Italian Botanist 5: 45–56 (2018)
doi: 10.3897/italianbotanist.5.25910
Notulae to the Italian alien vascular flora: 5
RESEARCH ARTICLE
45
http://italianbotanist.pensoft.net
Notulae to the Italian alien vascular flora: 5
Gabriele Galasso1, Gianniantonio Domina2, Michele Adorni3,
Nicola M.G. Ardenghi4, Gianmaria Bonari5, Sergio Buono6, Laura Cancellieri7,
Giuseppina Chianese8, Giulio Ferretti9, Tiberio Fiaschi10, Luigi Forte11,
Riccardo Guarino12, Rocco Labadessa13, Lorenzo Lastrucci9, Lorenzo Lazzaro9,
Sara Magrini14, Luigi Minuto15, Sara Mossini16, Nicola Olivieri17, Anna Scoppola7,
Adriano Stinca18, Claudia Turcato15, Chiara Nepi19
1 Sezione di Botanica, Museo di Storia Naturale di Milano, Corso Venezia 55, 20121 Milano, Italy 2 Dipartimento di Scienze Agrarie, Alimentari e Forestali (SAAF), Università di Palermo, Viale delle Scienze, ed. 4,
90128 Palermo, Italy 3 Via degli Alpini 7, 43037 Lesignano de’ Bagni (Parma), Italy 4 Dipartimento di Scienze della Terra e dell’Ambiente, Università di Pavia, Via Sant’Epifanio 14, 27100 Pavia, Italy 5 Department
of Botany and Zoology, Masaryk University, Kotlarska 2, CZ-611 37 Brno, Czech Republic 6 Via XXV Aprile
6, 01010 Oriolo Romano (Viterbo), Italy 7 Dipartimento di Scienze Agrarie e Forestali (DAFNE), Università
della Tuscia, Via San Camillo de Lellis snc, 01100 Viterbo, Italy 8 Musei delle Scienze Agrarie, Università di
Napoli Federico II, Via Università 100, 80055 Portici (Napoli), Italy 9 Dipartimento di Biologia, Università
di Firenze, Via G. La Pira 4, 50121 Firenze, Italy 10 Dipartimento di Scienze della Vita, Università di Siena,
Via P.A. Mattioli 4, 53100 Siena, Italy 11 Dipartimento di Biologia e Museo Orto Botanico, Università di
Bari Aldo Moro, Via E. Orabona 4, 70125 Bari, Italy 12 Sezione di Botanica ed Ecologia Vegetale, Dipartimento di Scienze e Tecnologie Biologiche, Chimiche e Farmaceutiche (STEBICEF), Università di Palermo,
Via Archirafi 20, 90123 Palermo, Italy 13 Associazione Centro Studi de Romita, Via G. Postiglione 9, 70126
Bari, Italy 14 Banca del Germoplasma della Tuscia, Università della Tuscia, Largo dell’Università snc, blocco
c, 01100 Viterbo, Italy 15 Dipartimento di Scienze della Terra, dell’Ambiente e della Vita (DISTAV), Università di Genova, Corso Dogali 1/m, 16136 Genova, Italy 16 Via E. Rovati 6, 27049 Stradella (Pavia), Italy
17 Via Maestri del Lavoro 40, 64100 Teramo, Italy 18 Dipartimento di Scienze e Tecnologie Ambientali,
Biologiche e Farmaceutiche, Università della Campania Luigi Vanvitelli, Via A. Vivaldi 43, 81100 Caserta,
Italy 19 Sezione di Botanica Filippo Parlatore, Museo di Storia Naturale, Università di Firenze, Via G. La
Pira 4, 50121 Firenze, Italy
Corresponding author: Gabriele Galasso (
[email protected])
Academic editor: Lorenzo Peruzzi | Received 17 April 2018 | Accepted 27 April 2018 | Published 14 May 2018
Citation: Galasso G, Domina G, Adorni M, Ardenghi NMG, Bonari G, Buono S, Cancellieri L, Chianese G, Ferretti G,
Fiaschi T, Forte L, Guarino R, Labadessa R, Lastrucci L, Lazzaro L, Magrini S, Minuto L, Mossini S, Olivieri N, Scoppola
A, Stinca A, Turcato C, Nepi C (2018) Notulae to the Italian alien vascular flora: 5. Italian Botanist 5: 45–56. https://doi.
org/10.3897/italianbotanist.5.25910
Copyright Gabriele Galasso et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
46
Gabriele Galasso et al. / Italian Botanist 5: 45–56 (2018)
Abstract
In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented.
It includes new records, and confirmations for Italy or for Italian administrative regions of taxa in the
genera Albizia, Anredera, Bougainvillea, Cardamine, Cenchrus, Cephalaria, Ceratochloa, Cytisus, Datura,
Delosperma, Euonymus, Freesia, Hylotelephium, Lantana, Musa, Physalis, Rotala, Styphnolobium, Trachycarpus, and Tradescantia. Nomenclature and distribution updates, published elsewhere, and corrections are
provided as supplementary material.
Keywords
Alien species, floristic data, Italy, nomenclature
How to contribute
The text for the new records should be submitted electronically to Chiara Nepi (chi
[email protected]). The corresponding specimen along with its scan or photograph
has to be sent to FI Herbarium: Sezione di Botanica Filippo Parlatore del Museo
di Storia Naturale, Via G. La Pira 4, 50121 Firenze (Italy). Those texts concerning
nomenclatural novelties (typifications only for accepted names), status changes, ex
clusions, and confirmations should be submitted electronically to: Gabriele Galasso
(
[email protected]). Each text should be within 2,000 characters
(spaces included).
Floristic records
Albizia julibrissin Durazz. (Fabaceae)
+ (CAS) CAL: Montegiordano (Cosenza), fraz. Montegiordano Marina, massicciata
ferroviaria presso Via Canale G. Garibaldi (WGS84: 40.031813°N; 16.600047°E),
massicciata ferroviaria, ca. 10 m, 21 August 2017, N. Olivieri (FI). – Casual alien spe
cies new for the flora of Calabria.
A young specimen of the species grows on pebbly ground at the railway roadbed
crossing the village’s suburban area near an overpass. It originated from the seeds
produced by some trees cultivated along a neighboring road. The native range of
Albizia julibrissin includes southern Caucasus, Iran, Pakistan, India, Nepal, Bangla
desh, Sri Lanka, Myanmar, China, Korea, Japan, Ethiopia, Eritrea, and Somalia. The
species was introduced in Italy for ornamental purposes in 1745 (Targioni Tozzetti
1896) and it is currently widely planted as ornamental in parks and gardens around
the world.
N. Olivieri
Notulae to the Italian alien vascular flora: 5
47
Anredera cordifolia (Ten.) Steenis (Basellaceae)
+ (CAS) MAR: Cupra Marittima (Ascoli Piceno), presso la stazione ferroviaria
(WGS84: 43.023750°N; 13.860402°E), incolto, ca. 6 m, 3 August 2017, N. Olivieri
(FI). – Casual alien species confirmed for the flora of Marche.
At Cupra Marittima, this species grows on sandy soil at the edge of a small neglected
area, along with Broussonetia papyrifera (L.) Vent. and Ailanthus altissima (Mill.) Swin
gle. The species was no longer recorded in the Marche after 1950 (Galasso et al. 2018).
According to Viegi et al. (2004), reporting BrilliCattarini’s personal communications,
Anredera cordifolia was previously observed in Ancona in 1945, and in Pesaro in 1939.
N. Olivieri
Bougainvillea spectabilis Willd. (Nyctaginaceae)
+ (CAS) ABR: Pescara (Pescara), presso Viale V. Pepe (WGS84: 42.458347°N;
14.231300°E), bordo di marciapiede, ca. 3 m, NE, 9 August 2017, N. Olivieri (FI). –
Casual alien species new for the flora of Abruzzo.
A young plant has developed on the inner edge of a sidewalk at the base of the
perimetral wall of a private home in the urban area of Pescara. The location is partially
shaded and at a short distance from the Adriatic Sea, but it is sheltered from sea winds
by the buildings. In Italy, Boungavillea spectabilis is also reported as a casual alien in
Sardegna (Bacchetta et al. 2009, Galasso et al. 2018).
N. Olivieri
Cardamine occulta Hornem. (Brassicaceae)
+ (CAS) LAZ: Roma (Roma), lungo Via del Biscione (WGS84: 41.895524°N;
12.472969°E), aiuola, 18 m, 28 October 2017, A. Stinca (FI, PORUN). – Casual
alien species new for the flora of Lazio.
In Italy, Cardamine occulta is recorded for Piemonte, Lombardia, TrentinoAlto
Adige, Veneto, Friuli Venezia Giulia, EmiliaRomagna, Toscana, Campania, and
Sardegna (Galasso et al. 2018). According to Stinca et al. (2017), this species has been
largely spread by nurseries and soil transportation.
A. Stinca, G. Chianese
Cenchrus setaceus (Forssk.) Morrone (Poaceae)
+ (CAS) TOS: Piombino (Livorno), fraz. Baratti (WGS84: 42.998797°N;
10.516870°E), margine stradale, 8 m, 21 October 2017, T. Fiaschi (FI). – Casual alien
species new for the flora of Toscana.
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Gabriele Galasso et al. / Italian Botanist 5: 45–56 (2018)
The occurrence of this species in Toscana does not seem to be linked to direct hu
man cultivation. Seeds of Cenchrus setaceus are known to have a very efficient wind
dispersal strategy. However, given that the nearest known populations occur a few
hundred kilometres away from Baratti, wind is not the most likely vector. The spe
cies is included in the list of Invasive Alien Species of Union Concern (Commission
Implementing Regulation EU 2016/1141) and was found in three separate nuclei 10
m apart, in a parking area of a renowned tourist location, where seeds could have been
introduced via pet fur or car mats. According to Galasso et al. (2018), it occurs in
Lazio, Puglia, Calabria, Sicilia, and Sardegna.
G. Bonari, T. Fiaschi, R. Guarino
Cephalaria syriaca (L.) Schrad. (Dipsacaceae)
+ (NAT) LAZ: Roma (Roma), Scalo ferroviario Ostiense (WGS84: 41.871550°N;
12.488699°E ± 700 m), ca. 20 m, 28 May 1954, A. Cacciato (RO); Tarquinia (Viter
bo), loc. Pian di Spille, zona militare (WGS84: 42.255734°N; 11.678989°E), incolto
erboso al margine di campo coltivato a fieno greco, 4 m, 17 April 2017, A. Scoppola
(FI, UTV no. 35200); Monte Romano (Viterbo), valle del Fiume Mignone (WGS84:
42.251059°N; 11.877385°E), margine di campi lungo strada sterrata in luogo arido,
109 m, 20 April 2017, A. Scoppola, L. Cancellieri (UTV no. 35202, Herb. L. Cancellieri). – Naturalized alien species confirmed for the flora of Lazio.
Cephalaria syriaca belongs to the MediterraneanTuranian element, growing as a
weed in cereal fields and waste places (Matthews 1972). It is not listed for Lazio by An
zalone et al. (2010), while Lucchese (2017) records the species for this administrative
region, but without providing information on recent records or herbarium specimens.
Accordingly, it has been excluded from the confirmed regional alien flora (Galasso et
al. 2018). It is not reported in the flora of the Pian di Spille coast (Iocchi and Bar
tolucci 2008). However, it was reported by Cacciato (1955) in the Ostiense railway
station in Roma, an area deeply transformed due to the urban development of the last
decades. This ancient discovery is attested by a 1954 herbarium specimen preserved in
RO. However, this record was ignored by Lucchese (2017). In the two new localities,
C. syriaca occurs with a conspicuous number of fertile individuals, spread at the edge
of cultivated fields and waste places.
L. Cancellieri, A. Scoppola
Ceratochloa cathartica (Vahl) Herter (Poaceae)
+ (NAT) PUG: Bari (Bari), foce del Torrente Lamasinata (WGS84: 41.134654°N;
16.827638°E), prateria umida su substrato sabbioso, 1 m, 30 September 2017,
R. Labadessa (FI); Bari (Bari), Torrente Lamasinata (WGS84: 41.134647°N;
Notulae to the Italian alien vascular flora: 5
49
16.827632°E), prati umidi su sabbia, 1 m, 30 September 2017, R. Labadessa (BI no.
40486, no. 40487). – Naturalized alien species new for the flora of Puglia.
Ceratochloa cathartica is an alien plant from South America, whose European dis
tribution ranges from Portugal to Ukraine, and from Great Britain to Italy (Ryves et
al. 1996). This species was probably introduced as fodder in Europe, where it may be
found as naturalized in the warmer regions (Ryves et al. 1996). Several individuals were
found within an area of about two hectares, in a wet sandy meadow dominated by an
nual and perennial grass species.
R. Labadessa, L. Forte
Cytisus striatus (Hill) Rothm. (Fabaceae)
+ (NAT) ITALIA (LIG): Noli (Savona), fraz. Tosse, fra la SP8 e l’Autostrada dei Fiori
A10 (WGS84: 44.227616°N; 8.393363°E), macchia mediterranea, 177 m, 20 June
2017, leg. L. Minuto, det. C. Turcato (FI, GE). – Naturalized alien species new for the
flora of Italy (Liguria).
Cytisus striatus is native to Morocco, Portugal, and Spain. It has been introduced
into a number of northwestern European countries (England, Scotland, Wales, and
France), and in the Americas (California and Oregon in the U.S.A., and Argentina),
where it is considered to be an invasive shrub. A large population, identified follow
ing the key published by Frodin and Heywood (1968), was detected growing on the
motorway embankment. Other plants were observed along the Autostrada A10 from
Albenga to Savona Vado, and along the Autostrada A12 near Sestri Levante (loc. Roc
che di Sant’Anna).
C. Turcato, L. Minuto
Datura wrightii Regel (Solanaceae)
+ (CAS) LAZ: Bracciano (Roma), fraz. Vigna di Valle, Museo Storico dell’Aeronautica
Militare, presso l’Aeroporto di Vigna di Valle (WGS84: 42.085366°N; 12.218396°E),
suolo sabbioso presso il lago, 158 m, 16 July 2017, S. Buono (FI, UTV). – Casual alien
species new for the flora of Lazio.
Datura wrightii is a perennial plant native to the southwestern United States and
Mexico. In southern Europe, it was widely confused with the closely related species
D. inoxia Mill. (Verloove 2008). Reports of the occurrence of this species as casual or
naturalized alien in some Italian regions are very recent (Banfi and Galasso 2010, Ver
loove et al. 2010, Ardenghi et al. 2011, Del Guacchio 2011, Cerutti and Motta 2012,
D’Aleo and Bonanno 2016, Galasso et al. 2018). A single individual was observed
growing on sandy soil near Lake Bracciano, along with other alien species as Abutilon theophrasti Medik., Ludwigia peploides (Kunth) P.H.Raven subsp. montevidensis
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Gabriele Galasso et al. / Italian Botanist 5: 45–56 (2018)
(Spreng.) P.H.Raven, Pavonia hastata Cav. (see also Galasso et al. 2017), and Physalis
peruviana L. (see beyond in this contribution). The plant regularly developed a high
number of flowers and fruits from July to December 2017.
S. Buono, S. Magrini, A. Scoppola
Delosperma cooperi (Hook.f.) L.Bolus (Aizoaceae)
+ (CAS) EMR: Castel San Giovanni (Piacenza), Via Fratelli Bandiera (SP10R), angolo
con Via Bottarone (WGS84: 45.06085°N; 9.43153°E), ciglio stradale, con Setaria
italica subsp. viridis e Lactuca sativa subsp. serriola, 75 m, 3 September 2017, N.
Ardenghi, S. Mossini (FI). – Casual alien species new for the flora of EmiliaRomagna.
A single flowering individual was found in the growing site.
N.M.G. Ardenghi, S. Mossini
Euonymus japonicus Thunb. (Celastraceae)
+ (CAS) ABR: Pescara (Pescara), aiuola lungo Viale T. Patini (WGS84: 42.452488°N;
14.241713°E), epifita su stipite di Phoenix canariensis, ca. 4 m, 29 October 2017, N.
Olivieri (FI). – Casual alien species new for the flora of Abruzzo.
A young individual of this species grows as an epiphyte among the stumps of the cut
leafy rachis of a Phoenix canariensis H.Wildpret, at about 1.5 m from the ground. The
settlement site is located in a coastal, partially shaded, suburban area, not far from the
Adriatic Sea. In the surrounding gardens, Euonymus japonicus is cultivated as ornamen
tal and the recorded plant may have originated from seeds dispersed by ornithochory.
N. Olivieri
Freesia alba (G.L.Mey.) Gumbl. (Iridaceae)
+ (CAS) ABR: San Vito Chietino (Chieti), fraz. Marina di San Vito (WGS84:
42.307275°N; 14.446527°E), prato presso giardino privato, ca. 20 m, 29 October
2017, N. Olivieri (FI). – Casual alien species new for the flora of Abruzzo.
A young individual of the species has developed in a meadow near a private gar
den where the species is cultivated. It grows on sandypelitic soil, dry in summer. The
locality is close to the Adriatic Sea, and has a Mediterranean climate, but it is exposed
to damp atmospheric currents coming from the sea. Freesia alba and its horticultural
hybrids with F. corymbosa N.E.Br. and F. leichtlinii Klatt, erroneously attributed to F.
refracta (Jacq.) Klatt (Goldblatt and Manning 2008, Galasso et al. 2018), are bulbous
plants of South African origin cultivated as ornamentals, even outdoors and in the
ground along the Abruzzo coasts.
N. Olivieri
Notulae to the Italian alien vascular flora: 5
51
Hylotelephium spectabile (Boreau) H.Ohba (Crassulaceae)
+ (CAS) MAR: Pesaro (Pesaro e Urbino), Viale della Liberazione, Mura roveresche
(WGS84: 43.90999°N; 12.90483°E), parete in mattoni, con Capparis orientalis, Parietaria judaica, Convolvulus sepium, 28 August 2017, N. Ardenghi, S. Mossini (FI). –
Casual alien species new for the flora of Marche.
Two individuals (one of which with ripening fruits) were observed on the ancient
city walls of Pesaro, probably originating from the dissemination of cultivated plants
in the nearby dwellings.
N.M.G. Ardenghi, S. Mossini
Lantana camara L. subsp. aculeata (L.) R.W.Sanders (Verbenaceae)
+ (CAS) PUG: Ugento (Lecce), fraz. Torre San Giovanni, lungo un canale artificiale
presso la costa ionica (WGS84: 39.875752°N; 18.146011°E), vegetazione disturbata
lungo un canale, ca. 4 m, 24 August 2017, N. Olivieri (FI). – Casual alien subspecies
new for the flora of Puglia.
Some individuals grow near an artificial channel along with Phragmites australis
(Cav.) Trin. ex Steud. subsp. australis in a flat coastal area, on a red soil with good
water availability, characterized by disturbed vegetation and partially shaded by some
Eucalyptus camaldulensis Dehnh. subsp. camaldulensis. The previous report for Puglia
(Olivieri 2012) of Lantana camara L. for the same area of Ugento has to be referred to
the same taxon recorded here. L. camara subsp. aculeata is widely cultivated for orna
mental purposes and naturalized in the tropics and subtropics; in Italy it is reported in
Abruzzo, Molise, Basilicata, Calabria, Sicilia, and Sardegna (Galasso et al. 2018).
N. Olivieri
Lantana depressa Small (Verbenaceae)
+ (CAS) PUG: Taranto (Taranto), presso la stazione ferroviaria (WGS84: 40.484680°N;
17.223055°E), incolto, ca. 17 m, S, 23 August 2017, N. Olivieri (FI). – Casual alien
species new for the flora of Puglia.
One individual of the species grows at the base of the steps leading to an old
abandoned building near the railway embankment, not far from the railway station. It
has developed inside a fissure at the base of the rise of a partially eroded concrete step,
in a context of ruderal vegetation dominated by Dittrichia viscosa (L.) Greuter subsp.
viscosa. Lantana depressa is native to southern Florida and has also been reported in
Sicilia (Galasso et al. 2018). According to Sanders (2012), the individual belongs to L.
depressa var. depressa, originally widespread along the limestone outcrop of the Miami
Rock Ridge.
N. Olivieri
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Gabriele Galasso et al. / Italian Botanist 5: 45–56 (2018)
Musa basjoo Siebold & Zucc. ex Iinuma (Musaceae)
+ (NAT) TOS: Montignoso (MassaCarrara), fraz. Cinquale, sommità dell’argine di un
canale in destra del Fiume Versilia, in prossimità di una villetta residenziale (WGS84:
43.986072°N; 10.161817°E), argine di canale, 2 m, 19 July 2017, L. Lastrucci, L. Lazzaro (FI). – Naturalized alien species new for the flora of Toscana.
In the site of collection, the species is present with several individuals, of different
ages. The population appears to have originated by vigorous resprout from pruning
residues. Indeed, the species is widely cultivated in the gardens of the residential areas
near the canals surrounding Lake Porta. Several localized stands of this species have
also been observed in other sites of the MassaCarrara and Lucca provinces.
L. Lastrucci, G. Ferretti, L. Lazzaro
Physalis peruviana L. (Solanaceae)
+ (CAS) LAZ: Bracciano (Roma), fraz. Vigna di Valle, Museo Storico dell’Aeronautica
Militare, presso l’Aeroporto di Vigna di Valle, in riva al lago (WGS84: 42.085373°N;
12.218674°E), riva di lago, 157 m, 28 July 2017, S. Buono (FI, UTV). – Casual alien
species new for the flora of Lazio.
Physalis peruviana is a herbaceous perennial species, which has been very widely
introduced across the world from South America as a cultivated plant for its fruit, as a
medicinal plant, and as an ornamental (CABI 2018). It is reported as a casual alien in
several administrative regions, especially in northern Italy, and as naturalized in Sicilia
(Galasso et al. 2018). It is classified as an invasive plant at the global level (Global
Invasive Species Database 2018). Some individuals of this species grow on sandy soil
near Lake Bracciano, along with Abutilon theophrasti Medik, Datura wrightii Regel (see
a previous record in this contribution), Ludwigia peploides (Kunth) P.H.Raven subsp.
montevidensis (Spreng.) P.H.Raven, Pavonia hastata Cav. (see also Galasso et al. 2017),
Portulaca oleracea L., Solanum nigrum L., and other alien species. The plants regularly
develop flowers and fruits.
S. Buono, S. Magrini, A. Scoppola
Rotala ramosior (L.) Koehne (Lythraceae)
+ (NAT) EMR: Colorno (Parma), golena del Po presso la fraz. Sacca (WGS84:
44.976183°N; 10.383083°E), fanghi di lanca, 25 m, 20 August 2017, M. Adorni (FI).
– Naturalized alien species new for the flora of EmiliaRomagna.
In Galasso et al. (2018), Rotala ramosior is reported only for Piemonte, Lombar
dia, where the species grows in rice fields (Banfi and Galasso 2010), and for Veneto
along the River Po (Masin and Scortegagna 2012). The population from Sacca, con
Notulae to the Italian alien vascular flora: 5
53
sisting of several dozens of plants growing on muddy and damp soil, was also reported
in the Acta Plantarum Forum (http://www.floraitaliae.actaplantarum.org/viewtopic.
php?t=98599).
M. Adorni
Styphnolobium japonicum (L.) Schott (Fabaceae)
+ (CAS) PUG: Foggia (Foggia), Villa Comunale Parco Karol Wojtyla (WGS84:
41.555366°N; 15.186515°E), ca. 64 m, 19 August 2017, N. Olivieri (FI). – Casual
alien species new for the flora of Puglia.
Some young individuals of the species grow along the perimetral wall of the Villa
Comunale, near Via Galliani, on a calcareous alluvial vertisol subjected to partial
desiccation of the herbaceous vegetation in summer, partially shaded by Styphnolobium
japonicum and Pinus halepensis Mill. subsp. halepensis.
N. Olivieri
Trachycarpus fortunei (Hook.) H.Wendl. (Arecaceae)
+ (CAS) MAR: Pesaro (Pesaro e Urbino), Viale C. Battisti (WGS84: 43.91577°N;
12.91205°E), aiuola con Acer pseudoplatanus coltivato, 4 m, un individuo, 28 Au
gust 2017, N. Ardenghi, S. Mossini (FI). – Casual alien species new for the flora of
Marche.
A single, aged individual was found in a public flowerbed, probably grown from
seeds originated from cultivated plants in the surroundig public and private gardens.
N.M.G. Ardenghi, S. Mossini
Tradescantia fluminensis Vell. (Commelinaceae)
+ (CAS) MAR: Pesaro (Pesaro e Urbino), Viale della Liberazione, Mura roveresche
(WGS84: 43.90999°N; 12.90483°E), parete in mattoni, con Capparis orientalis, Parietaria judaica, Convolvulus sepium, 28 August 2017, N. Ardenghi, S. Mossini (FI). –
Casual alien species new for the flora of Marche.
A group of plants, without flowers or fruits, was found on the ancient city walls of
Pesaro, probably deriving from the dissemination of cultivated plants in the surround
ing dwellings.
N.M.G. Ardenghi, S. Mossini
54
Gabriele Galasso et al. / Italian Botanist 5: 45–56 (2018)
Nomenclature and distribution updates from other literature sources
Nomenclature, status, and distribution updates according to Béguinot (1903), Devesa
(2007), Kilian et al. (2009+), Göktürk and Sümbül (2014), Liu et al. (2017), Banfi
(2018), Güzel et al. (2018), and Martini and Viciani (2018), and corrections to Ga
lasso et al. (2018) are provided in Supplementary material 1.
G. Galasso
Acknowledgements
We gratefully acknowledge colleagues who provided distribution, nomenclatural and
taxonomic advices: Alessandro Alessandrini, Enrico Banfi, Fabrizio Bartolucci, Giordano
Martini, and Daniele Viciani.
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Supplementary material 1
Supplementary data
Edited by: Gabriele Galasso
Data type: species data
Explanation note: 1. Nomenclature updates; 2. Status/Note updates; 3 Distribution
updates; 4. Synonyms, misapplied or included names.
Copyright notice: This dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/italianbotanist.5.25910.suppl1