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History of Earth

The history of Earth concerns the


development of planet Earth from
its formation to the present
day.[1][2] Nearly all branches of
natural science have contributed to
understanding of the main events of
Earth's past, characterized by
constant geological change and
biological evolution.

The geological time scale (GTS), as


defined by international
convention,[3] depicts the large
spans of time from the beginning of
the Earth to the present, and its
divisions chronicle some definitive
events of Earth history. (In the
graphic: Ga means "billion years
ago"; Ma, "million years ago".)
Earth formed around 4.54 billion
years ago, approximately one-third
the age of the universe, by accretion
from the solar nebula.[4][5][6]
Volcanic outgassing probably
created the primordial atmosphere and then the ocean, but the early atmosphere contained almost no oxygen. Much of the Earth was
molten because of frequent collisions with other bodies which led to extreme volcanism. While the Earth was in its earliest stage
(Early Earth), a giant impact collision with a planet-sized body named Theia is thought to have formed the Moon. Over time, the
Earth cooled, causing the formation of a solidcrust, and allowing liquid water on the surface.

The Hadean eon represents the time before a reliable (fossil) record of life; it began with the formation of the planet and ended 4.0
billion years ago. The following Archean and Proterozoic eons produced the beginnings of life on Earth and its earliest evolution.
The succeeding eon is the Phanerozoic, divided into three eras: the Palaeozoic, an era of arthropods, fishes, and the first life on land;
the Mesozoic, which spanned the rise, reign, and climactic extinction of the non-avian dinosaurs; and the Cenozoic, which saw the
rise of mammals. Recognizable humans emerged at most 2 million years ago, a vanishingly small period on the geological scale.

The earliest undisputed evidence of life on Earth dates at least from 3.5 billion years ago,[7][8][9] during the Eoarchean Era, after a
geological crust started to solidify following the earlier molten Hadean Eon. There are microbial mat fossils such as stromatolites
found in 3.48 billion-year-old sandstone discovered in Western Australia.[10][11][12] Other early physical evidence of a biogenic
substance is graphite in 3.7 billion-year-old metasedimentary rocks discovered in southwestern Greenland[13] as well as "remains of
biotic life" found in 4.1 billion-year-old rocks in Western Australia.[14][15] According to one of the researchers, "If life arose
relatively quickly on Earth … then it could be common in theuniverse."[14]

Photosynthetic organisms appeared between 3.2 and 2.4 billion years ago and began enriching the atmosphere with oxygen. Life
remained mostly small and microscopic until about 580 million years ago, when complex multicellular life arose, developed over
time, and culminated in the Cambrian Explosion about 541 million years ago. This sudden diversification of life forms produced
most of the major phyla known today, and divided the Proterozoic Eon from the Cambrian Period of the Paleozoic Era. It is estimated
that 99 percent of all species that ever lived on Earth, over five billion,[16] have gone extinct.[17][18] Estimates on the number of
Earth's current species range from 10 million to 14 million,[19] of which about 1.2 million are documented, but over 86 percent have
not been described.[20] However, it was recently claimed that 1 trillion species currently live on Earth, with only one-thousandth of
one percent described.[21]

The Earth's crust has constantly changed since its formation, as has life has since its first appearance. Species continue to evolve,
taking on new forms, splitting into daughter species, or going extinct in the face of ever
-changing physical environments. The process
of plate tectonics continues to shape the Earth's continents and oceans and the life they harbor. Human activity is now a dominant
force affecting global change, harming the biosphere, the Earth's surface, hydrosphere, and atmosphere with the loss of wild lands,
over-exploitation of the oceans, production of greenhouse gases, degradation of the ozone layer, and general degradation of soil, air,
and water quality.

Contents
Eons
Geologic time scale
Solar System formation
Hadean and Archean Eons
Formation of the Moon
First continents
Oceans and atmosphere
Origin of life
Proterozoic Eon
Oxygen revolution
Snowball Earth
Emergence of eukaryotes
Supercontinents in the Proterozoic
Late Proterozoic climate and life
Phanerozoic Eon
Tectonics, paleogeography and climate
Cambrian explosion
Colonization of land
Evolution of tetrapods
Extinctions
Diversification of mammals
Human evolution
See also
Notes
References
Further reading
External links

Eons
In geochronology, time is generally measured in mya (megayears or million years), each unit representing the period of
approximately 1,000,000 years in the past. The history of Earth is divided into four great eons, starting 4,540 mya with the formation
of the planet. Each eon saw the most significant changes in Earth's composition, climate and life. Each eon is subsequently divided
into eras, which in turn are divided intoperiods, which are further divided intoepochs.
Time
Eon Description
(mya)
The Earth is formed out of debris around the solarprotoplanetary disk. There is no life.
4,540– Temperatures are extremely hot, with frequent volcanic activity and hellish environments.
Hadean
4,000 The atmosphere is nebular. Possible early oceans or bodies of liquid water. The moon is
formed around this time, probably due to aprotoplanet's collision into Earth.
Prokaryote life, the first form of life, emerges at the very beginning of this eon, in a process
4,000–
Archean known as abiogenesis. The continents of Ur, Vaalbara and Kenorland may have been
2,500
formed around this time. The atmosphere is composed of volcanic and greenhouse gases.
Eukaryotes, a more complex form of life, emerge, including some forms ofmulticellular
organisms. Bacteria begin producing oxygen, shaping the third and current of Earth's
2,500– atmospheres. Plants, later animals and possibly earlier forms of fungi form around this time.
Proterozoic
541 The early and late phases of this eon may have undergoneSnowball
" Earth" periods, in
which all of the planet suffered below-zero temperatures. The early continents ofColumbia,
Rodinia and Pannotia may have formed around this time, in that order .
Complex life, including vertebrates, begin to dominate the Earth's ocean in a process
known as the Cambrian explosion. Pangaea forms and later dissolves intoLaurasia and
Gondwana. Gradually, life expands to land and all familiar forms of plants, animals and
541–
Phanerozoic fungi begin appearing, including annelids, insects and reptiles. Severalmass extinctions
present
occur, among which birds, the descendants of dinosaurs, and more recently mammals
emerge. Modern animals—including humans—evolve at the most recent phases of this
eon.

Geologic time scale


The history of the Earth can be organized chronologically according to the geologic time scale, which is split into intervals based on
stratigraphic analysis.[2][22] The following four timelines show the geologic time scale. The first shows the entire time from the
formation of the Earth to the present, but this gives little space for the most recent eon. Therefore, the second timeline shows an
expanded view of the most recent eon. In a similar way, the most recent era is expanded in the third timeline, and the most recent
period is expanded in the fourth timeline.

Millions of Years
Solar System formation
The standard model for the formation of the Solar System
(including the Earth) is the solar nebula hypothesis.[23] In
this model, the Solar System formed from a large, rotating
cloud of interstellar dust and gas called the solar nebula. It
was composed of hydrogen and helium created shortly after
the Big Bang 13.8 Ga (billion years ago) and heavier
elements ejected by supernovae. About 4.5 Ga, the nebula
began a contraction that may have been triggered by the
shock wave from a nearby supernova.[24] A shock wave
would have also made the nebula rotate. As the cloud began
to accelerate, its angular momentum, gravity, and inertia An artist's rendering of aprotoplanetary disk

flattened it into a protoplanetary disk perpendicular to its


axis of rotation. Small perturbations due to collisions and
the angular momentum of other large debris created the means by which kilometer-sized protoplanets began to form, orbiting the
nebular center.[25]

The center of the nebula, not having much angular momentum, collapsed rapidly, the compression heating it until nuclear fusion of
hydrogen into helium began. After more contraction, a T Tauri star ignited and evolved into the Sun. Meanwhile, in the outer part of
the nebula gravity caused matter to condense around density perturbations and dust particles, and the rest of the protoplanetary disk
began separating into rings. In a process known as runaway accretion, successively larger fragments of dust and debris clumped
together to form planets.[25] Earth formed in this manner about 4.54 billion years ago (with an uncertainty of 1%)[26][27][4][28] and
was largely completed within 10–20 million years.[29] The solar wind of the newly formed T Tauri star cleared out most of the
material in the disk that had not already condensed into larger bodies. The same process is expected to produce accretion disks
planets.[30]
around virtually all newly forming stars in the universe, some of which yield

The proto-Earth grew by accretion until its interior was hot enough to melt the heavy, siderophile metals. Having higher densities
than the silicates, these metals sank. This so-called iron catastrophe resulted in the separation of a primitive mantle and a (metallic)
core only 10 million years after the Earth began to form, producing the layered structure of Earth and setting up the formation of
Earth's magnetic field.[31] J.A. Jacobs [32] was the first to suggest that the inner core—a solid center distinct from the liquid outer
core—is freezing and growing out of the liquid outer core due to the gradual cooling of Earth's interior (about 100 degrees Celsius
per billion years[33] ).

Hadean and Archean Eons


The first eon in Earth's history, the Hadean, begins with the Earth's formation and is followed by the Archean eon at 3.8 Ga.[2]:145
The oldest rocks found on Earth date to about 4.0 Ga, and the oldest detrital zircon crystals in rocks to about 4.4 Ga,[34][35][36] soon
after the formation of the Earth's crust and the Earth itself. The giant impact hypothesis for the Moon's formation states that shortly
after formation of an initial crust, the proto-Earth was impacted by a smaller protoplanet, which ejected part of the mantle and crust
into space and created the Moon.[37][38][39]

From crater counts on other celestial bodies, it is inferred that a period of intense meteorite impacts, called the Late Heavy
[40] In addition, volcanism was severe due
Bombardment, began about 4.1 Ga, and concluded around 3.8 Ga, at the end of the Hadean.
to the large heat flow and geothermal gradient.[41] Nevertheless, detrital zircon crystals dated to 4.4 Ga show evidence of having
[34]
undergone contact with liquid water, suggesting that the Earth already had oceans or seas at that time.

By the beginning of the Archean, the Earth had cooled significantly. Present life forms could not have survived at Earth's surface,
because the Archean atmosphere lacked oxygen hence had no ozone layer to block ultraviolet light. Nevertheless, it is believed that
primordial life began to evolve by the early Archean, with candidate fossils dated to around 3.5 Ga.[42] Some scientists even
speculate that life could have begun during the early Hadean, as far back as 4.4 Ga,
surviving the possible Late Heavy Bombardment period in hydrothermal vents
below the Earth's surface.[43]

Formation of the Moon


Earth's only natural satellite, the Moon, is larger relative to its planet than any other
satellite in the solar system.[nb 1] During the Apollo program, rocks from the Moon's
surface were brought to Earth. Radiometric dating of these rocks shows that the
Moon is 4.53 ± 0.01 billion years old,[46] formed at least 30 million years after the
solar system.[47] New evidence suggests the Moon formed even later,
4.48 ± 0.02 Ga, or 70–110 million years after thestart of the Solar System.[48] Artist's conception of Hadean Eon
Earth, when it was much hotter and
Theories for the formation of the Moon must explain its late formation as well as the
inhospitable to all forms of life.
following facts. First, the Moon has a low density (3.3 times that of water, compared
to 5.5 for the earth[49] ) and a small metallic core. Second, there is virtually no water
or other volatiles on the moon. Third, the Earth and Moon have the same oxygen
isotopic signature (relative abundance of the oxygen isotopes). Of the theories
proposed to account for these phenomena, one is widely accepted: The giant impact
hypothesis proposes that the Moon originated after a body the size of Mars
(sometimes named Theia[47] ) struck the proto-Earth a glancing blow.[1]:256[50][51]

The collision released about 100 million times more energy than the more recent
Chicxulub impact that is believed to have caused the extinction of the dinosaurs. It
was enough to vaporize some of the Earth's outer layers and melt both
bodies.[50][1]:256 A portion of the mantle material was ejected into orbit around the Artist's impression of the enormous
Earth. The giant impact hypothesis predicts that the Moon was depleted of metallic collision that probably formed the
material,[52] explaining its abnormal composition.[53] The ejecta in orbit around the Moon
Earth could have condensed into a single body within a couple of weeks. Under the
influence of its own gravity, the ejected material became a more spherical body: the
Moon.[54]

First continents
Mantle convection, the process that drives plate tectonics, is a result of heat flow
from the Earth's interior to the Earth's surface.[55]:2 It involves the creation of rigid
tectonic plates at mid-oceanic ridges. These plates are destroyed by subduction into
the mantle at subduction zones. During the early Archean (about 3.0 Ga) the mantle
was much hotter than today, probably around 1,600 °C (2,910 °F),[56]:82 so
convection in the mantle was faster. Although a process similar to present-day plate
tectonics did occur, this would have gone faster too. It is likely that during the
Hadean and Archean, subduction zones were more common, and therefore tectonic
plates were smaller.[1]:258[57]

The initial crust, formed when the Earth's surface first solidified, totally disappeared Geologic map of North America,
from a combination of this fast Hadean plate tectonics and the intense impacts of the color-coded by age. The reds and
Late Heavy Bombardment. However, it is thought that it was basaltic in pinks indicate rock from theArchean.
composition, like today's oceanic crust, because little crustal differentiation had yet
taken place.[1]:258 The first larger pieces of continental crust, which is a product of
differentiation of lighter elements during partial melting in the lower crust, appeared at the end of the Hadean, about 4.0 Ga. What is
left of these first small continents are called cratons. These pieces of late Hadean and early Archean crust form the cores around
which today's continents grew.[58]

The oldest rocks on Earth are found in theNorth American cratonof Canada. They are tonalites from about 4.0 Ga. They show traces
of metamorphism by high temperature, but also sedimentary grains that have been rounded by erosion during transport by water,
showing that rivers and seas existed then.[59] Cratons consist primarily of two alternating types of terranes. The first are so-called
greenstone belts, consisting of low-grade metamorphosed sedimentary rocks. These "greenstones" are similar to the sediments today
found in oceanic trenches, above subduction zones. For this reason, greenstones are sometimes seen as evidence for subduction
during the Archean. The second type is a complex of felsic magmatic rocks. These rocks are mostly tonalite, trondhjemite or
granodiorite, types of rock similar in composition to granite (hence such terranes are called TTG-terranes). TTG-complexes are seen
[60]:Chapter 5
as the relicts of the first continental crust, formed by partial melting in basalt.

Oceans and atmosphere


Earth is often described as having had three atmospheres. The first
atmosphere, captured from the solar nebula, was composed of light
(atmophile) elements from the solar nebula, mostly hydrogen and
helium. A combination of the solar wind and Earth's heat would have
driven off this atmosphere, as a result of which the atmosphere is
now depleted of these elements compared to cosmic abundances.[62]
After the impact which created the moon, the molten Earth released
volatile gases; and later more gases were released by volcanoes,
Graph showing range of estimatedpartial pressure
completing a second atmosphere rich ingreenhouse gases but poor in of atmospheric oxygen through geologic time[61]
oxygen. [1]:256 Finally, the third atmosphere, rich in oxygen, emerged
when bacteria began to produce oxygen about
2.8 Ga.[63]:83–84, 116–117

In early models for the formation of the atmosphere and ocean, the second atmosphere was formed by outgassing of volatiles from
the Earth's interior. Now it is considered likely that many of the volatiles were delivered during accretion by a process known as
impact degassing in which incoming bodies vaporize on impact. The ocean and atmosphere would, therefore, have started to form
even as the Earth formed.[64] The new atmosphere probably contained water vapor, carbon dioxide, nitrogen, and smaller amounts of
other gases.[65]

Planetesimals at a distance of 1astronomical unit (AU), the distance of the Earth from the Sun, probably did not contribute any water
to the Earth because the solar nebula was too hot for ice to form and the hydration of rocks by water vapor would have taken too
long.[64][66] The water must have been supplied by meteorites from the outer asteroid belt and some large planetary embryos from
beyond 2.5 AU.[64][67] Comets may also have contributed. Though most comets are today in orbits farther away from the Sun than
[59]:130–132
Neptune, computer simulations show that they were originally far more common in the inner parts of the solar system.

As the Earth cooled, clouds formed. Rain created the oceans. Recent evidence suggests the oceans may have begun forming as early
as 4.4 Ga.[34] By the start of the Archean eon, they already covered much of the Earth. This early formation has been difficult to
explain because of a problem known as thefaint young Sun paradox. Stars are known to get brighter as they age, and at the time of its
formation the Sun would have been emitting only 70% of its current power. Thus, the Sun has become 30% brighter in the last 4.5
billion years.[68] Many models indicate that the Earth would have been covered in ice.[69][64] A likely solution is that there was
enough carbon dioxide and methane to produce agreenhouse effect. The carbon dioxide would have been produced by volcanoes and
the methane by early microbes. Another greenhouse gas, ammonia, would have been ejected by volcanos but quickly destroyed by
ultraviolet radiation.[63]:83

Origin of life
One of the reasons for interest in the early atmosphere and
Ice Ages Life timeline
ocean is that they form the conditions under which life first
0—
Quaternary Primates ←Earliest apes
arose. There are many models, but little consensus, on how P Flowers Birds Mammals
h
life emerged from non-living chemicals; chemical systems Karoo
– a Plants Dinosaurs
n ←Tetrapoda
created in the laboratory fall well short of the minimum Andean e Arthropods and Molluscs
-500 — r ←Cambrian explosion
complexity for a living organism.[70][71] ←Ediacaran biota
Cryogenian o
–z
o ←Earliest plants
The first step in the emergence of life may have been i
-1000 — c Multicellular life
chemical reactions that produced many of the simpler
organic compounds, including nucleobases and amino

←Sexual
acids, that are the building blocks of life. An experiment in P
reproduction
r
1953 by Stanley Miller and Harold Urey showed that such -1500 — o
t
molecules could form in an atmosphere of water, methane,
– er
ammonia and hydrogen with the aid of sparks to mimic the Eukaryotes
o
effect of lightning.[72] Although atmospheric composition -2000 — z
o
i
was probably different from that used by Miller and Urey,
Huronian – c
later experiments with more realistic compositions also
←Oxygen crisis
←Atmospheric oxygen
managed to synthesize organic molecules.[73] Computer -2500 —
simulations show that extraterrestrial organic molecules
– photosynthesis
could have formed in the protoplanetary disk before the Pongola

formation of the Earth.[74] -3000 — A


r
c
Additional complexity could have been reached from at –h
e
least three possible starting points: self-replication, an a
-3500 — n ←Earliest oxygen
organism's ability to produce offspring that are similar to Single-celled life
itself; metabolism, its ability to feed and repair itself; and –
external cell membranes, which allow food to enter and -4000 — ←Earliest life
H
waste products to leave, but exclude unwanted a
–d
water
substances.[75] e
a ←Earliest water
-4500 —n ←Earth (−4540)
Replication first: RNA world Axis scale : million years
Also see: Human timeline and Nature timeline
Even the simplest members of the three modern domains
of life use DNA to record their "recipes" and a complex
array of RNA and protein molecules to "read" these instructions and use them for growth, maintenance, and self-replication.

The discovery that a kind of RNA molecule called a ribozyme can catalyze both its own replication and the construction of proteins
led to the hypothesis that earlier life-forms were based entirely on RNA.[76] They could have formed an RNA world in which there
were individuals but no species, as mutations and horizontal gene transfers would have meant that the offspring in each generation
were quite likely to have different genomes from those that their parents started with.[77] RNA would later have been replaced by
DNA, which is more stable and therefore can build longer genomes, expanding the range of capabilities a single organism can
have.[78] Ribozymes remain as the main components ofribosomes, the "protein factories" of modern cells.[79]

Although short, self-replicating RNA molecules have been artificially produced in laboratories,[80] doubts have been raised about
whether natural non-biological synthesis of RNA is possible.[81][82][83] The earliest ribozymes may have been formed of simpler
nucleic acids such as PNA, TNA or GNA, which would have been replaced later by RNA.[84][85] Other pre-RNA replicators have
been posited, includingcrystals[86]:150 and even quantum systems.[87]

In 2003 it was proposed that porous metal sulfide precipitates would assist RNA synthesis at about 100 °C (212 °F) and at ocean-
bottom pressures near hydrothermal vents. In this hypothesis, the proto-cells would be confined in the pores of the metal substrate
until the later development of lipid membranes.[88]
Metabolism first: iron–sulfur world
Another long-standing hypothesis is that the first life was composed of protein molecules.
Amino acids, the building blocks of proteins, are easily synthesized in plausible prebiotic
conditions, as are small peptides (polymers of amino acids) that make good
catalysts.[89]:295–297 A series of experiments starting in 1997 showed that amino acids and
peptides could form in the presence of carbon monoxide and hydrogen sulfide with iron
sulfide and nickel sulfide as catalysts. Most of the steps in their assembly required
temperatures of about 100 °C (212 °F) and moderate pressures, although one stage required
250 °C (482 °F) and a pressure equivalent to that found under 7 kilometers (4.3 mi) of rock.
[90]
Hence, self-sustaining synthesis of proteins could have occurred near hydrothermal vents.

A difficulty with the metabolism-first scenario is finding a way for organisms to evolve.
Without the ability to replicate as individuals, aggregates of molecules would have
"compositional genomes" (counts of molecular species in the aggregate) as the target of
natural selection. However, a recent model shows that such a system is unable to evolve in
response to natural selection.[91]

Membranes first: Lipid world


The replicator in virtually all
It has been suggested that double-walled "bubbles" of lipids like those that form the external
known life is
membranes of cells may have been an essential first step.[92] Experiments that simulated the deoxyribonucleic acid. DNA
conditions of the early Earth have reported the formation of lipids, and these can is far more complex than the
spontaneously form liposomes, double-walled "bubbles", and then reproduce themselves. original replicator and its
Although they are not intrinsically information-carriers as nucleic acids are, they would be replication systems are
highly elaborate.
subject to natural selection for longevity and reproduction. Nucleic acids such as RNA might
[93]
then have formed more easily within the liposomes than they would have outside.

The clay theory


Some clays, notably montmorillonite, have properties that make them plausible accelerators for the emergence of an RNA world:
they grow by self-replication of their crystalline pattern, are subject to an analog of natural selection (as the clay "species" that grows
fastest in a particular environment rapidly becomes dominant), and can catalyze the formation of RNA molecules.[94] Although this
[95]:150–158 [86]
idea has not become the scientific consensus, it still has active supporters.

Research in 2003 reported that montmorillonite could also accelerate the conversion of fatty
acids into "bubbles", and that the bubbles could encapsulate RNA attached to the clay.
Bubbles can then grow by absorbing additional lipids and dividing. The formation of the
earliest cells may have been aided by similar processes.[96]

A similar hypothesis presents self-replicating iron-rich clays as the progenitors of nucleotides,


lipids and amino acids.[97]

Last universal ancestor


It is believed that of this multiplicity of protocells, only one line survived. Current
phylogenetic evidence suggests that the last universal ancestor (LUA) lived during the early
Cross-section through a
Archean eon, perhaps 3.5 Ga or earlier.[98][99] This LUA cell is the ancestor of all life on
liposome
Earth today. It was probably a prokaryote, possessing a cell membrane and probably
ribosomes, but lacking a nucleus or membrane-bound organelles such as mitochondria or
chloroplasts. Like modern cells, it used DNA as its genetic code, RNA for information transfer and protein synthesis, and enzymes to
catalyze reactions. Some scientists believe that instead of a single organism being the last universal common ancestor, there were
populations of organisms exchanging genes bylateral gene transfer.[98]

Proterozoic Eon
The Proterozoic eon lasted from 2.5 Ga to 542 Ma (million years) ago.[2]:130 In this time span, cratons grew into continents with
modern sizes. The change to an oxygen-rich atmosphere was a crucial development. Life developed from prokaryotes into eukaryotes
and multicellular forms. The Proterozoic saw a couple of severe ice ages called snowball Earths. After the last Snowball Earth about
600 Ma, the evolution of life on Earth accelerated. About 580 Ma, the Ediacaran biota formed the prelude for the Cambrian
Explosion.

Oxygen revolution
The earliest cells absorbed energy and food from the surrounding environment. They
used fermentation, the breakdown of more complex compounds into less complex
compounds with less energy, and used the energy so liberated to grow and
reproduce. Fermentation can only occur in an anaerobic (oxygen-free) environment.
The evolution of photosynthesis made it possible for cells to derive energy from the
Sun.[100]:377

Most of the life that covers the surface of the Earth depends directly or indirectly on
photosynthesis. The most common form, oxygenic photosynthesis, turns carbon
Lithified stromatolites on the shores
dioxide, water, and sunlight into food. It captures the energy of sunlight in energy-
of Lake Thetis, Western Australia.
rich molecules such as ATP, which then provide the energy to make sugars. To Archean stromatolites are the first
supply the electrons in the circuit, hydrogen is stripped from water, leaving oxygen direct fossil traces of life on Earth.
as a waste product.[101] Some organisms, including purple bacteria and green sulfur
bacteria, use an anoxygenic form of photosynthesis that uses alternatives to
hydrogen stripped from water as electron donors; examples are hydrogen sulfide,
sulfur and iron. Such extremophile organisms are restricted to otherwise inhospitable
[100]:379–382 [102]
environments such as hot springs and hydrothermal vents.

The simpler anoxygenic form arose about 3.8 Ga, not long after the appearance of
life. The timing of oxygenic photosynthesis is more controversial; it had certainly
appeared by about 2.4 Ga, but some researchers put it back as far as 3.2 Ga.[101] The
A banded iron formation from the
latter "probably increased global productivity by at least two or three orders of
3.15 Ga Moories Group,Barberton
magnitude".[103][104] Among the oldest remnants of oxygen-producing lifeforms are Greenstone Belt, South Africa. Red
fossil stromatolites.[103][104][61] layers represent the times when
oxygen was available; gray layers
At first, the released oxygen was bound up with limestone, iron, and other minerals. were formed in anoxic
The oxidized iron appears as red layers in geological strata called banded iron circumstances.
formations that formed in abundance during the Siderian period (between 2500 Ma
and 2300 Ma).[2]:133 When most of the exposed readily reacting minerals were
oxidized, oxygen finally began to accumulate in the atmosphere. Though each cell only produced a minute amount of oxygen, the
combined metabolism of many cells over a vast time transformed Earth's atmosphere to its current state. This was Earth's third
atmosphere.[105]:50–51[63]:83–84, 116–117

Some oxygen was stimulated by solar ultraviolet radiation to form ozone, which collected in a layer near the upper part of the
atmosphere. The ozone layer absorbed, and still absorbs, a significant amount of the ultraviolet radiation that once had passed
through the atmosphere. It allowed cells to colonize the surface of the ocean and eventually the land: without the ozone layer,
[106][59]:219–220
ultraviolet radiation bombarding land and sea would have caused unsustainable levels of mutation in exposed cells.
Photosynthesis had another major impact. Oxygen was toxic; much life on Earth probably died out as its levels rose in what is known
as the oxygen catastrophe. Resistant forms survived and thrived, and some developed the ability to use oxygen to increase their
metabolism and obtain more energy from the same food.[106]

Snowball Earth
The natural evolution of the Sunmade it progressively more luminous during the Archean and Proterozoic eons; the Sun's luminosity
increases 6% every billion years.[59]:165 As a result, the Earth began to receive more heat from the Sun in the Proterozoic eon.
However, the Earth did not get warmer. Instead, the geological record suggests it cooled dramatically during the early Proterozoic.
Glacial deposits found in South Africa date back to 2.2 Ga, at which time, based on paleomagnetic evidence, they must have been
located near the equator. Thus, this glaciation, known as the Huronian glaciation, may have been global. Some scientists suggest this
, a hypothesis called Snowball Earth.[107]
was so severe that the Earth was frozen over from the poles to the equator

The Huronian ice age might have been caused by the increased oxygen concentration in the atmosphere, which caused the decrease
of methane (CH4) in the atmosphere. Methane is a strong greenhouse gas, but with oxygen it reacts to form CO2, a less effective
greenhouse gas.[59]:172 When free oxygen became available in the atmosphere, the concentration of methane could have decreased
dramatically, enough to counter the effect of the increasing heat flow from the Sun.[108]

However, the term Snowball Earth is more commonly used to describe later extreme ice ages during the Cryogenian period. There
were four periods, each lasting about 10 million years, between 750 and 580 million years ago, when the earth is thought to have
been covered with ice apart from the highest mountains, and average temperatures were about −50 °C (−58 °F).[109] The snowball
may have been partly due to the location of the supercontintent Rodinia straddling the Equator. Carbon dioxide combines with rain to
weather rocks to form carbonic acid, which is then washed out to sea, thus extracting the greenhouse gas from the atmosphere. When
the continents are near the poles, the advance of ice covers the rocks, slowing the reduction in carbon dioxide, but in the Cryogienian
the weathering of Rodinia was able to continue unchecked until the ice advanced to the tropics. The process may have finally been
reversed by the emission of carbon dioxide from volcanoes or the destabilization of methane gas hydrates. According to the
.[110][111]
alternative Slushball Earth theory, even at the height of the ice ages there was still open water at the Equator

Emergence of eukaryotes
Modern taxonomy classifies life into three domains. The time of their origin is
uncertain. The Bacteria domain probably first split off from the other forms of life
(sometimes called Neomura), but this supposition is controversial. Soon after this,
by 2 Ga,[112] the Neomura split into the Archaea and the Eukarya. Eukaryotic cells
(Eukarya) are larger and more complex than prokaryotic cells (Bacteria and
Archaea), and the origin of that complexity is only now becoming known.

Around this time, the first proto-mitochondrion was formed. A bacterial cell related
to today's Rickettsia,[113] which had evolved to metabolize oxygen, entered a larger
Chloroplasts in the cells of a moss
prokaryotic cell, which lacked that capability. Perhaps the large cell attempted to
digest the smaller one but failed (possibly due to the evolution of prey defenses).
The smaller cell may have tried toparasitize the larger one. In any case, the smaller cell survived inside the larger cell. Using oxygen,
it metabolized the larger cell's waste products and derived more energy. Part of this excess energy was returned to the host. The
smaller cell replicated inside the larger one. Soon, a stable symbiosis developed between the large cell and the smaller cells inside it.
Over time, the host cell acquired some genes from the smaller cells, and the two kinds became dependent on each other: the larger
cell could not survive without the energy produced by the smaller ones, and these, in turn, could not survive without the raw materials
provided by the larger cell. The whole cell is now considered a single organism, and the smaller cells are classified as organelles
called mitochondria.[114]
A similar event occurred with photosynthetic cyanobacteria[115] entering large heterotrophic cells and becoming
chloroplasts.[105]:60–61[116]:536–539 Probably as a result of these changes, a line of cells capable of photosynthesis split off from the
other eukaryotes more than 1 billion years ago. There were probably several such inclusion events. Besides the well-established
endosymbiotic theory of the cellular origin of mitochondria and chloroplasts, there are theories that cells led to peroxisomes,
spirochetes led to cilia and flagella, and that perhaps a DNA virus led to the cell nucleus,[117][118] though none of them are widely
accepted.[119]

Archaeans, bacteria, and eukaryotes continued to diversify and to become more complex and better adapted to their environments.
Each domain repeatedly split into multiple lineages, although little is known about the history of the archaea and bacteria. Around 1.1
Ga, the supercontinent Rodinia was assembling.[120][121] The plant, animal, and fungi lines had split, though they still existed as
solitary cells. Some of these lived in colonies, and gradually a division of labor began to take place; for instance, cells on the
periphery might have started to assume different roles from those in the interior. Although the division between a colony with
specialized cells and a multicellular organism is not always clear, around 1 billion years ago[122] , the first multicellular plants
emerged, probably green algae.[123] Possibly by around 900 Ma[116]:488 true multicellularity had also evolved in animals.

At first, it probably resembled today's sponges, which have totipotent cells that allow a disrupted organism to reassemble
itself.[116]:483–487 As the division of labor was completed in all lines of multicellular organisms, cells became more specialized and
more dependent on each other; isolated cells would die.

Supercontinents in the Proterozoic


Reconstructions of tectonic plate movement in the past 250 million years (the
Cenozoic and Mesozoic eras) can be made reliably using fitting of continental
margins, ocean floor magnetic anomalies and paleomagnetic poles. No ocean crust
dates back further than that, so earlier reconstructions are more difficult.
Paleomagnetic poles are supplemented by geologic evidence such as orogenic belts,
which mark the edges of ancient plates, and past distributions of flora and fauna. The
further back in time, the scarcer and harder to interpret the data get and the more
uncertain the reconstructions.[124]:370

Throughout the history of the Earth, there have been times when continents collided
and formed a supercontinent, which later broke up into new continents. About 1000
to 830 Ma, most continental mass was united in the supercontinent A reconstruction of Pannotia
Rodinia.[124]:370[125] Rodinia may have been preceded by Early-Middle Proterozoic (550 Ma).
continents called Nuna and Columbia.[124]:374[126][127]

After the break-up of Rodinia about 800 Ma, the continents may have formed another short-lived supercontinent around 550 Ma. The
hypothetical supercontinent is sometimes referred to as Pannotia or Vendia.[128]:321–322 The evidence for it is a phase of continental
collision known as the Pan-African orogeny, which joined the continental masses of current-day Africa, South America, Antarctica
and Australia. The existence of Pannotia depends on the timing of the rifting between Gondwana (which included most of the
landmass now in the Southern Hemisphere, as well as the Arabian Peninsula and the Indian subcontinent) and Laurentia (roughly
equivalent to current-day North America).[124]:374 It is at least certain that by the end of the Proterozoic eon, most of the continental
[129]
mass lay united in a position around the south pole.

Late Proterozoic climate and life


The end of the Proterozoic saw at least two Snowball Earths, so severe that the surface of the oceans may have been completely
frozen. This happened about 716.5 and 635 Ma, in the Cryogenian period.[130] The intensity and mechanism of both glaciations are
still under investigation and harder to explain than the early Proterozoic Snowball Earth.[131] Most paleoclimatologists think the cold
episodes were linked to the formation of the supercontinent Rodinia.[132] Because Rodinia was centered on the equator, rates of
chemical weathering increased and carbon dioxide (CO2) was taken from the atmosphere. Because CO2 is an important greenhouse
gas, climates cooled globally. In the same way, during the Snowball Earths most of
the continental surface was covered with permafrost, which decreased chemical
weathering again, leading to the end of the glaciations. An alternative hypothesis is
that enough carbon dioxide escaped through volcanic outgassing that the resulting
greenhouse effect raised global temperatures.[132] Increased volcanic activity
resulted from the break-up of Rodinia at about the same time.

The Cryogenian period was followed by the Ediacaran period, which was
characterized by a rapid development of new multicellular lifeforms.[133] Whether
there is a connection between the end of the severe ice ages and the increase in
A 580 million year old fossil of
diversity of life is not clear, but it does not seem coincidental. The new forms of life, Spriggina floundensi, an animal from
called Ediacara biota, were larger and more diverse than ever. Though the taxonomy the Ediacaran period. Such life forms
of most Ediacaran life forms is unclear, some were ancestors of groups of modern could have been ancestors to the
life.[134] Important developments were the origin of muscular and neural cells. None many new forms that originated in
of the Ediacaran fossils had hard body parts like skeletons. These first appear after the Cambrian Explosion.

the boundary between the Proterozoic and Phanerozoic eons or Ediacaran and
Cambrian periods.

Phanerozoic Eon
The Phanerozoic is the current eon on Earth, which started approximately 542 million years ago. It consists of three eras: The
Paleozoic, Mesozoic, and Cenozoic,[22] and is the time when multi-cellular life greatly diversified into almost all the organisms
known today.[135]

The Paleozoic ("old life") era was the first and longest era of the Phanerozoic eon, lasting from 542 to 251 Ma.[22] During the
Paleozoic, many modern groups of life came into existence. Life colonized the land, first plants, then animals. Two major extinctions
occurred. The continents formed at the break-up of Pannotia and Rodinia at the end of the Proterozoic slowly moved together again,
forming the supercontinentPangaea in the late Paleozoic.

The Mesozoic ("middle life") era lasted from 251 Ma to 66 Ma.[22] It is subdivided into the Triassic, Jurassic, and Cretaceous
periods. The era began with the Permian–Triassic extinction event, the most severe extinction event in the fossil record; 95% of the
species on Earth died out.[136] It ended with the Cretaceous–Paleogene extinction eventthat wiped out the dinosaurs..

The Cenozoic ("new life") era began at 66 Ma,[22] and is subdivided into the Paleogene, Neogene, and Quaternary periods. These
three periods are further split into seven sub-divisions, with the Paleogene composed of The Paleocene, Eocene, and Oligocene, the
Neogene divided into the Miocene, Pliocene, and the Quaternary composed of the Pleistocene, and Holocene.[137] Mammals, birds,
amphibians, crocodilians, turtles, and lepidosaurs survived the Cretaceous–Paleogene extinction event that killed off the non-avian
dinosaurs and many other forms of life, and this is the era during which they diversified into their modern forms.

Tectonics, paleogeography andclimate


At the end of the Proterozoic, the supercontinent Pannotia had broken apart into the smaller continents Laurentia, Baltica, Siberia and
Gondwana.[138] During periods when continents move apart, more oceanic crust is formed by volcanic activity. Because young
volcanic crust is relatively hotter and less dense than old oceanic crust, the ocean floors rise during such periods. This causes the sea
level to rise. Therefore, in the first half of the Paleozoic, lar
ge areas of the continents were below sea level.

Early Paleozoic climates were warmer than today


, but the end of the Ordovician saw a shortice age during which glaciers covered the
south pole, where the huge continent Gondwana was situated. Traces of glaciation from this period are only found on former
Gondwana. During the Late Ordovician ice age, a few mass extinctions took place, in which many brachiopods, trilobites, Bryozoa
.[139]
and corals disappeared. These marine species could probably not contend with the decreasing temperature of the sea water
The continents Laurentia and Baltica collided between 450 and 400 Ma, during the
Caledonian Orogeny, to form Laurussia (also known as Euramerica).[140] Traces of
the mountain belt this collision caused can be found in Scandinavia, Scotland, and
the northern Appalachians. In the Devonian period (416–359 Ma)[22] Gondwana and
Siberia began to move towards Laurussia. The collision of Siberia with Laurussia
caused the Uralian Orogeny, the collision of Gondwana with Laurussia is called the
Variscan or Hercynian Orogeny in Europe or the Alleghenian Orogeny in North
America. The latter phase took place during the Carboniferous period (359–
299 Ma)[22] and resulted in the formation of the last supercontinent, Pangaea.
[60]

By 180 Ma, Pangaea broke up intoLaurasia and Gondwana.

Cambrian explosion Pangaea was a supercontinent that


The rate of the evolution of life as recorded by fossils accelerated in the Cambrian existed from about 300 to 180 Ma.
The outlines of the modern
period (542–488 Ma).[22] The sudden emergence of many new species, phyla, and
continents and other landmasses are
forms in this period is called the Cambrian Explosion. The biological fomenting in
indicated on this map.
the Cambrian Explosion was unpreceded before and since that time.[59]:229 Whereas
the Ediacaran life forms appear yet primitive and not easy to put in any modern
group, at the end of the Cambrian most modern phyla were already present. The
development of hard body parts such as shells, skeletons or exoskeletons in animals
like molluscs, echinoderms, crinoids and arthropods (a well-known group of
arthropods from the lower Paleozoic are the trilobites) made the preservation and
fossilization of such life forms easier than those of their Proterozoic ancestors. For
this reason, much more is known about life in and after the Cambrian than about that
of older periods. Some of these Cambrian groups appear complex but are seemingly
quite different from modern life; examples are Anomalocaris and Haikouichthys.
More recently, however, these seem to have found a place in modern classification.
Trilobites first appeared during the
During the Cambrian, the first vertebrate animals, among them the first fishes, had Cambrian period and were among
appeared.[116]:357 A creature that could have been the ancestor of the fishes, or was the most widespread and diverse
probably closely related to it, was Pikaia. It had a primitive notochord, a structure groups of Paleozoic organisms.
that could have developed into a vertebral column later. The first fishes with jaws
(Gnathostomata) appeared during the next geological period, the Ordovician. The
colonisation of new niches resulted in massive body sizes. In this way, fishes with increasing sizes evolved during the early
Paleozoic, such as the titanicplacoderm Dunkleosteus, which could grow 7 meters (23 ft) long.

The diversity of life forms did not increase greatly because of a series of mass extinctions that define widespread biostratigraphic
units called biomeres.[141] After each extinction pulse, the continental shelf regions were repopulated by similar life forms that may
have been evolving slowly elsewhere.[142] By the late Cambrian, the trilobites had reached their greatest diversity and dominated
nearly all fossil assemblages.[143]:34

Colonization of land
Oxygen accumulation from photosynthesis resulted in the formation of an ozone layer that absorbed much of the Sun's ultraviolet
radiation, meaning unicellular organisms that reached land were less likely to die, and prokaryotes began to multiply and become
better adapted to survival out of the water. Prokaryote lineages[144] had probably colonized the land as early as 2.6 Ga[145] even
before the origin of the eukaryotes. For a long time, the land remained barren of multicellular organisms. The supercontinent
Pannotia formed around 600 Ma and then broke apart a short 50 million years later.[146] Fish, the earliest vertebrates, evolved in the
oceans around 530 Ma.[116]:354 A major extinction event occurred near the end of the Cambrian period,[147] which ended 488
Ma.[148]
Several hundred million years ago, plants (probably resembling algae) and fungi
started growing at the edges of the water, and then out of it.[149]:138–140 The oldest
fossils of land fungi and plants date to 480–460 Ma, though molecular evidence
suggests the fungi may have colonized the land as early as 1000 Ma and the plants
700 Ma.[150] Initially remaining close to the water's edge, mutations and variations
resulted in further colonization of this new environment. The timing of the first
animals to leave the oceans is not precisely known: the oldest clear evidence is of
arthropods on land around 450 Ma,[151] perhaps thriving and becoming better
adapted due to the vast food source provided by the terrestrial plants. There is also Artist's conception of Devonian flora
unconfirmed evidence that arthropods may have appeared on land as early as 530
Ma.[152]

Evolution of tetrapods
At the end of the Ordovician period, 443 Ma,[22] additional extinction events
occurred, perhaps due to a concurrent ice age.[139] Around 380 to 375 Ma, the first
tetrapods evolved from fish.[153] Fins evolved to become limbs that the first
tetrapods used to lift their heads out of the water to breathe air. This would let them
Tiktaalik, a fish with limb-like fins and
live in oxygen-poor water, or pursue small prey in shallow water.[153] They may
a predecessor of tetrapods.
have later ventured on land for brief periods. Eventually, some of them became so
Reconstruction from fossils about
well adapted to terrestrial life that they spent their adult lives on land, although they 375 million years old.
hatched in the water and returned to lay their eggs. This was the origin of the
amphibians. About 365 Ma, another period of extinction occurred, perhaps as a
result of global cooling.[154] Plants evolved seeds, which dramatically accelerated their spread on land, around this time (by
approximately 360 Ma).[155][156]

About 20 million years later (340 Ma[116]:293–296 ), the amniotic egg evolved, which could be laid on land, giving a survival
advantage to tetrapod embryos. This resulted in the divergence of amniotes from amphibians. Another 30 million years
(310 Ma[116]:254–256 ) saw the divergence of the synapsids (including mammals) from the sauropsids (including birds and reptiles).
Other groups of organisms continued to evolve, and lines diverged—in fish, insects, bacteria, and so on—but less is known of the
details.

After yet another, the most severe extinction of the period (251~250 Ma), around
230 Ma, dinosaurs split off from their reptilian ancestors.[157] The Triassic–Jurassic
extinction event at 200 Ma spared many of the dinosaurs,[22][158] and they soon
became dominant among the vertebrates. Though some mammalian lines began to
separate during this period, existing mammals were probably small animals
resembling shrews.[116]:169

The boundary between avian and non-avian dinosaurs is not clear, but
Archaeopteryx, traditionally considered one of the first birds, lived around
150 Ma.[159] Dinosaurs were the dominant
terrestrial vertebrates throughout
The earliest evidence for the angiosperms evolving flowers is during the Cretaceous most of the Mesozoic
period, some 20 million years later (132 Ma).[160]

Extinctions
The first of five great mass extinctions was the Ordovician-Silurian extinction. Its possible cause was the intense glaciation of
Gondwana, which eventually led to asnowball earth. 60% of marine invertebrates became extinct and 25% of all families.
The second mass extinction was the Late Devonian extinction, probably caused by the evolution of trees, which could have led to the
depletion of greenhouse gases (like CO2) or theeutrophication of water. 70% of all species became extinct.

The third mass extinction was the Permian-Triassic, or the Great Dying, event was possibly caused by some combination of the
Siberian Traps volcanic event, an asteroid impact, methane hydrate gasification, sea level fluctuations, and a major anoxic event.
Either the proposed Wilkes Land crater[161] in Antarctica or Bedout structure off the northwest coast of Australia may indicate an
impact connection with the Permian-Triassic extinction. But it remains uncertain whether either these or other proposed Permian-
Triassic boundary craters are either real impact craters or even contemporaneous with the Permian-Triassic extinction event. This was
by far the deadliest extinction ever, with about 57% of allfamilies and 83% of all genera killed.[162][163]

The fourth mass extinction was the Triassic-Jurassic extinction event in which almost all synapsids and archosaurs became extinct,
probably due to new competition from dinosaurs.

The fifth and most recent mass extinction was the K-T extinction. In 66 Ma, a 10-kilometer (6.2 mi) asteroid struck Earth just off the
Yucatán Peninsula—somewhere in the south western tip of then Laurasia—where the Chicxulub crater is today. This ejected vast
quantities of particulate matter and vapor into the air that occluded sunlight, inhibiting photosynthesis. 75% of all life, including the
non-avian dinosaurs, became extinct,[164] marking the end of the Cretaceous period and Mesozoic era.

Diversification of mammals
The first true mammals evolved in the shadows of dinosaurs and other large archosaurs that filled the world by the late Triassic. The
first mammals were very small, and were probably nocturnal to escape predation. Mammal diversification truly began only after the
Cretaceous-Paleogene extinction event.[165] By the early Paleocene the earth recovered from the extinction, and mammalian diversity
increased. Creatures like Ambulocetus took to the oceans to eventually evolve into whales,[166] whereas some creatures, like
primates, took to the trees.[167] This all changed during the mid to late Eocene when the circum-Antarctic current formed between
Antarctica and Australia which disrupted weather patterns on a global scale. Grassless savannas began to predominate much of the
landscape, and mammals such as Andrewsarchus rose up to become the largest known terrestrial predatory mammal ever,[168] and
early whales like Basilosaurus took control of the seas.

The evolution of grass brought a remarkable change to the Earth's landscape, and the new open spaces created pushed mammals to
get bigger and bigger. Grass started to expand in the Miocene, and the Miocene is where many modern- day mammals first appeared.
Giant ungulates like Paraceratherium and Deinotherium evolved to rule the grasslands. The evolution of grass also brought primates
down from the trees, and started human evolution. The first big cats evolved during this time as well.[169] The Tethys Sea was closed
off by the collision of Africa and Europe.[170]

The formation of Panama was perhaps the most important geological event to occur in the last 60 million years. Atlantic and Pacific
currents were closed off from each other, which caused the formation of the Gulf Stream, which made Europe warmer. The land
bridge allowed the isolated creatures of South America to migrate over to North America, and vice versa.[171] Various species
migrated south, leading to the presence in South America ofllamas, the spectacled bear, kinkajous and jaguars.

Three million years ago saw the start of the Pleistocene epoch, which featured dramatic climactic changes due to the ice ages. The ice
ages led to the evolution of modern man in Saharan Africa and expansion. The mega-fauna that dominated fed on grasslands that, by
now, had taken over much of the subtropical world. The large amounts of water held in the ice allowed for various bodies of water to
shrink and sometimes disappear such as the North Sea and the Bering Strait. It is believed by many that a huge migration took place
along Beringia which is why, today, there are camels (which evolved and became extinct in North America), horses (which evolved
and became extinct in North America), and Native Americans. The ending of the last ice age coincided with the expansion of man,
along with a massive die out of ice age mega-fauna. This extinction, nicknamedthe
" Sixth Extinction", has been going ever since.

Human evolution

Hominin timeline
0— ←Modern humans
Homo sapiens
0— ←Modern humans
A small African ape living around 6 Ma was the last P Neanderthals Homo sapiens ←Modern speech
animal whose descendants would include both modern – l H. heidelbergensis ←Earliest clothes
e ←Earliest cooking
humans and their closest relatives, the -1 — i
s Homo erectus
chimpanzees.[116]:100–101 Only two branches of its family
– t ←Earliest fire use
tree have surviving descendants. Very soon after the split, o ←Exit from Africa
for reasons that are still unclear, apes in one branch -2 — c
e Homo habilis
developed the ability to walk upright.[116]:95–99 Brain size – n
e
increased rapidly, and by 2 Ma, the first animals classified
-3 —
in the genus Homo had appeared.[149]:300 Of course, the
←Stone tools
line between different species or even genera is somewhat – Australopithecus
P
arbitrary as organisms continuously change over -4 — l ←Earliest bipedal
generations. Around the same time, the other branch split i
Ardipithecus H
–o
into the ancestors of the common chimpanzee and the c
ancestors of the bonobo as evolution continued -5 — e o
n
simultaneously in all life forms.[116]:100–101 m
–e Hominini
The ability to control fire probably began in Homo erectus -6 — Orrorin i ←Chimpanzee split
(or Homo ergaster), probably at least 790,000 years n

ago[172] but perhaps as early as 1.5 Ma.[116]:67 The use
Sahelanthropus i ←
and discovery of controlled fire may even predate Homo -7 — M Possibly bipedal
i d
erectus. Fire was possibly used by the early Lower –o
Paleolithic (Oldowan) hominid Homo habilis or strong c s
-8 — e
australopithecines such as Paranthropus.[173] n
–e

-9 — Ouranopithecus ←Gorilla split



Nakalipithecus
-10 — ←Earlier apes
Axis scale : million years
Also see: Life timeline and Nature timeline

It is more difficult to establish the origin of language; it is unclear whether Homo


erectus could speak or if that capability had not begun until Homo sapiens.[116]:67
As brain size increased, babies were born earlier
, before their heads grew too large to
pass through the pelvis. As a result, they exhibited more plasticity, and thus
possessed an increased capacity to learn and required a longer period of dependence.
A reconstruction of human history Social skills became more complex, language became more sophisticated, and tools
based on fossil data.[174] became more elaborate. This contributed to further cooperation and intellectual
development.[175]:7 Modern humans (Homo sapiens) are believed to have originated
around 200,000 years ago or earlier in Africa; the oldest fossils date back to around
160,000 years ago.[176]

The first humans to show signs of spirituality are the Neanderthals (usually classified as a separate species with no surviving
descendants); they buried their dead, often with no sign of food or tools.[177]:17 However, evidence of more sophisticated beliefs,
such as the early Cro-Magnon cave paintings (probably with magical or religious significance)[177]:17–19 did not appear until
32,000 years ago.[178] Cro-Magnons also left behind stone figurines such as Venus of Willendorf, probably also signifying religious
belief.[177]:17–19 By 11,000 years ago, Homo sapiens had reached the southern tip of South America, the last of the uninhabited
continents (except for Antarctica, which remained undiscovered until 1820 AD).[179] Tool use and communication continued to
improve, and interpersonal relationships became more intricate.
Civilization
Throughout more than 90% of its history, Homo sapiens lived in small bands as
nomadic hunter-gatherers.[175]:8 As language became more complex, the ability to
remember and communicate information resulted, according to a theory proposed by
Richard Dawkins, in a new replicator: the meme.[180] Ideas could be exchanged
quickly and passed down the generations. Cultural evolution quickly outpaced
biological evolution, and history proper began. Between 8500 and 7000 BC, humans
in the Fertile Crescent in the Middle East began the systematic husbandry of plants
and animals: agriculture.[181] This spread to neighboring regions, and developed
independently elsewhere, until most Homo sapiens lived sedentary lives in
permanent settlements as farmers. Not all societies abandoned nomadism, especially
those in isolated areas of the globe poor in domesticable plant species, such as
Australia.[182] However, among those civilizations that did adopt agriculture, the
relative stability and increased productivity provided by farming allowed the
population to expand.

Agriculture had a major impact; humans began to affect the environment as never Vitruvian Man by Leonardo da Vinci
before. Surplus food allowed a priestly or governing class to arise, followed by epitomizes the advances in art and
increasing division of labor. This led to Earth's first civilization at Sumer in the science seen during the
Middle East, between 4000 and 3000 BC.[175]:15 Additional civilizations quickly Renaissance.
arose in ancient Egypt, at the Indus River valley and in China. The invention of
writing enabled complex societies to arise: record-keeping and libraries served as a
storehouse of knowledge and increased the cultural transmission of information. Humans no longer had to spend all their time
working for survival, enabling the first specialized occupations (e.g. craftsmen, merchants, priests, etc...). Curiosity and education
drove the pursuit of knowledge and wisdom, and various disciplines, including
science (in a primitive form), arose. This in turn led to
the emergence of increasingly larger and more complex civilizations, such as the first empires, which at times traded with one
another, or fought for territory and resources.

By around 500 BC, there were advanced civilizations in the Middle East, Iran, India, China, and Greece, at times expanding, at times
entering into decline.[175]:3 In 221 BC, China became a single polity that would grow to spread its culture throughout East Asia, and
it has remained the most populous nation in the world. The fundamentals of Western civilization were largely shaped in Ancient
Greece, with the world's first democratic government and major advances in philosophy, science, and mathematics, and in Ancient
Rome in law, government, and engineering.[183] The Roman Empire was Christianized by Emperor Constantine in the early 4th
century and declined by the end of the 5th. Beginning with the 7th century, Christianization of Europe began. In 610, Islam was
founded and quickly became the dominant religion in Western Asia. The House of Wisdom was established in Abbasid-era Baghdad,
Iraq.[184] It is considered to have been a major intellectual center during the Islamic Golden Age, where Muslim scholars in Baghdad
and Cairo flourished from the ninth to the thirteenth centuries until the Mongol sack of Baghdad in 1258 AD. In 1054 AD the Great
Schism between the Roman Catholic Church and the Eastern Orthodox Church led to the prominent cultural differences between
Western and Eastern Europe.

In the 14th century, the Renaissance began in Italy with advances in religion, art, and science.[175]:317–319 At that time the Christian
Church as a political entity lost much of its power. In 1492, Christopher Columbus reached the Americas, initiating great changes to
the new world. European civilization began to change beginning in 1500, leading to the scientific and industrial revolutions. That
continent began to exert political and cultural dominance over human societies around the world, a time known as the Colonial era
(also see Age of Discovery).[175]:295–299 In the 18th century a cultural movement known as the Age of Enlightenmentfurther shaped
the mentality of Europe and contributed to its secularization. From 1914 to 1918 and 1939 to 1945, nations around the world were
embroiled in world wars. Established following World War I, the League of Nations was a first step in establishing international
institutions to settle disputes peacefully. After failing to prevent World War II, mankind's bloodiest conflict, it was replaced by the
United Nations. After the war, many new states were formed, declaring or being granted independence in a period of decolonization.
The United States and Soviet Union became the world's dominant superpowers for a time, and they held an often-violent rivalry
known as the Cold War until the dissolution of the latter. In 1992, several European nations joined in the European Union. As
transportation and communication improved, the economies and political affairs of nations around the world have become
increasingly intertwined. Thisglobalization has often produced both conflict and cooperation.

Recent events
Change has continued at a rapid pace from the mid-1940s to today. Technological
developments include nuclear weapons, computers, genetic engineering, and
nanotechnology. Economic globalization, spurred by advances in communication
and transportation technology, has influenced everyday life in many parts of the
world. Cultural and institutional forms such as democracy, capitalism, and
environmentalism have increased influence. Major concerns and problems such as
disease, war, poverty, violent radicalism, and recently, human-caused climate change
have risen as the world population increases.

In 1957, the Soviet Union launched the first artificial satellite into orbit and, soon
afterward, Yuri Gagarin became the first human in space. Neil Armstrong, an
American, was the first to set foot on another astronomical object, the Moon. Astronaut Bruce McCandless II
Unmanned probes have been sent to all the known planets in the solar system, with outside of the space shuttle
some (such as Voyager) having left the solar system. Five space agencies, Challenger in 1984
representing over fifteen countries,[185] have worked together to build the
International Space Station. Aboard it, there has been a continuous human presence
in space since 2000.[186] The World Wide Web became a part of everyday life in the 1990s, and since then has become an
indispensable source of information in thedeveloped world.

See also
Chronology of the universe– The history and future of Geological history of Earth– The sequence of major
the universe according to Big Bang cosmology geological events in Earth's past
Detailed logarithmic timeline Global catastrophic risk– Hypothetical future event
Evolutionary history of life– The processes by which that has the potential to damage human well-being on
organisms evolved on Earth a global scale
Future of Earth – Long term extrapolated geological Timeline of the evolutionary history of life– The
and biological changes current scientific theory outlining the major events
during the development of life
Timeline of natural history

Notes
1. Pluto's satellite Charon is relatively larger,[44] but Pluto is defined as adwarf planet.[45]

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Further reading
Dalrymple, G.B. (1991). The Age of the Earth. California: Stanford University Press.ISBN 978-0-8047-1569-0.
Dalrymple, G. Brent (2001)."The age of the Earth in the twentieth century: a problem (mostly) solved"
. Geological
Society, London, Special Publications. 190 (1): 205–221. Bibcode:2001GSLSP.190..205D.
doi:10.1144/GSL.SP.2001.190.01.14. Retrieved 2012-04-13.
Dawkins, Richard (2004). The Ancestor's Tale: A Pilgrimage to the Dawn of Life. Boston: Houghton Mifflin Company.
ISBN 978-0-618-00583-3.
Gradstein, F.M.; Ogg, James George; Smith,Alan Gilbert, eds. (2004).A Geological Time Scale 2004. Reprinted
with corrections 2006. Cambridge University Press.ISBN 978-0-521-78673-7.
Gradstein, Felix M.; Ogg, James G.; van Kranendonk, Martin (2008).On the Geological Time Scale 2008 (PDF)
(Report). International Commission on Stratigraphy
. Fig. 2. Archived fromthe original (PDF) on 28 October 2012.
Retrieved 20 April 2012.
Levin, H.L. (2009). The Earth through time(9th ed.). Saunders College Publishing.ISBN 978-0-470-38774-0.
Lunine, J.I. (1999). Earth: evolution of a habitable world. United Kingdom: Cambridge University Press. ISBN 978-0-
521-64423-5.
McNeill, Willam H. (1999) [1967]. A World History (4th ed.). New York: Oxford University Press. ISBN 978-0-19-
511615-1.
Melosh, H.J.; Vickery, A.M. & Tonks, W.B. (1993). Impacts and the early environment and evolution of the terrestrial
planets, in Levy, H.J. & Lunine, J.I. (eds.):Protostars and Planets III, University of Arizona Press, Tucson, pp. 1339–
1370.
Stanley, Steven M. (2005). Earth system history (2nd ed.). New York: Freeman. ISBN 978-0-7167-3907-4.
Stern, T.W.; Bleeker, W. (1998). "Age of the world's oldest rocks refined using Canada's SHRIMP: The Acasta
Gneiss Complex, Northwest Territories, Canada". Geoscience Canada. 25: 27–31.
Wetherill, G.W. (1991). "Occurrence of Earth-Like Bodies in Planetary Systems".Science. 253 (5019): 535–538.
Bibcode:1991Sci...253..535W. doi:10.1126/science.253.5019.535. PMID 17745185.

External links
Davies, Paul. "Quantum leap of life". The Guardian. 2005 December 20. – discusses speculation on the role of
quantum systems in the origin of life
Evolution timeline (uses Shockwave). Animated story of life shows everything from the big bang to the formation of
the earth and the development of bacteria and other organisms to the ascent of man.
25 biggest turning points in earth HistoryBBC
Evolution of the Earth. Timeline of the most important events in theevolution of the Earth.
The Earth's Origins on In Our Time at the BBC
Ageing the Earth, BBC Radio 4 discussion with Richard Corfield, Hazel Rymer & Henry GeeIn( Our Time, Nov. 20,
2003)

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