Speciation Can Occur Rapidly or Slowly and Can Result From Changes in Few or Many Genes

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Speciation can occur rapidly or

slowly and can result from changes in


few or many genes.
 Ways to determine how long it takes for new
species to form:
• Broad patterns in fossil records
• Studies using morphological data (with fossils)
• Molecular data to asses time intervals between
speciation
 Punctuated equilibria: periods of apparent
punctuated sudden change in species across
geologic stratum.
• Change in species may be relatively quick and therefore
unable to be determined according to fossil records.
• This would be then attributed to punctuated equilibria
 Punctuated patterns suggest that in speciation,
once started, is a relatively quick process.
• Helianthus anomalus: a change in the hybrid fertility
changing from 5% to 90% in 4 generations
• Natural selection eliminated species that were not
compatible genetically with one another
 Total length of time between speciation events is the time
that elapses before populations of newly formed species start
to diverge from one another plus the time it takes for
speciation to be complete once divergence begins.
• Total time between speciation events varies (4,000 years to 6.5 million
years)

 Importance of this data:
• Suggests that it takes millions of years for before a newly formed
species will give rise to a new species
• Great variability shows there is not a “speciation clock” that regulates
speciation. Speciation is most likely at random
• Interruption of gene flow leads to diverging population until they
become reproductively isolated.
 Scientists study genes that control traits which may
lead to reproductive barriers.
 Japanese snails of genus Euhadra alleles of a single gene
controls the direction of the spiral of the shells, which changes
the orientation of the snails genitals, thus preventing mating
 Two species of Monkey flower, M. lewisii (Pink) and
M. Cardinalis (Red), are separated by pollinator’s
choice, which is affected by flower color. Bumblebees
tend to stop at the pink flowers and hummingbirds
tend to stop at the red flowers. A single gene controls
this trait and experiments were done by switching the
two genes in the flowers.
 Results: The pink flower had a 68 times more frequent visit by the
hummingbird. The red flower had a 74 times more frequent visit
from the bumblebee.
 Hybrid sterility between two species of
Drosophilia pseudoobscura results form gene
interaction between four loci.

 Postzygotic isolation in sunflower hybrid zone


is influenced by at least 26 chromosomal
segments.
 Speciation is the accumulation of small
physical changes which make different groups
of organism more and more different from
each other.

 Evolutionary changes are affected by both


extinction and speciation.

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