Soil Bacteria - NRCS Soils
Soil Bacteria - NRCS Soils
Soil Bacteria - NRCS Soils
MNOPQRSPTSRSUPTNVSPWPXNYZPTSQZ[\PWPXNYZP]YNZN^_PPWPXNYZP]Q`[SRYQ
óôõö÷øùúúûüôûýP P P P P
18,>0h.2>-i
yz{|}~{
]_PZQYgSP
Pg^\QV
18,>0h.2>-i0j0h87.
18,>0k,8>8;B }}y}~
18,>0h.2>-i0:33.337.+- ]Q`[SRYQPQRSP[Yg_PNgS`SZZSPNR^QgYbVbPP^SgSRQZZ_PWPNPQgPYg`\PYSPPV PQgPbNVS\Q[PZNg^SRPYg
18,>0h.2>-i0l2+2;.7.+- ZSg^[\P¡\Q[PdQ`[SRYQPZQ`¢PYgPbY£SP[\S_PVQ¢SPOcPYgPgOVdSRbP¤P[SQbcNNgPNPcRNO`[Y¥SPbNYZP^SgSRQZZ_P`Ng[QYgb
C.38=6<.30m0n=?>,<2-,8+3 dS[SSgPPVYZZYNgPQgPPdYZZYNgPdQ`[SRYQP¦\Q[PYbPQbPVO`\PVQbbPQbP[NP`NbPcSRPQ`RS
¤P[NgPNPVỲRNb`NcỲPdQ`[SRYQPVQ_PdSPQ`[Y¥SPYgPSQ`\PQ`RSPNPbNYZ
18,>0k,8>8;B0n6,7.60Ei25-.63
§¨©ª«¬®¯«°±²©³®´µ®¶·³̧ ©¹º®»¨©¼·½®¾¬²¬©®¿½«À©¨¹«¬Áº®§·¨À²³³«¹µ®Â³©²¹©®°·½¬²°¬®¬±©
o7 ¾·«³®²½ª®Ã²¬©¨®§·½¹©¨À²¬«·½®¾·°«©¬Á®²¬®ÄÅƹǹȰ¹µ·¨¼®É·¨®²¹¹«¹¬²½°©®È«¬±
5678 p66 q °·ÄÁ¨«¼±¬©ª®Ê°¨©ª«¬©ªË®«¸²¼©¹µ
p66 q r 5678 L8
aỲRNb`NcỲPdQ`[SRYQ
5678
7
5678 p7
5678 s6
6t6 ]Q`[SRYQPN[P[\SPbORQ`SPNPb[RQgbPNPOg^QZP\_c\QS
5678 u
6
5678 G
66 §¨©ª«¬®Ìµ®§²¸ÄÆ©³³µ®Í½®Ìµ®§²¸ÄÆ©³³µ®ÎÏÐѵ®wZQg[PaỲRNdYNZN^_µ®ÒªÈ²¨ª®Ó¨½·³ªÔ
v
6u Õ·½ª·½µ®Âµ®ÎÖϵ®Ì©Ä¨«½¬©ª®È«¬±®¬±©®Ä©¨¸«¹¹«·½®·É®§²¸Æ¨«ª¼©®¿½«À©¨¹«¬Á®Â¨©¹¹µ
³©²¹©®°·½¬²°¬®¬±©®¾·«³®²½ª®Ã²¬©¨®§·½¹©¨À²¬«·½®¾·°«©¬Á®²¬®ÄÅƹǹȰ¹µ·¨¼®É·¨
²¹¹«¹¬²½°©®È«¬±®°·ÄÁ¨«¼±¬©ª®Ê°¨©ª«¬©ªË®«¸²¼©¹µ
]Q`[SRYQPeN[PfOg^QZ
T_c\QS
]Q`[SRYQPQZZPYg[NPNORPOg`[YNgQZP^RNOcbPaNb[PQRSPS`NVcNbSRbP[\Q[P`NgbOVSPbYVcZSP`QRdNgP`NVcNOgbPbO`\PQb
RNN[PS×OQ[SbPQgPRSb\PcZQg[PZY[[SRP]_P[\YbPcRN`SbbPdQ`[SRYQP`Ng¥SR[PSgSR^_PYgPbNYZPNR^QgỲPVQ[[SRPYg[NPNRVb
ObSOZP[NP[\SPRSb[PNP[\SPNR^QgYbVbPYgP[\SPbNYZPNNPSdP¤PgOVdSRPNPS`NVcNbSRbP`QgPdRSQ¢PNgPcSb[ỲYSbPQg
cNZZO[Qg[bPYgPbNYZPeS`NVcNbSRbPQRSPSbcS`YQZZ_PYVcNR[Qg[PYgPYVVNdYZY£Yg^PNRPRS[QYgYg^PgO[RYSg[bPYgP[\SYRP`SZZbP[\Ob
cRS¥Sg[Yg^P[\SPZNbbPNPgO[RYSg[bPbO`\PQbPgY[RN^SgPRNVP[\SPRNN[Yg^P£NgS
XNYZP]YNZN^_PwRYVSRPw\N[NPxQZZSR_ ¤PbS`NgP^RNOcPNPdQ`[SRYQPQRSP[\SP¸Å¬Å²³«¹¬¹P[\Q[PNRVPcQR[gSRb\YcbPY[\PcZQg[bP¦\SPVNb[PSZZ¢gNgPNP[\SbS
QRSP[\SPgY[RN^SgY×Yg^PdQ`[SRYQP¦\SP[\YRP^RNOcPNPdQ`[SRYQPYbP[\SPIJ¬±·¼©½¹P]Q`[SRYQZPcQ[\N^SgbPYg`ZOS
ØÁ¸·¸·½²¹PQgPҨȫ½«²PbcS`YSbPQgPbcS`YSbPNPÓ¼¨·Æ²°¬©¨«Å¸P[\Q[P`QObSP^QZZPNRVQ[YNgPYgPcZQg[bP¤PNOR[\
^RNOcP`QZZSP³«¬±·¬¨·Ä±¹PNRP°±©¸·²Å¬·¬¨·Ä±¹PNd[QYgbPY[bPSgSR^_PRNVP`NVcNOgbPNPgY[RN^SgPbOZORPYRNgPNR
\_RN^SgPYgb[SQPNPRNVP`QRdNgP`NVcNOgbPXNVSPNP[\SbSPbcS`YSbPQRSPYVcNR[Qg[P[NPgY[RN^SgP`_`ZYg^PQg
S^RQQ[YNgPNPcNZZO[Qg[bPP
ÙÚ}Ûz}~{}ÛzÜ
]Q`[SRYQPRNVPQZZPNORP^RNOcbPcSRNRVPYVcNR[Qg[PbSR¥ỲSbPRSZQ[SP[NPQ[SRP_gQVỲbPgO[RYSg[P`_`ZYg^PQgPYbSQbS
bOccRSbbYNgPXNVSPdQ`[SRYQPQS`[PQ[SRPVN¥SVSg[Pd_PcRNO`Yg^PbOdb[Qg`SbP[\Q[P\SZcPdYgPbNYZPcQR[ỲZSbPYg[N
bVQZZPQ^^RS^Q[SbP[\NbSPY[\PYQVS[SRbPNPWWPNPQgPYg`\PNRPÝÝV PX[QdZSPQ^^RS^Q[SbPYVcRN¥S
Q[SRPYgYZ[RQ[YNgPQgP[\SPbNYZÞbPQ[SR\NZYg^PQdYZY[_PgPQPY¥SRbSPdQ`[SRYQZP`NVVOgY[_PVQg_PNR^QgYbVbPYZZ
`NVcS[SPY[\PYbSQbS`QObYg^PNR^QgYbVbPYgPRNN[bPQgPNgPQdN¥S^RNOgPbORQ`SbPNPcZQg[b
}ßà}áâzã}~{
{zäãåæ{ç{ãä}è{}NRVPb_VdYN[ỲPQbbN`YQ[YNgbPY[\P[\SPRNN[bPNPZS^OVSbPZY¢SP`ZN¥SRPQgPZOcYgSPQgP[RSSb
bO`\PQbPQZSRPQgPZN`Ob[PéYbYdZSPgNOZSbPQRSP`RSQ[SP\SRSPdQ`[SRYQPYgS`[PQP^RNYg^PRNN[P\QYRP¦\SPcZQg[PbOccZYSb
bYVcZSP`QRdNgP`NVcNOgbP[NP[\SPdQ`[SRYQPQgP[\SPdQ`[SRYQP`Ng¥SR[PgY[RN^SgPêÝ PRNVPQYRPYg[NPQPNRVP[\SPcZQg[
XNYZPTSQZ[\ \Nb[P`QgPObSP¡\SgPZSQ¥SbPNRPRNN[bPRNVP[\SP\Nb[PcZQg[PS`NVcNbSPbNYZPgY[RN^SgPYg`RSQbSbPYgP[\SPbORRNOgYg^
QRSQ
{{æë{ãä}è{}`\Qg^SPQVVNgYOVPêTì P[NPgY[RY[SPêíÝ P[\SgP[NPgY[RQ[SPêíî PPQPcRSSRRSPNRVPN
gY[RN^SgPNRP^RQbbSbPQgPVNb[PRNP`RNcbPêY[RQ[SPYbPZSQ`\SPVNRSPSQbYZ_PRNVP[\SPbNYZPbNPbNVSPQRVSRbPObS
gY[RYỲQ[YNgPYg\YdY[NRbP[NPRSO`SP[\SPQ`[Y¥Y[_PNPNgSP[_cSPNPgY[RY_Yg^PdQ`[SRYQPêY[RY_Yg^PdQ`[SRYQPQRSPbOccRSbbSPYg
NRSb[PbNYZbPbNP[\Q[PVNb[PNP[\SPgY[RN^SgPRSVQYgbPQbPQVVNgYOV
116116
811
7888167818
17686!1"7#$%33&4'()23 $13
01213434 5678
7 5 5678
>?@ABCADEA@FGHIJB?CAIG4.8K67528,5735625.28,57.L682MNOP2.728,57.0;2.Q,962MNORP2L3;S2T68,57,U,67;2376238367./,4V
1638,8L25+6=23762345,K62W+6762.Q=L682,;23/;685V2;04+23;2,82;350735692;.,:;2.72,8;,962;.,:23LL76L356;S
XJBA@YZEJ?B?[G376232:37L62L7.0\2.U2/34567,325+352L7.W23;2+=\+362:,]62U08L,S2^+6=2376276;\.8;,/:62U.725+6
4+3734567,;5,43::=2_6375+=`2;16::2.U2U76;+:=2507869V2+63:5+=2;.,:S2<45,8.1=4656;2964.1\.;6232W,9623773=2.U
;0/;57356;V2/05237626;\64,3::=2,1\.753852,8296L739,8L27643:4,573852M+379a5.a964.1\.;6P24.1\.089;V2;04+23;24+,5,8
389246::0:.;6V238923762345,K62352+,L+2\b2:6K6:;S2c08L,237621.762,1\.753852,8296L739,8L25+6;624.1\.089;2352:.W
\bS2<2801/672.U2385,/,.5,4;23762\7.904692/=2345,8.1=4656;2;04+23;2d576\5.1=46;S
N.90:6;2U.71692W+6762*+,-./,012/34567,32,8U645692;.=/63827..5;S
efghijklmjgnogplqgrnsgtlugvlwgxiyzlmj{jgl|pi}gf~ijlsg{~glyzpj{yjljoglmzis
{phl{jgflezp~gf}{jizplmzyigjl{jln~
~vy~zflzfl{~~i~j{pyglvijo
yznfiojghlyfghijghlir{g~
*+,-./,012/34567,3
8.90:6;
<45,8.1=4656;V2;04+23;25+,;2d576\5.1=46;V2L,K62;.,:2,5;26375+=2;16::S
efghijkluzllfzrld.,:2,47./,.:.L=23892,.4+61,;57=2d:,962d65ll
w{fjiptlgjl{stlgh~lmmmtlw{hi~zptllsg{~glyzpj{yjljoglmzisl{phl{jgf
ezp~gf}{jizplmzyigjl{jln~
~vy~zflzfl{~~i~j{pyglvijolyznfiojgh
<45,8.1=4656; yfghijghlir{g~
?C?GXC?GIJB?CAI
37,.0;2;\64,6;2.U2/34567,325+7,K62.829,UU676852U..92;.0746;23892,829,UU6768521,47.68K,7.81685;S282L68673:V
/34567,3237621.7624.1\65,5,K62W+682:3/,:62M63;=a5.a1653/.:,-6P2;0/;57356;23762\76;685S2^+,;2,84:096;2U76;+V2=.08L
\:385276;,906238925+624.1\.089;2U.089286372:,K,8L27..5;S234567,3237626;\64,3::=24.84685735692,825+627+,-.;\+676V
5+628377.W276L,.8286Q525.23892,825+627..5S2^+6762,;26K,9684625+352\:385;2\7.9046246753,825=\6;2.U27..526Q09356;25.
684.073L625+62L7.W5+2.U2\7.5645,K62/34567,3S
34567,323:56725+62;.,:268K,7.8168525.25+626Q568525+3525+62;.,:268K,7.816852W,::2U3K.7246753,82\:38524.1108,5,6;
.K672.5+67;S26U.762\:385;24382/64.1626;53/:,;+692.82U76;+2;69,1685;V25+62/34567,3:24.1108,5=210;526;53/:,;+
U,7;5V2;5375,8L2W,5+2\+.5.;=85+65,42/34567,3S2^+6;62U,Q2351.;\+67,428,57.L6823892437/.8V2\7.90462.7L38,42135567V
3892,11./,:,-6268.0L+28,57.L6823892.5+6728057,685;25.2,8,5,35628,57.L6824=4:,8L2\7.46;;6;2,825+62=.08L2;.,:S2^+68V
637:=2;0446;;,.83:2\:3852;\64,6;24382L7.WS2<;25+62\:38524.1108,5=2,;26;53/:,;+69V29,UU6768525=\6;2.U2.7L38,4
13556726856725+62;.,:238924+38L625+625=\62.U2U..923K3,:3/:625.2/34567,3S2825078V25+623:567692/34567,3:24.1108,5=
4+38L6;2;.,:2;57045076238925+6268K,7.816852U.72\:385;S2d.16276;6374+67;25+,8]2,5213=2/62\.;;,/:625.24.857.:25+6
\:3852;\64,6;2,8232\:3462/=21383L,8L25+62;.,:2/34567,324.1108,5=S
FGAYFCIEGIJB?CAIGIBGCYZYB?G I@BG¡CY¢BGG
£lppl¤gppghtl|m¥lfiysjf{sl¦g~g{fyolmgf}iygtlssr{ptlll
§6753,82;573,8;2.U25+62;.,:2/34567,32¨;609.1.83;2U:0.76;468;2+3K62385,aU08L3:2345,K,5=25+352,8+,/,5;2;.162\:385
\35+.L68;S2¨S2U:0.76;468;23892.5+672¨;609.1.83;23892©385+.1.83;2;\64,6;24382,84763;62\:3852L7.W5+2,82;6K673:
W3=;S2^+6=213=2\7.90462324.1\.08925+352,8+,/,5;25+62L7.W5+2.U2\35+.L68;2.72769046;2,8K3;,.82.U25+62\:3852/=23
\35+.L68S2^+6=213=23:;.2\7.904624.1\.089;2ML7.W5+2U345.7;P25+3529,7645:=2,84763;62\:3852L7.W5+S22
^+6;62\:3852L7.W5+a68+384,8L2/34567,32.44072835073::=2,82;.,:;V2/0528.523:W3=;2,82+,L+268.0L+2801/67;25.2+3K623
973135,426UU645S2825+62U05076V2U37167;213=2/623/:625.2,8.40:3562;669;2W,5+2385,aU08L3:2/34567,3V2;04+23;2¨S
U:0.76;468;V25.268;07625+3525+62/34567,327690462\35+.L68;237.08925+62;669238927..52.U25+6247.\S
N*§d2b.162ª2«dT<SL.K2ª2d,5623\2ª2§,K,:2*,L+5;2ª2cR<2ª2̈:3,82¬7,5,8L2ª2<446;;,/,:,5=2d53561685
¨.:,4=23892,8];ª2N.8aT,;47,1,835,.82d535616852ª28U.7135,.82®03:,5=2ª2«d<SL.K2ª2¬+,56b.0;6SL.K
116116 811 7888167818 17686!1"7#$%33&4'()23 313