]

Bornean shrew


C. foetida
Peters, 1870

Three subspecies
  • C. f. doriae
  • C. f. foetida
  • C. f. kelabit
Borneo
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[36]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [36]

Bottego's shrew


C. bottegi
Thomas, 1898
Ethiopia
 
Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[37]

Habitat: Unknown[38]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [38]

Buettikofer's shrew


C. buettikoferi
Jentink, 1888
West-central Africa
 
Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[39]

Habitat: Forest[40]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [40]

Butiaba naked-tailed shrew


C. littoralis
Heller, 1910
Central Africa
 
Size: 8–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[41]

Habitat: Forest[42]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [42]

Cameroonian shrew


C. picea
Sanderson, 1940
West-central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[43]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [43]

Canarian shrew

 

C. canariensis
Hutterer, López-Jurado, & Vogel, 1987
Canary Islands
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Coastal marine, desert, and shrubland[44]

Diet: Snails and insects[44]
 EN 


Unknown  [44]

Caspian shrew


C. caspica
Thomas, 1907
Iran and Azerbaijan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[45]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [45]

Chinese white-toothed shrew


C. rapax
Allen, 1923

Four subspecies
  • C. r. kurodai
  • C. r. lutaoensis
  • C. r. rapax
  • C. r. tadae
Eastern Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[46]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [46]

Christmas Island shrew


C. trichura
Dobson, 1889
Christmas Island
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[47]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


0–200  [47]

Cinderella shrew


C. cinderella
Thomas, 1911
Western Africa
 
Size: 5–7 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[48]

Habitat: Savanna[49]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [49]

Congo white-toothed shrew


C. congobelgica
Hollister, 1916
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[50]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [50]

Cranbrook's white-toothed shrew


C. cranbrooki
Jenkins, Lunde, & Moncrieff, 2009
Northern Myanmar Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[51]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [51]

Cretan shrew


C. zimmermanni
Wettstein, 1953
Island of Crete in Greece
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown and rocky areas[52]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [52]

Crosse's shrew


C. crossei
Thomas, 1895
Western Africa
 
Size: 6–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[53]

Habitat: Forest[54]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [54]

Cyrenaica shrew


C. aleksandrisi
Vesmanis, 1977
Northern Libya
 
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[55]

Habitat: Shrubland[56]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [56]

Dent's shrew

 

C. denti
Dollman, 1915
Western and central Africa
 
Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[57]

Habitat: Forest and savanna[58]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [58]

Desert musk shrew


C. smithii
Thomas, 1895

Two subspecies
  • C. s. debalsaci
  • C. s. smithii
Western and Eastern Africa
 
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[59]

Habitat: Savanna[60]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [60]

Desperate shrew


C. desperata
Hutterer, Jenkins, & Verheyen, 1991
Tanzania
 
Size: 8–11 cm (3–4 in) long, plus about 7 cm (3 in) tail[61]

Habitat: Forest[62]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [62]

Dhofar shrew


C. dhofarensis
(Hutterer, 1988)
Yemen and Oman Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[63]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [63]

Doucet's musk shrew


C. douceti
Heim de Balsac, 1958
Western Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[64]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [64]

Dsinezumi shrew

 

C. dsinezumi
(Temminck, 1842)
Japan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[65]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [65]

East African highland shrew


C. allex
Osgood, 1910
Kenya and Tanzania
 
Size: 5–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[66]

Habitat: Grassland and inland wetlands[67]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [67]

Egyptian pygmy shrew


C. religiosa
(Geoffroy, 1827)
Northern Egypt
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[68]

Habitat: [69]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [69]

Eisentraut's shrew


C. eisentrauti
Heim de Balsac, 1957
Cameroon
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and forest[70]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [70]

Elgon shrew


C. elgonius
Osgood, 1910
Kenya and Tanzania
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[71]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [71]

Elongated shrew


C. elongata
Miller & Hollister, 1921
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[72]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [72]

Fingui white-toothed shrew


C. fingui
Ceríaco et al., 2015
Príncipe in São Tomé and Príncipe Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[73]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [73]

Fischer's shrew


C. fischeri
Pagenstecher, 1885
Kenya and Tanzania
 
Size: 6–10 cm (2–4 in) long, plus 4–6 cm (2 in) tail[74]

Habitat: Savanna[75]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [75]

Flat-headed shrew


C. planiceps
Heller, 1910
West-central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[76]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [76]

Flower's shrew


C. floweri
Dollman, 1915
Northern Egypt
 
Size: 5–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[77]

Habitat: Artificial terrain[78]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [78]

Fox's shrew


C. foxi
Dollman, 1915
Western and central Africa
 
Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[79]

Habitat: Savanna and forest[80]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [80]

Fraser's musk shrew


C. poensis
(Fraser, 1843)
Western Africa
 
Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[81]

Habitat: Shrubland and forest[82]

Diet: Arthropods and other invertebrates[81]
 LC 


Unknown  [82]

Gathorne's shrew


C. gathornei
Jenkins, 2013
Himalayas in India Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[83]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [83]

Glass's shrew


C. glassi
Heim de Balsac, 1966
Ethiopia
 
Size: 6–9 cm (2–4 in) long, plus 4–6 cm (2 in)tail[84]

Habitat: Shrubland, grassland, and inland wetlands[85]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [85]

Gmelin's white-toothed shrew


C. gmelini
Pallas, 1811
Central Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[86]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [86]

Goliath shrew


C. goliath
Thomas, 1906
Central Africa
 
Size: 15–18 cm (6–7 in) long, plus 10–14 cm (4–6 in) tail[87]

Habitat: Forest[88]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [88]

Gracile naked-tailed shrew


C. maurisca
Thomas, 1904
Central Africa
 
Size: 7–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[89]

Habitat: Inland wetlands and forest[90]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [90]

Grasse's shrew


C. grassei
Brosset, Dubost, & Heim de Balsac, 1965
West-central Africa
 
Size: 7–9 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[91]

Habitat: Forest[92]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [92]

Greater Mindanao shrew


C. grandis
Miller, 1911
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[93]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [93]

Greater red musk shrew

 

C. flavescens
(Geoffroy, 1827)
Southern Africa
 
Size: 8–12 cm (3–5 in) long, plus 3–6 cm (1–2 in) tail[94]

Habitat: Savanna and grassland[95]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [95]

Greater white-toothed shrew

 

C. russula
(Hermann, 1780)
Europe and northwestern Africa
 
Size: 6–9 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[96]

Habitat: Shrubland[97]

Diet: Invertebrates, as well as small mammals and small lizards[96]
 LC 


Unknown  [97]

Greenwood's shrew


C. greenwoodi
Heim de Balsac, 1966
Somalia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, and shrubland[98]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [98]

Guramba shrew


C. phaeura
Osgood, 1936
Ethiopia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[99]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [99]

Guy's shrew


C. guy
Jenkins, Lunde, & Moncrieff, 2009
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[100]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [100]

Güldenstädt's shrew


C. gueldenstaedtii
(Pallas, 1811)
Europe and western Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, inland wetlands, rocky areas, intertidal marine, and coastal marine[101]

Diet: Insects, as well as molluscs, amphipods, annelids, and small vertebrates[101]
 LC 


Unknown  [101]

Hainan Island shrew


C. wuchihensis
Sung, 1966
Southern China and northern Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[102]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [102]

Harenna shrew


C. harenna
Hutterer & Yalden, 1990
Ethiopia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[103]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown  [103]

Heather shrew


C. erica
Dollman, 1915
Angola
 
Size: About 10 cm (4 in) long, plus 5 cm (2 in) tail[104]

Habitat: Unknown[105]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [105]

Hildegarde's shrew


C. hildegardeae
Thomas, 1904
Central Africa
 
Size: 6–8 cm (2–3 in) long, plus 3–6 cm (1–2 in) tail[106]

Habitat: Forest[107]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [107]

Hill's shrew


C. hilliana
Jenkins & Smith, 1995
Thailand and Laos
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[108]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [108]

Horsfield's shrew


C. horsfieldii
(Tomes, 1856)
Scattered southern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[109]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [109]

Hun shrew


C. attila
Dollman, 1915
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[110]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [110]

Hutan shrew


C. hutanis
Ruedi & Vogel, 1995
Sumatra island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[111]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [111]

Indochinese shrew


C. indochinensis
Kloss, 1922
Southeastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[112]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [112]

Iranian shrew


C. susiana
Redding & Lay, 1978
Iran
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[113]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [113]

Jackass shrew


C. arispa
Spitzenberger, 1971
Turkey
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[114]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [114]

Jackson's shrew


C. jacksoni
Thomas, 1904
East-central Africa
 
Size: 5–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[115]

Habitat: Forest[116]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [116]

Javan ghost shrew


C. umbra
Demos, Achmadi, Handika, Maharadatunkamsi, Rowe, & Esselstyn, 2016
Java island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[117]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [117]

Javanese shrew


C. maxi
Sody, 1936
Indonesia and East Timor
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[118]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [118]

Jenkins's shrew


C. jenkinsi
Chakraborty, 1978
South Andaman Island in India
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[119]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown  [119]

Jouvenet's shrew


C. jouvenetae
Heim de Balsac, 1958
Western Africa
 
Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[120]

Habitat: Forest[121]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [121]

Kashmir white-toothed shrew


C. pullata
Miller, 1911
Northern India and Pakistan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[122]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [122]

Katinka's shrew


C. katinka
Bate, 1937
Western Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[123]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [123]

Kilimanjaro shrew


C. monax
Thomas, 1910

 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[124]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [124]

Kinabalu shrew


C. baluensis
Thomas, 1898
Northern Borneo
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[125]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [125]

Kivu long-haired shrew


C. lanosa
Heim de Balsac, 1968
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and inland wetlands[126]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [126]

Kivu shrew


C. kivuana
Heim de Balsac, 1968
Central Africa
 
Size: 6–9 cm (2–4 in) long, plus 6–8 cm (2–3 in) tail[127]

Habitat: Forest and inland wetlands[128]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [128]

Lamotte's shrew


C. lamottei
Heim de Balsac, 1968
Western Africa
 
Size: 8–9 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[127]

Habitat: Forest and savanna[129]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [129]

Large-headed forest shrew


C. grandiceps
Hutterer, 1983
Western Africa
 
Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[130]

Habitat: Forest[131]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [131]

Latona's shrew


C. latona
Hollister, 1916
Democratic Republic of the Congo
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[132]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [132]

Lesser Ryukyu shrew


C. watasei
Kuroda, 1924
Southern Japan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[133]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [133]

Lesser gray-brown musk shrew


C. silacea
Thomas, 1895
Southern Africa
 
Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[134]

Habitat: Rocky areas, grassland, shrubland, and savanna[135]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [135]

Lesser red musk shrew

 

C. hirta
Peters, 1852
Southern and southeastern Africa
 
Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[136]

Habitat: Savanna, grassland, and desert[137]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [137]

Lesser rock shrew


C. serezkyensis
Laptev, 1929
Western Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[138]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [138]

Lesser white-toothed shrew

 

C. suaveolens
(Pallas, 1811)
Europe and Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine[139]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [139]

Long-footed shrew


C. crenata
Brosset, Dubost, & Heim de Balsac, 1965
West-central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[140]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [140]

Long-tailed musk shrew


C. dolichura
Peters, 1876
Central Africa
 
Size: 6–7 cm (2–3 in) long, plus 7–8 cm (3 in) tail[141]

Habitat: Forest[142]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [142]

Lucina's shrew


C. lucina
Dippenaar, 1980
Ethiopia
 
Size: 8–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[143]

Habitat: Grassland and inland wetlands[144]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [144]

Ludia's shrew


C. ludia
Hollister, 1916
Central Africa
 
Size: 6–7 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[145]

Habitat: Forest[146]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [146]

Luzon shrew


C. grayi
Dobson, 1890
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[147]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [147]

Lwiro shrew


C. lwiroensis
Peterhans & Hutterer, 2013
Democratic Republic of the Congo Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[148]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [148]

MacArthur's shrew


C. macarthuri
St. Leger, 1934
Eastern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[149]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [149]

MacMillan's shrew


C. macmillani
Dollman, 1915
Ethiopia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[150]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [150]

Makwassie musk shrew


C. maquassiensis
Roberts, 1946
Southern Africa
 
Size: 5–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[151]

Habitat: Grassland and rocky areas[152]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [152]

Malayan shrew


C. malayana
Robinson & Kloss, 1911
Malaysia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[153]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [153]

Mamfe shrew


C. virgata
Sanderson, 1940
West-central Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[154]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [154]

Manenguba shrew


C. manengubae
Hutterer, 1982
West-central Africa
 
Size: 7–9 cm (3–4 in) long, plus 6–7 cm (2–3 in) tail[151]

Habitat: Forest, shrubland, and grassland[155]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [155]

Mauritanian shrew


C. lusitania
Dollman, 1915
Western and eastern Africa
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[156]

Habitat: Savanna and shrubland[157]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [157]

Mduma's shrew


C. mdumai
Stanley, Hutterer, Giarla, & Esselstyn, 2015
Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[158]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [158]

Mindanao shrew

 

C. beatus
Miller, 1910
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[159]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [159]

Mindoro shrew


C. mindorus
Miller, 1910
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[160]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [160]

Montane white-toothed shrew


C. montis
Thomas, 1906
Eastern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[161]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [161]

Moonshine shrew


C. luna
Dollman, 1910
Southeastern Africa
 
Size: 6–11 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[156]

Habitat: Forest[162]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [162]

Mossy forest shrew


C. musseri
Ruedi & Vogel, 1995
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[163]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [163]

Mount Nimba giant forest shrew


C. nimbasilvanus
Hutterer, 2003
Western Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[164]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [164]

Munissi's shrew


C. munissii
Stanley, Hutterer, Giarla, & Esselstyn, 2015
Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[165]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [165]

Narrow-headed shrew


C. stenocephala
Heim de Balsac & Dieterlen, 1979
Central Africa
 
Size: 8–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[166]

Habitat: Inland wetlands[167]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [167]

Negev shrew


C. ramona
Ivanitskaya, Shenbrot, & Nevo, 1996
Israel
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[168]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [168]

Negros shrew


C. negrina
Rabor, 1952
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[169]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [169]

Nicobar shrew


C. nicobarica
Miller, 1902
Great Nicobar Island in India
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[170]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown  [170]

Nigerian shrew


C. nigeriae
Dollman, 1915
West-central Africa
 
Size: 9–11 cm (4 in) long, plus 5–7 cm (2–3 in) tail[171]

Habitat: Forest[172]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [172]

Nimba shrew


C. nimbae
Heim de Balsac, 1956
Western Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[173]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [173]

Niobe's shrew


C. niobe
Thomas, 1906
Central Africa
 
Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[174]

Habitat: Forest and grassland[175]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [175]

North African white-toothed shrew

 

C. pachyura
Küster, 1835
Northern Africa and southern Europe
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[176]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [176]

Nyiro shrew


C. macowi
Dollman, 1915
Kenya
 
Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[177]

Habitat: Unknown[178]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [178]

Oriental shrew

 

C. orientalis
Jentink, 1890

Two subspecies
  • C. o. lawuana
  • C. o. orientalis
Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[179]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [179]

Palawan shrew


C. palawanensis
Taylor, 1934
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[180]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [180]

Pale gray shrew


C. pergrisea
Miller, 1913
Pakistan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[181]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [181]

Panay shrew


C. panayensis
Hutterer, 2007
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[182]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [182]

Peninsular shrew


C. negligens
Kloss, 1914
Southeastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[183]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [183]

Peters's musk shrew


C. gracilipes
Peters, 1870
Tanzania
 
Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[87]

Habitat: Unknown[184]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [184]

Phan Luong's shrew


C. phanluongi
Jenkins, Abramov, Rozhnov, & Olsson, 2010
Vietnam and Cambodia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[185]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [185]

Phu Hoc shrew


C. phuquocensis
Abramov, Jenkins, Rozhnov, & Kalinin, 2008
Phú Quốc island in Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[186]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [186]

Pitman's shrew


C. pitmani
Barclay, 1932
Zambia
 
Size: About 5 cm (2 in) long, plus 4 cm (2 in) tail[187]

Habitat: Unknown[188]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [188]

Polia's shrew


C. polia
Hollister, 1916
Democratic Republic of the Congo
 
Size: About 6 cm (2 in) long, plus 7 cm (3 in) tail[81]

Habitat: Unknown[189]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [189]

Rainey's shrew


C. raineyi
Heller, 1912
Kenya
 
Size: 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[190]

Habitat: Forest[191]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [191]

Reddish-gray musk shrew


C. cyanea
(Duvernoy, 1838)
Southern Africa
 
Size: 6–10 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[192]

Habitat: Forest, grassland, and caves[193]

Diet: Invertebrates[193]
 LC 


Unknown  [193]

Roosevelt's shrew


C. roosevelti
(Heller, 1910)
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[194]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [194]

Ryukyu shrew


C. orii
Kuroda, 1924
Southern Japan
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[195]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [195]

Sa Pa shrew

 

C. sapaensis
Jenkins, Abramov, Bannikova, & Rozhnov, 2013
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[196]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [196]

Saharan shrew


C. tarfayensis
Vesmanis & Vesmanis, 1980
Mauritania and Morocco
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas and desert[197]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [197]

Sahelian tiny shrew


C. pasha
Dollman, 1915
Scattered northern Africa
 
Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[198]

Habitat: Savanna[199]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [199]

Savanna dwarf shrew


C. nanilla
Thomas, 1909
Western and eastern Africa
 
Size: 4–6 cm (2–2 in) long, plus 3–5 cm (1–2 in) tail[200]

Habitat: Savanna[201]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [201]

Savanna path shrew


C. viaria
(Geoffroy, 1834)
Western, central, and eastern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[202]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [202]

Savanna shrew


C. fulvastra
(Sundevall, 1843)
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[203]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [203]

Savanna swamp shrew


C. longipes
Hutterer & Happold, 1983
West-central Africa
 
Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[204]

Habitat: Inland wetlands[205]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [205]

Siberian shrew


C. sibirica
Dukelsky, 1930
Central Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, grassland, inland wetlands, and desert[206]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [206]

Sibuyan shrew


C. ninoyi
Esselstyn & Goodman, 2010
Philippines
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[207]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [207]

Sicilian shrew

 

C. sicula
Miller, 1900

Four subspecies
  • C. s. aegatensis
  • C. s. calypso
  • C. s. esuae
  • C. s. sicula
Italy and Malta
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[208]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [208]

Sinharaja white-toothed shrew


C. hikmiya
Meegaskumbura, Meegaskumbura, Pethiyagoda, Manamendra-Arachchi, & Schneider, 2007
Sri Lanka
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[209]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [209]

Small-footed shrew


C. parvipes
Osgood, 1910
Central Africa
 
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[210]

Habitat: Forest, savanna, and shrubland[211]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [211]

Smoky white-toothed shrew


C. fumosa
Thomas, 1904
Kenya
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[212]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [212]

Sokolov's shrew


C. sokolovi
Jenkins, Abramov, Rozhnov, & Makarova, 2007
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[213]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [213]

Somali dwarf shrew


C. nana
Dobson, 1890
Ethiopia and Somalia
 
Size: About 4 cm (2 in) long, plus 3 cm (1 in) tail[214]

Habitat: Grassland[215]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [215]

Somali shrew


C. somalica
Thomas, 1895
Eastern Africa
 
Size: 6–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[166]

Habitat: Savanna[216]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [216]

Southeast Asian shrew

 

C. fuliginosa
(Blyth, 1856)
Southeastern Asian
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, and rocky areas[217]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [217]

Sri Lankan long-tailed shrew


C. miya
Phillips, 1929
Sri Lanka
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[218]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [218]

Sulawesi shrew


C. lea
Miller & Hollister, 1921
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[219]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [219]

Sulawesi tiny shrew


C. levicula
Miller & Hollister, 1921
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[220]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [220]

Sulawesi white-handed shrew


C. rhoditis
Miller & Hollister, 1921
Sulawesi island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[221]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [221]

Sumatran giant shrew


C. lepidura
Lyon, 1908
Sumatra island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[222]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [222]

Sumatran long-tailed shrew


C. paradoxura
Dobson, 1886
Sumatra island in Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[223]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [223]

Sunda shrew

 

C. monticola
Peters, 1870
Southeastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[224]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [224]

Swamp musk shrew


C. mariquensis
(Smith, 1844)

Three subspecies
  • C. m. mariquensis
  • C. m. neavei
  • C. m. shortridgei
Southern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[225]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [225]

São Tomé shrew

 

C. thomensis
(Bocage, 1887)
São Tomé Island in São Tomé and Príncipe
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[226]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [226]

Taiwanese gray shrew


C. tanakae
Kuroda, 1938
Eastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[227]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [227]

Tanzanian shrew


C. tansaniana
Hutterer, 1986
Tanzania
 
Size: 8–11 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[228]

Habitat: Forest[229]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown  [229]

Tarella shrew


C. tarella
Dollman, 1915
Central Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[230]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown  [230]

Telford's shrew


C. telfordi
Hutterer, 1986
Tanzania
 
Size: 10–11 cm (4 in) long, plus 7–9 cm (3–4 in) tail[231]

Habitat: Forest[232]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [232]

Thalia's shrew


C. thalia
Dippenaar, 1980
Ethiopia
 
Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[231]

Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[233]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [233]

Therese's shrew


C. theresae
Heim de Balsac, 1968
Western Africa
 
Size: 7–11 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[234]

Habitat: Forest and savanna[235]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [235]

Thick-tailed shrew


C. brunnea
Jentink, 1888

Two subspecies
  • C. b. brunnea
  • C. b. pudjonica
Indonesia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[236]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [236]

Timor shrew


C. tenuis
(Müller, 1840)
Timor
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[237]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [237]

Turbo shrew


C. turba
Dollman, 1910
Central Africa
 
Size: 9–11 cm (4 in) long, plus 5–6 cm (2–2 in) tail[238]

Habitat: Forest[239]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [239]

Ugandan lowland shrew


C. selina
(Dollman, 1915)
Kenya and Uganda
 
Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[240]

Habitat: Forest and inland wetlands[241]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [241]

Ugandan musk shrew


C. mutesae
Heller, 1910
Central Africa
 
Size: About 11 cm (4 in) long, plus 6 cm (2 in) tail[214]

Habitat: Forest[242]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [242]

Ultimate shrew


C. ultima
Dollman, 1915
Kenya
 
Size: About 9 cm (4 in) long, plus 6 cm (2 in) tail[243]

Habitat: Forest[244]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [244]

Upemba shrew


C. zimmeri
Osgood, 1936
Democratic Republic of the Congo
 
Size: 11–13 cm (4–5 in) long, plus 4–6 cm (2–2 in) tail[245]

Habitat: Unknown[246]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [246]

Usambara shrew


C. usambarae
Dippenaar, 1980
Tanzania
 
Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[247]

Habitat: Forest[248]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown  [248]

Ussuri white-toothed shrew

 

C. lasiura
(Dobson, 1890)
Eastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[249]

Diet: Insects and other invertebrates, as well as small vertebrates[249]
 LC 


Unknown  [249]

Voi shrew


C. voi
Osgood, 1910
Central and eastern Africa
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[250]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [250]

Voracious shrew

 

C. vorax
Allen, 1923
Southeastern Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[251]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [251]

West African long-tailed shrew


C. muricauda
(Miller, 1900)
Western Africa
 
Size: 5–7 cm (2–3 in) long, plus tail[252]

Habitat: Forest[253]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [253]

West African pygmy shrew


C. obscurior
Heim de Balsac, 1958
Western Africa
 
Size: 4–5 cm (2–2 in) long, plus 3–4 cm (1–2 in) tail[254]

Habitat: Forest[255]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [255]

Whitaker's shrew


C. whitakeri
De Winton, 1898
Northwestern Africa
 
Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[256]

Habitat: Coastal marine, intertidal marine, and shrubland[257]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [257]

Wimmer's shrew


C. wimmeri
Heim de Balsac & Aellen, 1958
Ivory Coast
 
Size: 8–9 cm (3–4 in) long, plus 5–6 cm (2–2 in) tail[258]

Habitat: Forest[259]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown  [259]

Xanthippe's shrew


C. xantippe
Osgood, 1910
Kenya and Tanzania
 
Size: 8–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[260]

Habitat: Savanna and shrubland[261]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [261]

Yankari shrew


C. yankariensis
Hutterer & Jenkins, 1980
Central and eastern Africa
 
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[260]

Habitat: Savanna[262]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [262]

Zaphir's shrew


C. zaphiri
Dollman, 1915
Ethiopia and Kenya
 
Size: About 10 cm (4 in) long, plus 6 cm (2 in) tail[263]

Habitat: Unknown[264]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown  [264]

Zarudny's rock shrew

 

C. zarudnyi
Ognew, 1928
Central Asia
 
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[265]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown  [265]

Genus Diplomesodon Brandt, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Piebald shrew

 

D. pulchellum
(Lichtenstein, 1823)
Central Asia
 
Size: 5–8 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[266]

Habitat: Desert[267]

Diet: Insects, as well as small lizards[267]
 LC 


Unknown  [267]

Genus Feroculus Kelaart, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Kelaart's long-clawed shrew


F. feroculus
(Kelaart, 1850)
Southern India and Sri Lanka
 
Size: 10–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[268]

Habitat: Forest, grassland, and inland wetlands[269]

Diet: Earthworms, small vertebrates, and plants[268]
 EN 


Unknown  [269]

Genus Paracrocidura Heim de Balsac, 1956 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Grauer's large-headed shrew


P. graueri
Hutterer, 1986
Democratic Republic of the Congo
 
Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270]

Habitat: Forest[271]

Diet: Insects[270]
 DD 


Unknown  [271]

Greater large-headed shrew


P. maxima
Heim de Balsac, 1959
Central Africa
 
Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270]

Habitat: Forest and inland wetlands[272]

Diet: Insects[270]
 LC 


Unknown  [272]

Lesser large-headed shrew


P. schoutedeni
Heim de Balsac, 1956
West-central Africa
 
Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[270]

Habitat: Forest[273]

Diet: Insects[270]
 LC 


Unknown  [273]

Genus Ruwenzorisorex Hutterer, 1986 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Ruwenzori shrew


R. suncoides
(Osgood, 1936)
Central Africa
 
Size: 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[274]

Habitat: Inland wetlands and forest[275]

Diet: Mollusks and small vertebrates[268]
 VU 


Unknown  [275]

Genus Scutisorex Thomas, 1910 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hero shrew

 

S. somereni
(Thomas, 1910)
Central Africa
 
Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[276]

Habitat: Forest[277]

Diet: Insects, earthworms, and small vertebrates[276]
 LC 


Unknown  [277]

Thor's hero shrew


S. thori
Stanley, Malekani, &Gambalemoke, 2013
Democratic Republic of the Congo Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[278]

Habitat: Forest[279]

Diet: Small animals and plants[278]
 DD 


Unknown  [279]

Genus Solisorex Thomas, 1924 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Pearson's long-clawed shrew


S. pearsoni
Thomas, 1924
Sri Lanka
 
Size: 12–15 cm (5–6 in) long, plus 5–7 cm (2–3 in) tail[266]

Habitat: Forest and grassland[280]

Diet: Insects, earthworms, and small vertebrates[266]
 EN 


Unknown  [280]

Genus Suncus Ehrenberg, 1832 – eighteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Anderson's shrew


S. stoliczkanus
(Anderson, 1877)
Southern Asia
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest and grassland[282]

Diet: Insects[281]
 LC 


Unknown  [282]

Asian highland shrew


S. montanus
(Kelaart, 1850)
Southern India and Sri Lanka
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest and grassland[283]

Diet: Insects[281]
 LC 


Unknown  [283]

Asian house shrew

 

S. murinus
(Linnaeus, 1766)
Asia and Africa (introduced in red)
 
Size: 10–5 cm (4–2 in) long, plus 5–9 cm (2–4 in) tail[284]

Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[285]

Diet: Insects and small mammals, as well as plants[286]
 LC 


Unknown  [285]

Black shrew


S. ater
(Medway, 1965)
Northern Borneo
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest[287]

Diet: Insects[281]
 DD 


Unknown  [287]

Bornean pygmy shrew


S. hosei
(Thomas, 1893)
Borneo
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest[288]

Diet: Insects[281]
 DD 


Unknown  [288]

Climbing shrew


S. megalura
(Jentink, 1888)
Sub-Saharan Africa
 
Size: 5–6 cm (2 in) long, plus 8–10 cm (3–4 in) tail[284]

Habitat: Forest and savanna[289]

Diet: Insects[284]
 LC 


Unknown  [289]

Day's shrew


S. dayi
(Dodson, 1888)
Southern India
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest and grassland[290]

Diet: Insects[281]
 EN 


Unknown  [290]

Etruscan shrew

 

S. etruscus
(Savi, 1822)
Europe, northern Africa, and Asia
 
Size: 3–5 cm (1–2 in) long, plus 2–3 cm (1 in) tail[1]

Habitat: Forest, shrubland, grassland, and desert[291]

Diet: Insects[292]
 LC 


Unknown  [291]

Flores shrew


S. mertensi
Kock, 1974
Flores island in Indonesia
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest[293]

Diet: Insects[281]
 EN 


Unknown  [293]

Greater dwarf shrew

 

S. lixus
(Thomas, 1898)
Southern Africa
 
Size: 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[281]

Habitat: Forest, savanna, shrubland, and grassland[294]

Diet: Insects[281]
 LC 


Unknown  [294]

Hutu-Tutsi dwarf shrew


S. hututsi
Peterhans & Hutterer, 2009
Central Africa Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[281]

Habitat: Forest and shrubland[295]

Diet: Insects[281]
 DD 


Unknown  [295]

Jungle shrew


S. zeylanicus
Phillips, 1928
Sri Lanka
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest[296]

Diet: Insects[281]
 EN 


Unknown  [296]

Least dwarf shrew


S. infinitesimus
(Heller, 1912)
South Africa and Kenya
 
Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[281]

Habitat: Forest and grassland[297]

Diet: Insects[281]
 LC 


Unknown  [297]

Lesser dwarf shrew


S. varilla
(Thomas, 1895)
Southern Africa
 
Size: 5–7 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[281]

Habitat: Forest, savanna, shrubland, and grassland[298]

Diet: Insects[281]
 LC 


Unknown  [298]

Malayan pygmy shrew


S. malayanus
(Kloss, 1917)
Malaysia and Thailand
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest[299]

Diet: Insects[281]
 LC 


Unknown  [299]

Remy's pygmy shrew


S. remyi
Brosset, Dubost, & Heim de Balsac, 1965
West-central Africa
 
Size: 4–5 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail[281]

Habitat: Forest[300]

Diet: Insects[281]
 LC 


Unknown  [300]

Sri Lankan shrew


S. fellowesgordoni
Phillips, 1932
Sri Lanka
 
Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[281]

Habitat: Forest and grassland[301]

Diet: Insects[281]
 EN 


Unknown  [301]

Taita shrew


S. aequatorius
(Heller, 1912)
Kenya
 
Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[302]

Habitat: Forest[303]

Diet: Insects[304]
 EN 


Unknown  [303]

Genus Sylvisorex Thomas, 1904 – fifteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bamenda pygmy shrew


S. silvanorum
(Hutterer, Riegert, & Sedláček, 2009)
Cameroon Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest and grassland[305]

Diet: Insects[306]
 DD 


Unknown  [305]

Bioko forest shrew


S. isabellae
Heim de Balsac, 1968
West-central Africa
 
Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest and grassland[307]

Diet: Insects[306]
 VU 


Unknown  [307]

Cameroonian forest shrew


S. camerunensis
Heim de Balsac, 1968
West-central Africa
 
Size: 5–6 cm (2 in) long, plus 5–7 cm (2–3 in) tail[308]

Habitat: Forest and grassland[309]

Diet: Insects[306]
 VU 


Unknown  [309]

Corbet's forest shrew


S. corbeti
Hutterer, 2009
West-central Africa Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest[310]

Diet: Insects[306]
 DD 


Unknown  [310]

Dudu Akaibe's pygmy shrew


S. akaibei
Mukinzi, Hutterer, & Barriere, 2009
Democratic Republic of the Congo Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest[311]

Diet: Insects[306]
 DD 


Unknown  [311]

Grant's forest shrew


S. granti
Thomas, 1907

Two subspecies
  • S. g. granti
  • S. g. mundus
Central Africa
 
Size: 4–7 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[312]

Habitat: Forest[313]

Diet: Insects[306]
 LC 


Unknown  [313]

Greater forest shrew


S. ollula
Thomas, 1913
Central Africa
 
Size: 5–8 cm (2–3 in) long, plus 1–2 cm (0–1 in) tail[314]

Habitat: Forest[315]

Diet: Insects[306]
 LC 


Unknown  [315]

Howell's forest shrew


S. howelli
Jenkins, 1984
Tanzania
 
Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[316]

Habitat: Forest[317]

Diet: Insects[306]
 LC 


Unknown  [317]

Johnston's forest shrew


S. johnstoni
(Dobson, 1888)
West-central and central Africa
 
Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[308]

Habitat: Forest[318]

Diet: Insects[306]
 LC 


Unknown  [318]

Kongana shrew


S. konganensis
Ray & Hutterer, 1996
Central Africa
 
Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest[319]

Diet: Insects[306]
 DD 


Unknown  [319]

Lesser forest shrew

 

S. oriundus
Hollister, 1916
Central Africa
 
Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[274]

Habitat: Forest[320]

Diet: Insects[306]
 DD 


Unknown  [320]

Moon forest shrew


S. lunaris
Thomas, 1906
Central Africa
 
Size: 7–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[321]

Habitat: Forest[322]

Diet: Insects[306]
 NT 


Unknown  [322]

Mount Cameroon forest shrew


S. morio
(Gray, 1862)
West-central Africa
 
Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2 in) tail[323]

Habitat: Forest[324]

Diet: Insects[306]
 EN 


Unknown  [324]

Rain forest shrew


S. pluvialis
Hutterer & Schlitter, 1996
West-central Africa
 
Size: 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[321]

Habitat: Forest[325]

Diet: Insects[306]
 DD 


Unknown  [325]

Volcano shrew


S. vulcanorum
Hutterer & Verheyan, 1985
Central Africa
 
Size: 4–6 cm (2 in) long, plus 4–7 cm (2–3 in) tail[308]

Habitat: Forest and inland wetlands[326]

Diet: Insects[306]
 LC 


Unknown  [326]

References

edit
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  64. ^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura douceti". IUCN Red List of Threatened Species. 2016: e.T40629A22295053. doi:10.2305/IUCN.UK.2016-3.RLTS.T40629A22295053.en.
  65. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura dsinezumi". IUCN Red List of Threatened Species. 2016: e.T40627A115176222. doi:10.2305/IUCN.UK.2016-3.RLTS.T40627A22294822.en.
  66. ^ Kingdon, p. 57
  67. ^ a b Dando, T. (2021). "Crocidura allex". IUCN Red List of Threatened Species. 2021: e.T5620A22301002. doi:10.2305/IUCN.UK.2021-2.RLTS.T5620A22301002.en.
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