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Tulipa (tulips) is a genus of spring-

blooming perennial herbaceous bulbiferous geophytes, dying back after flowering
to an underground storage bulb. Depending on the species, tulip plants can be
between 10 and 70 cm (4 and 28 inches) high.
Flowers: The tulip's flowers are usually large and are actinomorphic (radially
symmetric) and hermaphrodite (contain both male (androecium) and female
(gynoecium) characteristics), generally erect, or more rarely pendulous, and are
arranged more usually as a single terminal flower, or when pluriflor as two to three
(e.g. Tulipa turkestanica), but up to four, flowers on the end of
a floriferous stem (scape), which is single arising from amongst the basal leaf
rosette. In structure, the flower is generally cup or star shaped. As with other
members of Liliaceae the perianth is undifferentiated (perigonium) and biseriate
(two whorled), formed from six free (i.e. apotepalous) caducous tepals arranged
into two separate whorls of three parts (trimerous) each. The two whorls represent
three petals and three sepals, but are termed tepals because they are nearly
identical. The tepals are usually petaloid (petal like), being brightly coloured, but
each whorl may be different, or have different coloured blotches at their bases,
forming darker colouration on the interior surface. The inner petals have a small,
delicate cleft at the top, while the sturdier outer ones form uninterrupted ovals. [3]
Androecium: The flowers have six distinct, basifixed introrse stamens arranged in
two whorls of three, which vary in length and may be glabrous or hairy. The
filaments are shorter than the tepals and dilated towards their base. [4]
Gynoecium: The style is short or absent and each stigma has three distinct lobes,
and the ovaries are superior, with three chambers.[4]
Fruit: The tulip's fruit is a globose or ellipsoid capsule with a leathery covering and
an ellipsoid to globe shape. Each capsule contains numerous flat, disc-
shaped seeds in two rows per chamber.[5] These light to dark brown seeds have
very thin seed coats and endosperm that does not normally fill the entire seed.[6][4]
Leaves: Tulip stems have few leaves. Larger species tend to have multiple leaves.
Plants typically have two to six leaves, some species up to 12. The tulip's leaf
is cauline (born on a stem), strap-shaped, with a waxy coating, and the leaves are
alternate (alternately arranged on the stem), diminishing in size the further up the
stem. These fleshy blades are often bluish-green in colour. [7][8][9]The bulbs are
truncated basally and elongated towards the apex. They are covered by a protective
tunic (tunicate) which can be glabrous or hairy inside.[4][9]
Colours[edit]
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The "Semper Augustus" was the most expensive tulip during the 17th-century tulip
mania. “The colour is white, with Carmine on a blue base, and with an unbroken
flame right to the top” – wrote Nicolas van Wassenaer in 1624 after seeing the
tulip in the garden of one Dr Adriaen Pauw, a director of the new East India
Company. With limited specimens in existence at the time and most owned by
Pauw, his refusal to sell any flowers, despite wildly escalating offers, is believed
by some to have sparked the mania.[3]
Tulip flowers come in a wide variety of colours, except pure blue (several tulips
with "blue" in the name have a faint violet hue), and have absent nectaries.[7][8][9]
[4]
Tulip flowers are generally bereft of scent and are the coolest of floral
characters. The Dutch regarded this lack of scent as a virtue, as it demonstrates the
flower's chasteness.[3]
While tulips can be bred to display a wide variety of colours, black tulips have
historically been difficult to achieve. The Queen of the Night tulip is as close to
black as a flower gets, though it is, in fact, a dark and glossy maroonish purple -
nonetheless, an effect prized by the Dutch. [3] The first truly black tulip was bred in
1986 by a Dutch flower grower in Bovenkarspel, Netherlands. The specimen was
created by cross-breeding two deep purple tulips, the Queen of the Night and
Wienerwald tulips.[10]
Phytochemistry[edit]
Tulipanin is an anthocyanin found in tulips. It is the 3-rutinoside of delphinidin.
The chemical compounds named tuliposides and tulipalins can also be found in
tulips and are responsible for allergies.[11] Tulipalin A, or α-methylene-γ-
butyrolactone, is a common allergen, generated by hydrolysis of the glucoside
tuliposide A. It induces a dermatitis that is mostly occupational and affects tulip
bulb sorters and florists who cut the stems and leaves. [12] Tulipanin A and B are
toxic to horses, cats and dogs.[13] The colour of a tulip is formed from two pigments
working in concert; a base colour that is always yellow or white, and a second laid-
on anthocyanin colour. The mix of these two hues determines the visible unitary
colour. The breaking of flowers occurs when a virus suppresses anthocyanin and
the base colour is exposed as a streak.[3]
Fragrance
The great majority of tulips, both species and cultivars, have no discernable scent,
but a few of both are scented to a degree, and Anna Pavord describes T.
Hungarica as "strongly scented",[14] and among cultivars, some such as "Monte
Carlo" and "Brown Sugar" are "scented", and "Creme Upstar" "fragrant".[15]
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Taxonomy
Main article: Taxonomy of Tulipa
Tulipa is a genus of the lily family, Liliaceae, once one of the largest families
of monocots, but which molecular phylogenetics has reduced to
a monophyletic grouping with only 15 genera. Within Liliaceae, Tulipa is placed
within Lilioideae, one of three subfamilies, with two tribes. Tribe Lilieae includes
seven other genera in addition to Tulipa.
Subdivision[edit]
The genus, which includes about 75 species, is divided into four subgenera.[9]

 Clusianae (4 species)
 Orithyia (4 species)
 Tulipa (52 species)
 Eriostemones (16 species)
Etymology[edit]
The word tulip, first mentioned in western Europe in or around 1554 and
seemingly derived from the "Turkish Letters" of diplomat Ogier Ghiselin de
Busbecq, first appeared in English as tulipa or tulipant, entering the language by
way of French: tulipe and its obsolete form tulipan or by way of Modern
Latin tulipa, from Ottoman Turkish tülbend ("muslin" or "gauze"), and may be
ultimately derived from the Persian: ‫ دلبند‬delband ("Turban"), this name being
applied because of a perceived resemblance of the shape of a tulip flower to that of
a turban.[16] This may have been due to a translation error in early times when it
was fashionable in the Ottoman Empire to wear tulips on turbans. The translator
possibly confused the flower for the turban.[9]
Ogier Ghiselin de Busbecq stated that the "Turks" used the word "tulipan" to
describe the flower. Extensive speculation has tried to understand why he would
state this, given that the Turkish word for tulip is "lale". It is from this speculation
that "tulipan" being a translation error referring to turbans is derived. This
Etymology has been challenged and makes no assumptions about possible errors.
At no point does Busbecq state this was the word used in Turkey, he simply states
it was used by the "Turks". On his way to Constantinople Busbecq states he
travelled through Hungary and used Hungarian guides. Until recent times "Turk"
was a common term when referring to Hungarians. The word "tulipan" is in fact
the Hungarian word for tulip. As long as one recognizes "Turk" as a reference to
Hungarians, no amount of speculation is required to reconcile the word's origin or
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form. Busbecq was simply repeating the word used by his "Turk/Hungarian"
guides.[17]
The Hungarian word "tulipan" may be adopted from an Indo-Aryan reference to
the tulip as a symbol of resurrection, "tala" meaning bottom or underworld and
"pAna" meaning defence.[17] Prior to arriving in Europe the Hungarians, and other
Finno-Ugrians, embraced the Indo-Iranian cult of the dead, Yima/Yama, and
would have been familiar with all of its symbols including the tulip. [18]

Distribution and habitat

Eastern end of the tulip range from Turkmenistan on the eastern shore of
the Caspian Sea to the Pamir-Alai and Tien-Shan mountains
Tulips are mainly distributed along a band corresponding to latitude 40° north,
from southeast of Europe (Greece, Albania, North Macedonia, Kosovo,
Southern Serbia, Bulgaria, most part of Romania, Ukraine, Russia) and Turkey in
the west, through the Levant (Syria, Israel, Palestinian Territories, Lebanon and
Jordan) and the Sinai Peninsula. From there it extends eastwards through Jerevan,
(Armenia) and Baku (Azerbaijan) and on the eastern shore of the Caspian
Sea through Turkmenistan, Bukhara, Samarkand and Tashkent (Uzbekistan), to the
eastern end of the range in the Pamir-Alai and Tien-Shan mountains in Central
Asia, which form the centre of diversity.[19] Further to the east, Tulipa is found in
the western Himalayas, southern Siberia, Inner Mongolia, and as far as the
northwest of China. While authorities have stated that no tulips west of the Balkans
are native,[20] subsequent identification of Tulipa sylvestris subsp. australis as a
native of the Iberian peninsula and adjacent North Africa shows that this may be a
simplification. In addition to these regions in the west tulips have been identified in
Greece, Cyprus and the Balkans. In the south, Iran marks its furthest extent, while
the northern limit is Ukraine. [21] Although tulips are also throughout most of the
Mediterranean and Europe, these regions do not form part of the natural
distribution. Tulips were brought to Europe by travellers and merchants
from Anatolia and Central Asia for cultivation, from where they escaped and
naturalised (see map). For instance, less than half of those species found in Turkey
are actually native.[20] These have been referred to as neo-tulipae.[22][23][9]
Tulips are indigenous to mountainous areas with temperate climates, where they
are a common element of steppe and winter-rain Mediterranean vegetation. They
thrive in climates with long, cool springs and dry summers. Tulips are most
commonly found in meadows, steppes and chaparral, but also introduced in fields,
orchards, roadsides and abandoned gardens.[22][9]
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Ecologya

Variegation produced by the tulip breaking virus

Botrytis tulipae is a major fungal disease affecting tulips, causing cell death and
eventually the rotting of the plant.[24] Other pathogens
include anthracnose, bacterial soft rot, blight caused by Sclerotium rolfsii,
bulb nematodes, other rots including blue molds, black molds and mushy rot.[25]
The fungus Trichoderma viride can infect tulips, producing dried leaf tips and
reduced growth, although symptoms are usually mild and only present on bulbs
growing in glasshouses.[citation needed]
Variegated tulips admired during the Dutch tulipomania gained their delicately
feathered patterns from an infection with the tulip breaking virus, a mosaic
virus that was carried by the green peach aphid, Myzus persicae. While the virus
produces fantastically streaked flowers, it also weakens plants and reduces the
number of offsets produced. Dutch growers would go to extraordinary lengths
during tulipomania to make tulips break, borrowing alchemists’ techniques and
resorting to sprinkling paint powders of the desired hue or pigeon droppings onto
flower roots.[3]
Tulips affected by the mosaic virus are called "broken"; while such plants can
occasionally revert to a plain or solid colouring, they will remain infected and have
to be destroyed. Today the virus is almost eradicated from tulip growers' fields.
The multicoloured patterns of modern varieties result from breeding; they normally
have solid, un-feathered borders between the colours.
Tulip growth is also dependent on temperature conditions. Slightly germinated
plants show greater growth if subjected to a period of cool dormancy, known
as vernalisation. Furthermore, although flower development is induced at warmer
temperatures (20–25 °C or 68–77 °F), elongation of the flower stalk and proper
flowering is dependent on an extended period of low temperature (< 10 °C or
50 °F).[26] Tulip bulbs imported to warm-winter areas are often planted in autumn
to be treated as annuals.
The colour of tulip flowers also varies with growing conditions.[27]