Stock Recruitment Relationships

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STOCK-

RECRUITMENT
R E L AT I O N S H I P S

Ekwan Nofa Wiratno, S.Si., M.Si


[email protected]

Aquatic Resource Management Study Program -- Universitas Brawijaya


RECRUITMENT
• Annual recruitment is defined as the number of animals
“added to the population” each year.
• However, recruitment is also defined by when recruitment
occurs:
• at birth (mammals and birds);
• at age one (mammals and birds, some fish);
• at settlement (invertebrates / coral reef fishes);
• when it is first possible to detect animals using sampling gear; and
• when the animals enter the fishery.
• All of these definitions are “correct” but you need to be aware
which one is being used.
STOCK-RECRUITMENT
R E L AT I O N S H I P S
used to determine to what extent a population may be
• Stock harvested by either commercial or sport fisheries
• part of population under consideration
for actual or potential utilization; or
• sample of individuals with similar
production characteristics
• Recruitment
• the addition of new individuals to a
population through the birth process
• point in time (stage) after the egg
stage
RECRUITMEN
T Density-
R E L AT I O N S H I independent
PS Ricker
What determines
recruitment?
very few
Beverton-
recruits when
the spawning
Holt
stock is very
low

From: Wootton (1998). Ecology of teleost fishes.

Spawning stock biomass (SSB)


SOME HYPOTHESES FOR
D E N S I T Y- D E P E N D E N C E
• Habitat
• Some habitats lead to higher survival of offspring than others
(predators / food). Selection of habitat may be systematic (nest
selection) or random (location of settling individuals).
• Fecundity
• Animals are territorial – the total fecundity depends on getting a
territory.
• Feeding
• Given a fixed amount of food, sharing of food amongst
spawners will occur.
SELECTING AND FITTING
STOCK-RECRUITMENT
R E L AT I O N S H I P S
Skeena River sockeye

4,000

3,000
Recruites

2,000

1,000

0
0 500 1,000 1,500
Skeena river sockeye
Spawners
(Oncorhynchus nerka)
T H E B E V E R T O N - H O LT
R E L AT I O N S H I P
• The recruitment increases with the size of the spawning stock to a certain level,
then it flattens out. Further increase of the spawning stock does not lead to higher
recruitment.
• The survival rate of a cohort depends on the size of the cohort, i.e.:
dR Cohort is a
 (q  pR) R; R(0)  a S group
dt
of fish born in
R(t ) is the number of recruits at time t ,
the same year
S is the number (biomass) of spawners. within a
population or
• This can be integrated to give: stock.
a1 S S a3 S
R  
b1  S b2  a2 S 1  b3 S
T H E R I C K E R R E L AT I O N S H I P
• Recruitment increases with the size of the spawning stock to a
maximum, then recruitment decreases as the spawning stock increases.
• The survival rate of a cohort depends only on the initial abundance of
the cohort, i.e:
dR
 (q  pS ) R; R(0)  a S
dt
R(t ) is the number of recruits at time t ,
S is the number (biomass) of spawners.

• This can be integrated to give:


R  a1 S exp(b1S )  S exp(a 2  S / b2 )
THANK
YOU

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