Origins of Life
Origins of Life
Origins of Life
History of
Life
Learning Targets
We study the remains of things that existed many years ago. The
Ruins of Pompeii have given archeologists, historians, and other
scholars a tremendous amount of information about life two
thousand years ago. This section discusses studying things that
are many thousands of years older than these remains.
Earth in a Day
It’s hard to grasp the vast amounts of time since Earth formed
and life first appeared on its surface. It may help to think of History of Earth in a Day. In this model of Earth’s history, the planet formed at
midnight. What time was it when the first prokaryotes evolved?
Earth’s history as a 24-hour day, as shown in the picture on the
next page. Humans would have appeared only during the last Learning About the Past
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Much of what we know about the history of life on Earth is based To be preserved as fossils, remains must be covered quickly by
on the fossil record. Detailed knowledge of modern organisms sediments or preserved in some other way. For example, they
also helps us understand how life evolved. may be frozen in glaciers or trapped in tree resin. Sometimes
traces of organisms—such as footprints or burrows—are
The Fossil Record
preserved. The conditions required for fossils to form rarely occur.
Fossils are the preserved remains or traces of organisms that Therefore, the chance of an organism being preserved as a fossil
lived in the past. The soft parts of organisms almost always is very low.
decompose quickly after death. On occasion, the hard parts— The photo on the left
mainly bones, teeth, or shells—remain long enough to mineralize shows an ancient
and form fossils. An example of a complete fossil skeleton is frog trapped in
hardened tree resin,
shown in figure below. The fossil record is the record of life that
or amber. The photo
unfolded over four billion years and pieced back together through on the right shows
the analysis of fossils. the fossil footprints
of a dinosaur.
Video: Fossils
Extinct Lion Fossil. This fossilized skeleton represents an extinct lion species. It
is rare for fossils to be so complete and well preserved as this one.
Watch this video link to see in more detain how fossils
form.http://www.youtube.com/watch?v=A5i5Qrp6sJU
In order
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for fossils to “tell” us the story of life, they must be dated. Then Relative Dating Using Rock
they can help scientists reconstruct how life changed over time. Layers. Relative dating
establishes which of two fossils
Fossils can be dated in two different ways: relative dating and
is older than the other. It is
absolute dating. based on the rock layers in
which the fossils formed.
Relative dating determines which of two fossils is older or
younger than the other, but not their age in years. Relative dating
is based on the positions of fossils in rock layers. Lower layers
were laid down earlier, so they are assumed to contain older
fossils. This is illustrated in figure on the next page.
Molecular Clocks
Evidence from the fossil record can be combined with data from
molecular clocks. A molecular clock uses DNA sequences (or
the proteins they encode) to estimate relatedness among species.
Molecular clocks estimate the time in geologic history when
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related species diverged from a common ancestor. Molecular climate, and the evolution of life. It organizes Earth’s history and
clocks are based on the assumption that mutations accumulate the evolution of life on the basis of important events instead of
through time at a steady average rate for a given region of DNA. time alone. It also allows more focus to be placed on recent
Species that have accumulated greater differences in their DNA events, about which we know the most.
sequences are assumed to have diverged from their common
ancestor in the more distant past. Molecular clocks based on
different regions of DNA may be used together for more accuracy.
Consider the example in the table below. The table shows how
similar the DNA of several animal species is to human DNA.
Based on these data, which organism do you think shared the
most recent common ancestor with humans?
Chimpanzee 98
Mouse 85
Chicken 60
Fruit Fly 44
Another tool for understanding the history of Earth and its life is
the geologic time scale, shown on the next page . The geologic
time scale divides Earth’s history into divisions (such as eons, Geologic Time Scale. The geologic time scale divides Earth’s history into units
eras, and periods) that are based on major changes in geology, that reflect major changes in Earth and its life forms.
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Click on this link: A Brief History of Life; Explore 4.5 billion years Review
of life in just a few minutes.
1. What are fossils?
http://www.pbs.org/wgbh/nova/evolution/brief-history-life.html
2. Describe how fossils form
Summary
Species DNA Similarity with Species A
• Much of what we know about the history of life on Earth is Species B 42%
based on the fossil record.
Species C 85%
• Molecular clocks are used to estimate how long it has been Species D 67%
since two species diverged from a common ancestor.
Species E 91%
• The geologic time scale is another important tool for
understanding the history of life on Earth. 3. This table shows DNA sequence comparisons for some
hypothetical species. Based on the data, describe evolutionary
relationships between Species A and the other four species.
Explain your answer.
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Don't Call That A Duckbill!! Video: Leonardo the Mummified Dinosaur
Student Exploration
Watch this video to find out about a truly remarkable find, a
Because most of them went extinct and the Age of Dinosaurs ended dinosaur with it's last meal preserved! http://www.youtube.com/
and the Age of Mammals began, some people still think of dinosaurs watch?v=ihifMrV3-pY
as failed organisms. Somehow they weren't as good as mammals,
they were flawed and these flaws did them in. Science no longer sees
dinosaurs this way. Remember, science is an ongoing process where
theories and views are reevaluated as new information comes in. The Extension Investigation
ongoing work of scientists has led to the recognition of Dinosauria
as a distinct phylum all its own, as well as recognizing that not all Use the resources below to answer the following questions:
dinosaurs went extinct, just the non-avian ones, which is a fancy 1. Why does Dr. Bakker hate the name duckbill? Explain your an-
way of saying dinosaurs are still around, we just call them birds. swer as fully as you can.
One reason for these advances are new fossil finds. As more and 2. How do the bills of duckbill dinosaurs differ from those of
more finds were discovered and science advanced in other areas like ducks?
geology, ecology, paleoclimatology, evolution and genetics, it be- 3. Why do large herbivores need to have large guts? What do
came increasingly clear that dinosaurs were not just big lizards but they have in these guts that humans don't need?
something quite different and quite well adjusted to the world they 4. What was one suspicion of scientists that Leonardo confirmed?
lived in. 5. How do the teeth of duckbill dinosaurs differ from those of
cows or horses? How did the vegetation that duckbill dinosaurs
ate differ from the vegetation eaten by cows and horses today?
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Chapter 2
The Origin of
Life
Learning Targets
source:http://www.accessexcellence.org/RC/AB/BC/
Spontaneous_Generation.php
From the time of the ancient Romans, through the Middle Ages,
and until the late nineteenth century, it was generally accepted
that some life forms arose spontaneously from non-living matter.
Such "spontaneous generation" appeared to occur primarily in
decaying matter. For example, a seventeenth century recipe for
the spontaneous production of mice required placing sweaty
underwear and husks of wheat in an open-mouthed jar, then
waiting for about 21 days, during which time it was alleged that This was one of the first examples of an experiment in the
the sweat from the underwear would penetrate the husks of modern sense, in which controls are used. In spite of his well-
wheat, changing them into mice. Although such a concept may executed experiment, the belief in spontaneous generation
remained strong, and even Redi continued to believe it occurred
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under some circumstances. The invention of the microscope only Academy of Sciences sponsored a contest for the best
served to enhance this belief. Microscopy revealed a whole new experiment either proving or disproving spontaneous generation.
world of organisms that appeared to arise spontaneously. It was Pasteur's winning experiment was a variation of the methods of
quickly learned that to create "animalcules," as the organisms Needham and Spallanzani. He boiled meat broth in a flask,
were called, you needed only to place hay in water and wait a few heated the neck of the flask in a flame until it became pliable, and
days before examining your new creations under the microscope. bent it into the shape of an S. Air could enter the flask, but
airborne microorganisms could not - they would settle by gravity
The debate over spontaneous generation continued for centuries.
in the neck. As Pasteur had expected, no microorganisms grew.
In 1745, John Needham, an English clergyman, proposed what he
When Pasteur tilted the flask so that the broth reached the lowest
considered the definitive experiment. Everyone knew that boiling
point in the neck, where any airborne particles would have
killed microorganisms, so he proposed to test whether or not
settled, the broth rapidly became cloudy with life. Pasteur had
microorganisms appeared spontaneously after boiling. He boiled
both refuted the theory of spontaneous generation and
chicken broth, put it into a flask, sealed it, and waited - sure
convincingly demonstrated that microorganisms are everywhere -
enough, microorganisms grew. Needham claimed victory for
even in the air. The idea of spontaneous generation was replaced
spontaneous generation.
with the theory of biogenesis. This theory states that only living
An Italian priest, Lazzaro Spallanzani, was not convinced, and he organisms can produce other living organisms.
suggested that perhaps the microorganisms had entered the
Origins: Modern Ideas
broth from the air after the broth was boiled, but before it was
sealed. To test his theory, he modified Needham's experiment -
he placed the chicken broth in a flask, sealed the flask, drew off
the air to create a partial vacuum, then boiled the broth. No
microorganisms grew. Proponents of spontaneous generation
argued that Spallanzani had only proven that spontaneous
generation could not occur without air.
http://www.pbs.org/wgbh/nova/evolution/origins-life.html
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Which Organic Molecule Came First?
Some scientists speculate that RNA may have been the first
organic molecule to evolve. In fact, they think that early life was
based solely on RNA and that DNA and proteins evolved later.
You can learn more about the RNA world hypothesis and the evidence
This is called the RNA world hypothesis. Why RNA It can
for it by watching the video. http://www.youtube.com/watch?
encode genetic instructions (like DNA), and some RNAs can carry v=sAkgb3yNgqg
out chemical reactions (like proteins). Therefore, it solves the
chicken-and-egg problem of which of these two molecules came
first. Other evidence also suggests that RNA may be the most
ancient of the organic molecules.
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Summary
• RNA may have been the first organic molecule to form as well
as the basis of early life.
Review
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Chapter 3
First Cells
Learning Targets
What was needed for the first cell? Some sort of membrane
surrounding organic molecules? Probably.
You start with simple ones. The first cells were most likely
primitive prokaryotic-like cells, even more simplistic than these E.
coli bacteria. The first cells were probably no more than organic
compounds such as a simplistic RNA, surrounded by a
membrane. Was it a phospholipid bilayer membrane? Probably
not — it was likely a simplistic membrane able to separate the
inside from the outside. Over time, as other organic compounds
such as DNA and proteins developed, cells also evolved into
more complex structures. Once a cell was able to be stable,
reproduce itself, and pass its genetic information to the next
generation, then there was life
Hypothetical First Cell. The earliest cells may have consisted of little more than
RNA inside a lipid membrane.
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LUCA Photosynthesis and Cell Respiration
No doubt there were many early cells of this type. However, The earliest cells were probably heterotrophs. Most likely they
scientists think that only one early cell (or group of cells) got their energy from other molecules in the organic “soup.”
eventually gave rise to all subsequent life on Earth. That one cell However, by about 3 billion years ago, a new way of obtaining
is called the Last Universal Common Ancestor (LUCA). It energy evolved. This new way was photosynthesis. Through
probably existed around 3.5 billion years ago. LUCA was one of photosynthesis, organisms could use sunlight to make food from
the earliest prokaryotic cells. It would have lacked a nucleus and carbon dioxide and water. These organisms were the first
other membrane-bound organelles. autotrophs. They provided food for themselves and for other
organisms that began to consume them.
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To learn more about LUCA and universal common descent, watch this
video http://www.youtube.com/watch?v=G0UGpcea8Zg
Summary
• One cell (or group of cells), called the last universal common
ancestor (LUCA), gave rise to all subsequent life on Earth.
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Chapter 4
Evolution of
Eukaryotes
Learning Targets
Why can this fish live in these tentacles, but other fish cannot?
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The first eukaryotic cells - cells with a nucleus an internal energy not only to themselves but also to the large cell. They
membrane-bound organelles - probably evolved about 2 billion became the mitochondria of eukaryotic cells. Other small cells
years ago. This is explained by the endosymbiotic theory . As were able to use sunlight to make food. They shared the food
shown in the diagram on the next page, endosymbiosis came with the large cell. They became the chloroplasts of eukaryotic
about when large cells engulfed small cells. The small cells were cells.
not digested by the large cells. Instead, they lived within the large
Mitochondria and Chloroplasts
cells and evolved into organelles.
What is the evidence for this evolutionary pathway? Biochemistry
and electron microscopy provide convincing support. The
mitochondria and chloroplasts within our eukaryotic cells share
the following features with prokaryotic cells:
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The "host" cell membrane and biochemistry are more similar to
those of Archaebacteria, so scientists believe eukaryotes Video: Endosymbiosis
descended more directly from that major group. The timing of this
dramatic evolutionary event (more likely a series of events) is not
clear. The oldest fossil clearly related to modern eukaryotes is a
red alga dating back to 1.2 billion years ago. However, many
scientists place the appearance of eukaryotic cells at about 2
billion years. Some time within Proterozoic Eon, then, all three
major groups of life – Bacteria, Archea, and Eukaryotes – became
well established.
http://www.youtube.com/watch?v=-FQmAnmLZtE
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of early evolution. Often, as humans, we focus our attention on
plants and animals, and ignore bacteria. Our human senses cannot
directly perceive the unimaginable variety of single cells, the
architecture of organic molecules, or the intricacy of
biochemical pathways. Let your study of early evolution give you a
new perspective – a window into the beauty and diversity of unseen
worlds, now and throughout Earth’s history. In addition to the
mitochondria that call your 100 trillion cells home, your body
contains more bacterial cells than human cells. You, mitochondria,
and your resident bacteria share common ancestry.
Summary
Review
1. Describe the endosymbiotic theory.
2. Discuss the evidence for the evolution of mitochondria and
chloroplasts.
I
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I
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Absolute dating
the process of determining an approximate computed age in archaeology and geology; the
determination of the age of an object with reference to a specific time scale, such as a fixed
calendar or in years, based on measurable physical or chemical qualities
organism that captures energy from sunlight or inorganic substances to produce its own
food; provides the foundation of the food supply for other organisms; also called a
producer.
chemical process in which organic molecules are broken down to release energy for use by
the cell; process that releases energy by breaking down glucose and other food molecules
in the presence of oxygen.
theory that eukaryotic cells formed from a symbiosis among several different prokaryotic
organisms; explains the evolution of eukaryotic cells from prokaryotic cells.
cells with membrane-bound nucleus and organelles; generally larger and more complex
than prokaryotic cells.
organisms that cannot make its own food and gets its nutrients and energy requirements by
feeding on other organisms; also called a consumer.
is the most recent organism from which all organisms now living on Earth descend. Thus it
is the most recent common ancestor of all current life on Earth. The LUCA (last universal
common ancestor) is estimated to have lived some 3.5 to 3.8 billion years ago
model that uses comparisons of DNA sequences to estimate phylogeny and rate of
evolutionary change.
process by which plants and some other organisms use light energy to convert water and
carbon dioxide into oxygen and high-energy carbohydrates such as sugars and starches; a
two-phase anabolic pathway in which the sun’s light energy is converted to chemical
energy for use by the cell.
used to arrange geological events, and the rocks they leave behind, in a sequence. The
method of reading the order of rock layers. Relative dating does not provide actual
numerical dates for the rocks; the science of determining the relative order of past events,
without necessarily determining their absolute age.
proposes that a world filled with life based on ribonucleic acid (RNA) predates the current
world of life based on deoxyribonucleic acid (DNA) and protein. RNA, which can both store
information like DNA and act as an enzyme like proteins, may have supported cellular or
pre-cellular life. Some hypotheses as to the origin of life present RNA-based catalysis and
information storage as the first step in the evolution of cellular life.
proposes that self-replicating ribonucleic acid (RNA) molecules were precursors to current
life, which is based on deoxyribonucleic acid (DNA), RNA and proteins. It is generally
accepted that current life on Earth descends from an RNA world.
It was once believed that life could come from nonliving things, such as mice from corn,
flies from bovine manure, maggots from rotting meat, and fish from the mud of previously
dry lakes. Spontaneous generation is the incorrect hypothesis that nonliving things are
capable of producing life. Several experiments have been conducted to disprove
spontaneous generation;