Here Be Dragons: The Ecology of Invasive Predators
Here Be Dragons: The Ecology of Invasive Predators
Here Be Dragons: The Ecology of Invasive Predators
2019
MCM/ICM
Summary Sheet
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Our model works well because it allows for the dragons to grow forever, but also limits their
growth rate based on environmental factors in a meaningful way that is representative of real
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ecological pressures. Not only will dragons need different amounts of land in different
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climates, but they will also grow to different sizes based on the scarcity of prey. Also, our
model allows for individual dragons to die off due to starvation which lets the remaining
dragons have a better chance at survival. This flexibility in the model leads to some
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interesting results, such as one or more dragons surviving in a certain area, but if the area is
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In a hot arid region, the dragons need 110,000 km2 to survive and become stable at 86,000 kg
each. In an arctic region, the dragons need 42,000 km2 and reach 3,500,000kg, and in a
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temperate region, they need only 570km2 and grow to 4,000,000 kg. As expected, the
dragons' ability to survive is directly correlated with the availability and scarcity of prey.
While this three-dragon scenario is not likely to happen on earth, the beauty of this model is
that it is dynamic enough to be highly generalizable. The hunting preference is robust enough
that it can be applied to any animal in any position on the food chain. Based on the habits of
any animal we can fit our model to imitate the hunting strategy it would take and apply it to
novel scenarios to learn more about its behavior.
By slightly modifying the model used, we showed that we were able to model the growth of
an invasive apex predator and see what will happen to the prey population given initial
conditions. This could be used as a precaution to stop harmful invasive species from entering
a sustainable and stable ecosystem.
Finally, using our analysis of dragons in different scenarios we recommended to George R.
R. Martin that he not send Daenerys and Drogon to fight the white walkers in the North, as
Drogon would not be able to meet his basal metabolic rate in such a sparse climate. We
informed George that in order for Drogon to survive he must always stay in warm temperate
regions where food is plentiful, and he can live sustainably within the ecosystem.
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Contents
1 INTRODUCTION 2
3 DRAGON CHARACTERISTICS 4
3.1 Unrestricted Dragon Growth . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
3.2 Metabolism and Kleibler’s Law . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
3.3 Dragon Are Gluten Intolerant . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
3.4 ODE Model . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
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4.4 Generalizing the Model . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
5 SENSITIVITY ANALYSIS 15
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6 WEAKNESSES AND FUTURE IMPROVEMENTS 17
6.1 Better Population Data . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
6.2 Dragon-Dragon and Prey-Prey Interactions . . . . . . . . . . . . . . . . . . . 17
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6.3 Scarcity Coefficient Ambiguity . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
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8 CONCLUSIONS 18
1 INTRODUCTION
We are required to analyze dragons’ characteristics, behavior, habits, diet and interaction with
their environment. Then build a mathematical model to analyze the ecological impacts the dragons
from Game of Thrones could introduce to our world. We decompose the problem into four sub-
problems:
• Base our model on Lotka–Volterra equations, modify them and apply to multi-prey situations.
In the first step, we seek to determine the scaling law of a dragon’s physique, using known data
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to estimate its size at maximum growth rate.
In the second step, we seek to use a 43 allometric scaling power law to evaluate metabolic rate
of dragons at given body masses. We will use a modified version of the scaling law to improve the
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accuracy additionally.
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In the third step, we aim to propose a biased hunting strategy that is related to relative size of
the prey, population density, and climate.
Finally, based on the already matured predator-prey model, we consider multiple species in
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different regions, justify the determination of several parameters, and put our model to the test.
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• Given infinite resources the dragon will continue to grow indefinitely as mentioned
in the problem’s description.
• All animals in a species will die of natural causes at the same age unless they are
eaten by a dragon. This is a simplification
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• In each species population there is a uniform spread of ages throughout the group.
If dogs live for ten years and there are one hundred dogs in a sample population, we assume
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there are ten dogs who are between zero and one years old, ten dogs who are between one and
two years old, etc. In view of the fact that this simplification doesn’t affect the results much.
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• Dragons do not have a preference for any type of animal meat over another. The
mass and population density of prey are the only parameters that matter in the dragons
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hunting strategy. We made this assumption as the TV show doesn’t present the dragons’
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• Drogon displays the maximum growth rate for a dragon in seasons 1 through 7
in Game of Thrones. The dragon has free access to roam as far as possible. We know
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keeping dragons in captivity stunts their growth, this does not affect Drogon. We see from
the complaints of farmers that Drogon has access to livestock, which is essentially unlimited
food that he doesn’t have to hunt for. Therefore, we believe that it is reasonable to assume
that Drogon grew as fast as a dragon could reasonably expect to grow.
• Dragons, while they have the ability to grow much larger than other animals,
basal metabolic rates scale with mass as other animals metabolic rates do. We
are assuming that dragons are in the kingdom of Animalia and hence obey allometric scaling
laws.
• Although reptiles are mostly ectotherms, dragons breath fire, so we assume they
are warm blooded and that their internal body temperature is similar to other mammals.
• In our models the only interaction is between dragons and their prey. There are
no dragon-dragon or prey-prey interactions. We made this assumption because dragons don’t
appear to practice cannibalism and we believe because of the size dragons can grow to, prey-
prey interactions are insignificant compared to dragon-prey interactions.
• The birth and death rates that we assume in our models encapsulates all natural
factors including: perdition, disease, aging, and availability of resources.
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• The current population of animals has already reached a steady state equilibrium.
This assumption follows from the fact that we are assuming there is no prey-prey interaction.
• The dragons are dropped into their environments today and are hatched as baby
dragons. This assumption enables us to present a more complete model.
• Each dragon has a preference to hunt something that is its own size rather than
something smaller or larger than itself. However, we assume a dragon can hunt some-
thing up to about 3 times as large as itself as a dragon can breathe fire. The reason that a
dragon would not prefer a larger prey is because there is a high energy cost to breathing fire.
• There is a change in the amount of food a dragon can eat in a day based on how
scarce food is. The amount this changes is dependent on the environment the dragon is
in. We made this assumption because the density of food has an inverse relationship with
hunting difficulty.
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• Each species is uniformly distributed throughout the area specified in each sim-
ulation. This is a simplification, in a better model the distribution would be based on the
herd behavior of the species
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• If the mass of a dragon drops to 60% of its previous maximum mass, the dragon
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dies of starvation. Humans will die if they drop to 60% of their normal mass. Since dragons
do not stop growing, we assume at any point their max mass was at once their ”normal” mass.
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• Dragons all grow at the same rate if they are in the same environment. This is a
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simplification, there would likely be some random variation due to the different activity levels
of the dragons and different genetic makeup.
• Dragons are roughly 90% efficient when they eat. 10% of ingested mass becomes waste.
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• The energy density of meat is 2500 kcal/kg for all animals This is a simplification, in
a perfect model we would find the energy density for all different types of meat.
3 DRAGON CHARACTERISTICS
3.1 Unrestricted Dragon Growth
Our first goal is to measure the growth rate of a dragon with infinite resources. It is stated in
the problem that dragons ”continue to grow throughout their life depending on the conditions and
amount of food available to them.” We know based on the show, however, that the rate of growth
is not unrestricted. After a year of growth we know that the dragons have gained between 20 to 30
kg even thought they certainly had access to ample food and were not held in captivity. In order
to find this growth rate we first gathered concrete data points on the size of Drogon as functions of
his age, the results of which are shown in table 1. [7]
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Age Height Wingspan Mass Standard Devi- Standard Devia- Standard Devi-
(in) (in) (kg) ation of Height tion of Wingspan ation of Mass
(in) (in) (kg)
0 9.5 - 10 0.71 - 0.5
1 16 - 35 0.5 - 7.07
1.5 64 240 - 11.31 6 -
6 - 2520.5 - - 243.95 -
As mentioned in the assumptions and justifications section, we are assuming that Drogon grew
at the maximum rate a dragon could grow. We believe this is reasonable since Drogon had unlimited
range to travel and he was able to eat from livestock, he was able to find as much food as he wanted
without having to hunt for it.
We were able to get an estimation of the height of Drogon at age six from his wingspan, we
estimated his height to be 672 in. We then had four data points for height as a function of age. We
know that mass is proportional to volume and volume is proportional to height cubed. Therefore,
we were able to find the relationship of mass, m, to time, t, given a maximal growth rate: m = ah3
By fitting to the data we had, we found a = 0.01134408.
In order to model the growth of Drogon for a long time, we needed a sample dragon that grew at
a maximal rate and grew older than Drogon is now. To do this we read through Game of Thrones
lore and found Balerion, to be the best candidate. Balerion was the Aegon the Conqueror’s Dragon
first, and was the dragon of the current king until he died at age 200. Since he was the dragon
of the king, it is reasonable to assume that he was able to eat as much as he wanted and achieve
maximal growth.
Given that he grows maximally we would like to get an estimation for his size at a known age.
To do this we follow the analysis by technology reporter, Robert Price [3], of a picture of Balerion
that has been approved by George R. R. Martin as being realistic. The picture is shown below in
Figure 2:
In season 7 of game of thrones we see that the skull of Balerion is kept under the red keep and
his skull is comparable to the size shown in this picture. In this picture we know that Aegon was
considered tall, so we estimate that he is 6 foot and 4 inches. Based on the picture we can estimate
that the height of Balerion is about 4 times the height of Aegon. We use a Komodo dragon as
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a model for length-to-height of a dragon, which we know is 10:1. Therefor we can estimate the
length of Balerion to be 76 meters long. Since Drogon is 61 meters long we can find the mass of
Balerion using the ratio of the lengths and the cubic mass to length ratio used previously and find
that Balerion was roughly 6,000,000 kg. This is a picture of Aegon on his conquest for Westeros,
therefore we know that at the time of this picture Balerion was 100 years old. Since we know that
100 years later, when Balerion died, he was roughly the same size as he was in that picture. This
means that at a certain point, although dragons may not ever stop growing, they do significantly
slow their rate of growth. To model this we decided a sigmoidal curve would best fit this data. As
it allows for the exponential growth we see at the beginning while also accounting for Balerion’s
obvious slow growth later in life.
This is a logistic function which will give the ”S-shaped” sigmoidal curve that we are looking
for:
L
f (t) = (1)
1 + e−k(t−t0 )
[4] We use the mass, height, and length relationships defined earlier to get mass as a function of
time. Once we had a function for mass as a function of time we did a χ2 test to find the constants,
L and k, that best fit the data. We found that L = 843.717523 and k = 0.71173635. We then
take the derivative to get change in mass as a function of age when a dragon has been growing at
its maximum rate. However, we don’t want our model to simply have the dragons grow at their
maximum rate at all times, so we need to do a change of variables to convert to an equation that
gives change in mass as a function of current mass. Doing so results in equation 2 with mass in
kilograms and time in days.
dmD 3 m
m0D = = mk(1 − ( 3 )1/3 ) (2)
dt 365 aL
climates: a warm temperate climate, an arid climate, and an arctic climate. For each of these
different climates we will provide a sample population based on data from The International Union
for Conservation of Nature, which is the worlds most comprehensive information source on the
global conservation status of animals. We will use this data to simulate the dragons growth in each
climate.
In order to simulate an accurate hunting strategy we came up with a preference model that
we will use to determine how likely the dragon is to go after a specific animal. As stated in the
assumptions we assume that a dragon has a preference to hunt something that is its own size rather
than something smaller or larger than itself. We constructed a probability density that the dragon
will hunt an animal based on the animal’s mass and it is shown below:
0.8
probability that the dragon will prefer that species
0.6
0.4
0.2
0.0
0.0 0.5 1.0 1.5 2.0 2.5 3.0 3.5
ratio of species mass available to the mass of each dragon
( 81 2
Ae− 8 (x−1) 0≤x≤1
p (ms mD ) = 9 2
Ae 8 (x−1) 1≤x
Where ms is the mass of an individual in the species, mD is the mass of the dragon, and A =
0.8976 which was found from normalizing the function. We used a piece-wise Gaussian distribution
for the dragon preference because in our assumptions we state that each species will be uniformly
distributed throughout the area. When the dragon is roaming around looking for the next animal
to hunt he will likely find something that is close to his size, assuming he has not yet outgrown all
animals. This probability density accounts for the unlikely, but possible, scenario that the dragon
cannot find something close to his size and will opt to hunt an animal that is smaller or larger than
himself. We made the right side of the distribution wider than the left side because dragons are
the most dangerous animal for a given size, so we think it is much more likely to punch above its
weight class and take down something that is bigger than itself than it is to waste it’s time chasing
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squirrels. Notice that as the dragon gets bigger the size of each animal matters less and less. This
exemplifies the idea that to a giant a small ant is the same as a large ant.
We then multiplied this preference distribution by the weighted sum Σmi ni where mi is the mass
of an individual in a species and ni is the population of that species. After summing over all species
this will give us the total prey mass in the system. We raise this to the power of a scarcity factor
that we will call sf . This scarcity factor encapsulates the difficulty to find food in the area given.
This difficulty could come from many factors, but given our assumptions about the distribution of
prey, the best method for determining this factor is basing it on the mass density, σm , of available
prey.
Knowing this helps, but it says nothing of the nature of the relationship between σm and sf . To
find this relationship, we looked at the plots of a wide range of both the areas in each region and
the values for sf . We found values for each region that exhibited the behavior we expected, feeding
when ample food is available, and struggling to feed when food is scarce. Using these values, we fit
a power law function to the data as shown in equation 4, where α was found to be 13.2948 and γ
was found to be -0.425.
γ
sf = ασm (4)
# " PN #sf
m0D
" !
mi np j=1 mij nj
me = PN PN + 2500B (7)
j=1 mij nj pij j=1 mij n0j 0.9
The resulting system of equations for the dragon and each prey species are as follows:
N
" #
dmD X
= mei − 2500B (8)
dt i=1
dnj mej
= nj (rbirthj − rdeathj ) − (9)
dt m ij
We used a python function that acts as a wrapper to the Fortran solver from ODEPACK to
integrate this system of differential equations. It switches automatically between the nonstiff Adams
method and the stiff BDF method.
As we can see, given these initial conditions the community reaches a steady state where the
deer, sheep, and New Zealand Fur Seal all survive while the Beef and Dairy Cattle are completely
eaten. In this simulation the three dragons together reach a peak of about 6.5×106 kg and stabalize
together at a total of about 4.4 × 106 kg.
Our next simulation is a bit more interesting. We restrict the dragons to an area of 570 km2 ,
only 30km2 less area than previously, but something drastically different happens.
0.8
2
species total mass (kg)
dragons remaining
0.6
0.4
1
0.2
0.0 0
1.0
Team #1924781 Page 11 of 21
The dragons still peak at around 6.5 × 106 kg, however this time, after about 11,000 days the
dragons have eaten so much of the sheep population that one dragon (Viserion) dies. Recall from
the assumptions that this happens when the dragons drops to 60% of its max mass. Now that
there is one less dragon eating sheep, the other two dragons actually start to gain weight back.
Unfortunately, at this point the sheep population is too small and they are not able to repopulate
fast enough, the dragons wipe out most of the sheep population, and they both starve on the same
day.
dragons remaining
male
20000
Rüppell’s Fox 100,000 3.2 2.1
15000 Honey Badger 10,000 9.5 1.0
Egyptian Mon- 10,000 2.6 2.2 1
10000 goose
Caracal 13,600 11.7934 2.8
5000 Barbary Sheep 5,000 120 0.9
In this simulation we see a very interesting phenomenon. Since this was such a low density
region
0 we had to give the dragons a lot of room in order for them not to immediately starve. 0 We
2
restricted0 the dragons to2000an area of 80,000
4000 km and let the 6000 system evolve.
8000 10000
time (days)
200000 2
species total mass (kg)
dragons remaining
150000
100000
1
50000
0
0
0 2000 4000 6000 8000 10000
time (days)
1.0
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We see that at about 800 days the dragons reach a collective peak at around 320,000 kg. We see
that almost all of their mass gain has come from the Barbary Sheep, which are by far the biggest
animals in this climate.at around day 1,450 we see that one dragon has died. Now that there are two
dragons left they are able to both grow with renewed speed, while also not draining the community
they are in of its resources. Notice that the only population that is affected by the dragons is the
Barbary Sheep population. This is because the Barbary Sheep is two orders of magnitude more
massive than all the other animals and has the smallest initial population out of any other animal.
This means that once the dragon becomes larger he will be more likely to go after the Barbary
sheep, and when he does, the effect it will have on the population of the sheep will be greater. The
two dragons now reach a new equilibrium and achieve a steady state with the community with a
combined mass of 200,000 kg.
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0
0 2000 4000 6000 8000 10000
time (days)
dragons remaining
2000000
1500000
1
1000000
500000
0
0
0 2000 4000 6000 8000 10000
time (days)
1.0
We see that we get to a collective mass maximum at about day 1,200. Then both the Alaska
Moose and Caribou population start to fall quickly. Soon the moose population begins to grow
0.8 returns to it’s initial population while the Caribou continues to fall. Around day 4,800 one
and
dragon dies and the two remaining dragon grow slightly and end up reaching a steady state with
the community with a mass of about 270,000 kg. What’s interesting about this simulation is that
no0.6animal was driven extinct, since there were so many moose and Caribou, and the likelihood the
dragon eats a certain species is proportional to the number of species in that population the other
animals were hardly touched by the dragon, leaving a completely stable community.
2
0.4We now complete our final simulation. We restrict our three dragons to an area of 400 km in
”Alaska” and let the simulation run. Observe that this is significantly less area than we allowed
our dragons originally in Alaska.
0.2
0.0
0.0 0.2 0.4 0.6 0.8 1.0
20000
Bison 2
125000
dragons remaining
Region: Arctic, Area: 800(km2)
40000 Dragon
160000 Alaska moose 3
Arctic ground squirrel 175
30000 Brown bear
140000 Caribou 1
Arctic fox 150
Eurasian lynx
20000 Wolf
120000 Wolverine
Mountain goat
Polar Bear 125
10000
Bison 2
100000
species total mass (kg)
we0.8will turn off the dragon death function. Instead of killing a dragon when it falls to 60% of its
0
maximum mass, we allow the dragons to get arbitrarily smaller than its maximum mass. Observe
0
0
the results: 2000 4000 6000 8000 10000
time (days)
0.6
dragons remaining
40000
0.0
0.0 0.2 0.4 0.6 0.8 1.0
30000
1
20000
10000
0
0
0 2000 4000 6000 8000 10000
time (days)
1.0
We get damped oscillatory behavior in the dragon mass by time function. We see that Caribou
0.8
Team #1924781 Page 15 of 21
reaches a population number such that they are able to increase their population as the dragon
oscillates, however, the moose population was not so lucky. In this simulation it is not quite clear
whether the moose population would eventually go extinct. However, since the dragon’s mass
function is decreasing quickly at the end of this simulation, it is safe to say that the Caribou will
remain at full population and the moose will not quite go extinct. It is unknown whether it will be
able to return to its former glory, further simulations would be needed to determine end behavior.
5 SENSITIVITY ANALYSIS
When we ran our tests we modified the behavior around our results slightly to see what would
happen, in the figures below we see that as one increases the land available to the dragons the
longer the dragons are able to survive until they reach a critical point where both the dragon and
prey populations are stable.
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Region: Arctic, Area: 24000(km2) Region: Arctic, Area: 29000(km2) Region: Arctic, Area: 26000(km2) Region: Arctic, Area: 20000(km2)
Dragon Dragon 3 Dragon Dragon
3500000 Alaska moose 3 4000000 Alaska moose Alaska moose 3 3000000 Alaska moose 3
Arctic ground squirrel Arctic ground squirrel 3500000 Arctic ground squirrel Arctic ground squirrel
Brown bear Brown bear Brown bear Brown bear
Caribou Caribou Caribou Caribou
3000000 Arctic fox 3500000 Arctic fox Arctic fox Arctic fox
Eurasian lynx Eurasian lynx Eurasian lynx 2500000 Eurasian lynx
3000000
Wolf Wolf Wolf Wolf
Wolverine 3000000 Wolverine Wolverine Wolverine
2500000 Mountain goat Mountain goat Mountain goat Mountain goat
Polar Bear 2 Polar Bear 2 2500000 Polar Bear 2 Polar Bear
Bison Bison Bison 2000000 Bison 2
2500000
dragons remaining
dragons remaining
dragons remaining
2000000
2000000
2000000 1500000
1500000
1500000
1500000
1 1 1000000
1 1
1000000 1000000
1000000
500000
500000 500000
500000
0 0 0 0 0 0
0 0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000
time (days) time (days) time (days) time (days)
Region: Arctic, Area: 21000(km2) Region: Arctic, Area: 27000(km2) Region: Arctic, Area: 30000(km2) Region: Arctic, Area: 28000(km2)
Dragon 4000000 Dragon Dragon 3 Dragon 3
3 3 4000000
Alaska moose Alaska moose Alaska moose Alaska moose
3000000 Arctic ground squirrel Arctic ground squirrel 4000000 Arctic ground squirrel Arctic ground squirrel
Brown bear 3500000 Brown bear Brown bear Brown bear
Caribou Caribou Caribou 3500000 Caribou
Arctic fox Arctic fox Arctic fox Arctic fox
Eurasian lynx Eurasian lynx Eurasian lynx Eurasian lynx
2500000 Wolf 3000000 Wolf Wolf Wolf
Wolverine Wolverine Wolverine 3000000 Wolverine
Mountain goat Mountain goat 3000000 Mountain goat Mountain goat
Polar Bear 2 Polar Bear 2 Polar Bear 2 Polar Bear 2
2000000 Bison 2500000 Bison Bison 2500000 Bison
species total mass (kg)
dragons remaining
dragons remaining
dragons remaining
2000000 2000000
1500000 2000000
1500000 1500000
1000000 1 1
1 1
1000000 1000000
1000000
500000
500000 500000
0 0 0 0 0 0
0 0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000
time (days) time (days) time (days) time (days)
Region: Arctic, Area: 22000(km2) Region: Arctic, Area: 25000(km2) Region: Arctic, Area: 23000(km2)
1.0
Dragon Dragon 3500000 Dragon
Alaska moose 3 Alaska moose 3 Alaska moose 3
Arctic ground squirrel 3500000 Arctic ground squirrel Arctic ground squirrel
3000000 Brown bear Brown bear Brown bear
Caribou Caribou 3000000 Caribou
Arctic fox Arctic fox Arctic fox
Eurasian lynx 3000000 Eurasian lynx Eurasian lynx 0.8
Wolf Wolf Wolf
2500000 Wolverine Wolverine Wolverine
2500000
Mountain goat Mountain goat Mountain goat
Polar Bear 2500000 Polar Bear Polar Bear
2 2 2
Bison Bison Bison
2000000 0.6
species total mass (kg)
dragons remaining
dragons remaining
2000000
1500000 1500000
1500000
0.4
1 1 1
1000000 1000000
1000000
500000 0.2
500000 500000
0 0 0 0 0 0
0.0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0.0 0.2 0.4 0.6 0.8 1.0
time (days) time (days) time (days)
This simulated graphs ranging from areas of 20,000 km2 to 30,000 km2 in 1,000 km2 increments.
(Note: these graphs are not in order) In this simulation we see that at 27,000 km2 the dragons die
and at 28,000 km2 we reach a stable community. We wished to further investigate the behavior or
the model in between those ranges so we refined our search.
Region: Arctic, Area: 27400(km2) Region: Arctic, Area: 27900(km2) Region: Arctic, Area: 27600(km2) Region: Arctic, Area: 27000(km2)
4000000 Dragon Dragon 3 4000000 Dragon 4000000 Dragon
3 4000000 3 3
Alaska moose Alaska moose Alaska moose Alaska moose
Arctic ground squirrel Arctic ground squirrel Arctic ground squirrel Arctic ground squirrel
3500000 Brown bear Brown bear Brown bear 3500000 Brown bear
Caribou 3500000 Caribou 3500000 Caribou Caribou
Arctic fox Arctic fox Arctic fox Arctic fox
Eurasian lynx Eurasian lynx Eurasian lynx Eurasian lynx
3000000 Wolf Wolf 3000000 Wolf 3000000 Wolf
Wolverine 3000000 Wolverine Wolverine Wolverine
Mountain goat Mountain goat Mountain goat Mountain goat
Polar Bear 2 Polar Bear 2 Polar Bear 2 Polar Bear 2
2500000 Bison 2500000 Bison 2500000 Bison 2500000 Bison
species total mass (kg)
dragons remaining
dragons remaining
dragons remaining
2000000 2000000 2000000 2000000
0 0 0 0 0 0 0
0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000
time (days) time (days) time (days) time (days)
Region: Arctic, Area: 27100(km2) Region: Arctic, Area: 27700(km2) Region: Arctic, Area: 28000(km2) Region: Arctic, Area: 27800(km2)
4000000 Dragon 3 4000000 Dragon Dragon 3 Dragon
3 4000000 4000000 3
Alaska moose Alaska moose Alaska moose Alaska moose
Arctic ground squirrel Arctic ground squirrel Arctic ground squirrel Arctic ground squirrel
3500000 Brown bear Brown bear Brown bear Brown bear
Caribou 3500000 Caribou 3500000 Caribou 3500000 Caribou
Arctic fox Arctic fox Arctic fox Arctic fox
Eurasian lynx Eurasian lynx Eurasian lynx Eurasian lynx
3000000 Wolf 3000000 Wolf Wolf Wolf
Wolverine Wolverine 3000000 Wolverine 3000000 Wolverine
Mountain goat Mountain goat Mountain goat Mountain goat
Polar Bear 2 Polar Bear 2 Polar Bear 2 Polar Bear 2
2500000 Bison 2500000 Bison 2500000 Bison 2500000 Bison
species total mass (kg)
dragons remaining
dragons remaining
dragons remaining
2000000 2000000 2000000 2000000
0 0 0 0 0 0
0 0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000
time (days) time (days) time (days) time (days)
Region: Arctic, Area: 27200(km2) Region: Arctic, Area: 27500(km2) Region: Arctic, Area: 27300(km2)
4000000 3 3 1.0
Dragon 4000000 Dragon 4000000 Dragon
Alaska moose Alaska moose 3 Alaska moose
Arctic ground squirrel Arctic ground squirrel Arctic ground squirrel
3500000 Brown bear Brown bear 3500000 Brown bear
Caribou 3500000 Caribou Caribou
Arctic fox Arctic fox Arctic fox
Eurasian lynx Eurasian lynx Eurasian lynx 0.8
3000000 Wolf 3000000 Wolf 3000000 Wolf
Wolverine Wolverine Wolverine
Mountain goat Mountain goat Mountain goat
Polar Bear 2 Polar Bear 2 Polar Bear 2
2500000 Bison 2500000 Bison 2500000 Bison
0.6
species total mass (kg)
dragons remaining
dragons remaining
0.2
500000 500000 500000
0 0 0 0
0 0 0.0
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 0.0 0.2 0.4 0.6 0.8 1.0
time (days) time (days) time (days)
This simulated graphs ranging from areas of 27,000 km2 to 28,000 km2 in 100 km2 increments.
From simulation we are able to see more finely where the transition from sustainable to unsustain-
able. In this simulation 27,100 km2 is the area needed to sustain three dragons in alaska.
Team #1924781 Page 17 of 21
least 500m2 .
Figure 3: Mismatch between energy supply and demand. Blue line denotes energy supply, orange
line denotes energy demand.
8 CONCLUSIONS
We were tasked with analyzing the dragons from Game of Thrones to find the characteristics,
behavior, habits, diets, and interaction of the dragons with their environment if they were in our
world today. To do this we gathered size and mass data from the book and fit a logistic growth
curve to the data. We then derived an equation for the maximum mass growth rate of a dragon.
We simulated three dragons growing in three different climates: a hot arid region, an arctic region,
and a temperate region.
Using species population data from the IUCN Red List for each region, we built a profile of
the most common animals in each, their population density, mass, and birth/death rates. Using
this data and the mass growth rate we fit to the dragons, we modeled the interactions between the
dragons and the different environments.
We ran many simulations for each environment and found the area needed to support all three
dragons, as well as other area sizes that exhibit interesting patterns or may support only one or two
dragons. In a hot arid region, (western Sahara desert), the dragons need 110,000 km2 to survive
and become stable at 86,000 kg each. In an arctic region (Alaska), the dragons need 42,000 km2
and reach 3,500,000 kg, and in a temperate region (New Zealand), they need only 570 km2 and
grow to 4,000,000 kg. As expected, the dragons’ ability to survive is directly correlated with the
availability and scarcity of prey.
Team #1924781 Page 19 of 21
First off we’d like to say that it is an honor to have the opportunity to give you our input,
and although we may criticize some aspects of the books for its questionable and often estranged
relationship with physics; it does not mean that we think any less of your work or don’t appreciate
it for what it is: One of the best fantasy series of our time. Now onto our guidance:
Under the current laws you have set, there are about to be dragon deaths on your hands
depending on how you handle the upcoming season and whether you take our advice or not. While
we do not know the exact animal makeup of the Game of Thrones universe, we have no reason to
believe that it is much different from ours in terms of basic animal population, besides the dragons
of course. While there may be the rare dire-wolf sighting, or a more giants than we usually see in
our world, the baseline ecosystems seem comparable. We have built a model that simulates the
growth of dragons and how they interact with local ecosystems as they grow. We studied three
different climates: arid, temperate, and arctic, and after our analysis we see an alarming situation
about to unfold. The biggest problem that will face Drogon in the next season is not the Night
King and his ice dragon, it is migration and starvation.
The problem is that for the past year Drogon has lived around Dragonstone with unlimited space
to travel, eat, and grow. Because of this, as seen in the show, he was able to grow incredibly quickly
and gain an astounding amount of mass. While this extra mass is intimidating to all his foes, it
requires a massive amount of energy to keep running. Based off our calculations we were able to
estimate Drogon’s mass during season 7 at 3,444,574 kg. With some help with our friends at Nature,
we are then able to estimate Drogon’s basal metabolic rate and in turn find the amount of food
Drogon must eat every day to stay alive. Drogon must eat an astounding 5,392 kg per day in order
to remain at his same size, and this doesn’t even account for all the energy he would expand walking,
flying, and breathing fire. Now, in his current climate this level of food take, while adventurous, is
possible. Based on a simulation we ran using New Zealand as a substitute for the ecosystem around
Dragonstone we found that Drogon could feed himself indefinitely at Dragonstone, the community
is able to adapt and reaches a new steady state relationship with Drogon as the apex predator.
While Drogon can sustain himself in a temperate climate, if he moves to any other climate
he will quickly starve. Using Alaska as a model for the North in Game of Thrones we were able
to simulate the growth of Drogon in an arctic environment, and the results are not promising for
Drogon. If we put Drogon in the north starting as a baby, there is still a chance he may not survive.
It is possible if Drogon is not careful that he over eats. Since the population in arctic region is
much more sparse than in that of a temperate region; it is very easy for Drogon to eat so much that
his basal metabolic rate exceeds the rate at which animals in that region are able to reproduce.
In this scenario Drogon eats as much as he can for as long as he can, but eventually he eats all
the animals up and dies quickly behind them. While when doing our simulations this was not an
unlikely scenario, there were ways to avoid this. If Drogon does not grow as quickly as possible at
first, he is able to more sustainably integrate himself into the arctic climate.
However that is not the situation Drogon has found himself in. When we last left last season
Daenerys was planning on taking Drogon deep into the North to fight the Night King and his army
of the dead. This would be a fatal mistake. Drogon has far outgrown the size that he would be
sustainable in the North. Even if he spent all his time hunting the most calorie dense food in
the North, he would not be able to gather enough food. He would quickly eat through the entire
animal population of the North and then starve. On top of that the massive amounts of energy
Team #1924781 Page 20 of 21
it would take for Drogon to fight would make his uninevitable death approach much more quickly.
The nights watch nor the wildlings would be able to spare that much food to save Drogon, they
can barely feed themselves at the moment. Unfortunately, since the Dothraki Sea is surrounded by
an arid region, which is even more sparsely populated than an arctic region, Drogon will never be
able to return to his birthplace.
For better or worse, Drogon can never leave his temperate climate for more than a few days.
What he has gained in size he has lost in mobility. We would advise Daenerys to stay at Dragonstone
and wait for the Night King and his army of the dead to come to them, otherwise Drogon will starve,
for winter is coming.
Sincerely,
Team #1924781
Team #1924781 Page 21 of 21
References
[1] Feces | biology | Britannica.com.
[3] The illustrated guide to the best dragons in fantasy, April 2014.
[5] Erica Gonzales. The Dragons in ’Game of Thrones’ Season 7 Are Literally the Size of Airplanes,
March 2017.
[6] Tom Kolokotrones, Van Savage, Eric J. Deeds, and Walter Fontana. Curvature in metabolic
scaling. Nature, 464(7289):753–756, April 2010.
[7] George R. R. Martin. A clash of kings: a song of fire and ice [book 2]. Martin, George R. R.
bk. 2. Song of ice and fire ;. Bantam Books, New York, 1999.
[8] Geoffrey B. West. Scale: the universal laws of growth, innovation, sustainability, and the pace
of life in organisms, cities, economies, and companies. Penguin Press, New York, 2017.