<i>Trechus isabelae</i> Borges &amp; Serrano, new species (Figs. 1–3) <b>Ty... more <i>Trechus isabelae</i> Borges &amp; Serrano, new species (Figs. 1–3) <b>Type material:</b> Holotype, male, Azores, São Jorge, Algar do Morro Pelado (= Algar do Montoso), 5– 11.VII. 2004. (P.A.V. Borges &amp; F. Pereira leg.). Paratypes 1 female, same locality as holotype, 7–11.V. 2004 (P.A.V. Borges &amp; F. Pereira leg.); 2 males and 1 female, same locality as holotype, 5–11.VII. 2004 (P.A.V. Borges &amp; F. Pereira leg.); The Holotype and two paratypes (1 male, 1 female) are deposited at the University of the Azores, Terceira ("Arruda Furtado" Collection); one paratype (female) is deposited at the Faculty of Sciences of the University of Lisbon, and the other one (male) at the University of La Laguna, Dep. of Animal Biology. <b>Additional material:</b> One male was collected in the same locality (7–11.V. 2004) for molecular analysis. <b>Etymology.</b> The species is named after Isabel Amorim who has discovered several new species of <i>Trechus</i> in the Azores. <b>Diagnosis.</b> Large species (4.8–5.38 mm) (Fig. 1), dark brown pigmentation, with some lighter parts; appendages less pigmented (light brown), tibiae and tarsomeres with the same colour patterns as the femora; pronotum as dark as elytra. Eyes slightly convex; 3 rd antennomere longer than 2 nd; pronotum rather cordiform and transverse, with lateral furrows larger than in <i>T. terrabravensis</i>; wingless. <b>Description.</b> Length of holotype: 5.18 mm. Length of female specimens: 4.80–5.38 mm. Large size, convex and ovate. Head almost 1.30 x as wide as long (females: 1.29–1.42), 0.78 x narrower than pronotum (females: 0.74–0.78); eyes well-developed (0.025 mm), moderately convex, diameter of eyes as long as temples; temples with microchaetae. Frons slightly convex, the frontal furrows deeply curved and rounded. Anterior margin of labrum very concave. Maxillary palpi long and slender. Chaetotaxy: two pairs of supraocular setae; four setae in clypeus (outer ones larger); six setae on labrum, the outer ones being the largest. Antennae 2.99 mm long (f [...]
Genus<i> Oplostomus</i> MacLeay, 1838 The genus was not revised. There are ten recogn... more Genus<i> Oplostomus</i> MacLeay, 1838 The genus was not revised. There are ten recognized species ( Holm & Marais 1992, Krajcik 2008a), three of them already recorded for Angola ( Ferreira 1965).
Genus<i> Mausoleopsis</i> Lansberg, 1882 Holm & Marais (1992) and Antoine (1997) refe... more Genus<i> Mausoleopsis</i> Lansberg, 1882 Holm & Marais (1992) and Antoine (1997) refer that the genus is justified itself by the prominently protruding last abdominal spiracles (="élévation des stigmates propygidiaux"). Among the twelve species recognized in the nominotypical subgenus ( Beinhundner 2017a), there are one species and one subspecies recorded for Angola ( Paulino de Oliveira 1882; Antoine 1989a).
<i> Scaptobius kochi</i> Schein, 1957<b> Distribution</b>: ANG.<b> ... more <i> Scaptobius kochi</i> Schein, 1957<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) NAM; 2) HUI.<b> Historic records:</b> —Ongueria (= Namibe) (NAM) ( Schein 1957); —Hunguéria (= Namibe) (NAM) ( Ferreira 1965).<b> New localities reported</b>: Kipungo ( 40 km W) (coord. not available,? alt., 338) ( HUI), 26.XI.2013, 1 ex., P. Schüle leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> none<b> Remarks.</b> The species is endemic of Angola ( Schein 1957). Biological and ecological data are not available. Presumably it presents the same myrmecophilic habits of the other species of this genus. It is a new species record for the Huíla Province.
<i> Gnathocera cruda gracilis</i> (Janson, 1877) ( Fig. 22a)<b> Distribution<... more <i> Gnathocera cruda gracilis</i> (Janson, 1877) ( Fig. 22a)<b> Distribution</b>: ANG, CON, DOC.<b> Distribution in Angola (Provinces):</b> 1) LNO, UIG.<b> Historic records:</b> — Angola (without locality), Bembe (UIG) ( Janson 1877b); —Dundo (LNO) ( Gomes Alves 1959 sub<i> Gnathocera lurida</i> Janson); —Dundo, Chicapa (LNO), Angola (LNO, without locality) ( Antoine 2002a).<b> Material examined:</b> Angola, 2 ♂, Nº 1907 (blue labels), nº 146a (white labels), 1 ♀, Nº 1907 (blue label), nº 147a (white label), 1 ♂, Nº 1907 (blue label), nº 148a (white label), old collection, box nº IX-78, MZUC; Dundo ( 7º 22´S, 20º 50´E, 1470 m alt., 51) ( LNO), -. III.1949, 5 ♂, 4 ♀, Nº 2132, 2 ♀, Nº 2151, Junta de Investigações Coloniais (Entomologia), IICT, MUHNAC.<b> Remarks.</b> The status of this taxon is very controversial. Its validity was established by Antoine (2002a), a criterion followed by Beinhundner (2017a). However, Krajcik (2008a) as well as Schoolmeesters (2018) keep it as a synonym of<i> G. lurida</i> Janson, 1877 [the name<i> lucida</i> used by Krajcik (2008a) is a misspelling of<i> lurida</i>, since Janson (1877b) did not describe any taxon named<i> lucida</i>].<i> Gnathocera cruda gracilis</i> was described based on one specimen from Angola, but Janson (1877b) referred that "there are also specimens in Mr. Higgins's collection found by Monteiro, at Bembe", a municipality within the Uíge Province. The adult specimens of IICT were collected by light (registered information) probably at the same time of<i> G. trivittata ardoini</i> (see remarks under this taxon). Ecological data are not available for this subspecies.
<i> Pseudinca incoides</i> (J. Thomson, 1860)<b> Distribution</b>: ANG, B... more <i> Pseudinca incoides</i> (J. Thomson, 1860)<b> Distribution</b>: ANG, BEN, CAM, CON, DOC, EQG, GAB, GHA, RCA, RCI, NRE, TOG.<b> Distribution in Angola (Provinces):</b> 1) CAB, ZAI.<b> Historic records:</b> — Cabinda (CAB) ( Paulino de Oliveira 1882 sub<i> Diplognatha incoides</i> Thoms.); —S. Salvador (ZAI) ( Valck Lucassen 1933).<b> Material examined:</b> none.<b> Remarks.</b> The species in Angola seems restricted to its northwesternmost territory. Tchibozo<i> et al</i>. (2008) and Joly (2018) provided biodiversity information (abundance, habitat preference) of this species for Benin and Ghana, respectively.
<i> Coenochilus ventricosus</i> (Gyllenhal, 1817) ( Fig. 12f)<b> Distribution&l... more <i> Coenochilus ventricosus</i> (Gyllenhal, 1817) ( Fig. 12f)<b> Distribution</b>: ANG, CAM, DOC, EQG, GAB, GAM, KEN, MOZ, NAB, RCI, RSA, SEN, SLN, TAN, TOG, ZIM.<b> Distribution in Angola (Provinces):</b> 1) LNO, MAL; 2) CNN.<b> Historic records:</b> —Quango (=Cuango) (LNO) ( Quedenfeldt 1884 and Ferreira 1965); —Malange (MAL) ( Schein 1954a).<b> New localities reported</b>: Cahama ( 20 km NE) (coord. not available, 1315 m alt., 399) ( CNN), 8.XII.2013, 1♂, P. Schüle leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> Cacuso ( 9º 25´ 28.03´´S, 15º 44´ 37.63´´ E, 1078m alt., 112) ( MAL), 16–18.XI.2017, 1♂, DO, A. Serrano leg., ASC; Quitota ( 9º 37´ 32.39´´ S, 15º 25´ 22.57´´ E, 898m alt., 129) ( MAL), 20.XI.2017, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC.<b> Remarks.</b> It is the type species of a group with its own name (<i> C. ventricosus</i> group<i> sensu</i> Schein 1954a). The Cacuso adult specimen was found on the ground during night near public lights. Like some<i> C. procerus</i> specimens, the Cacuso specimen had phoretic acarina on some parts of the body (<i> e.g.</i> legs). The Quitota adult specimen was collected by LT (<i> e.g.</i> Fig. 1c) in an open secondary forest. Joly (2018) provided biodiversity information (abundance, habitat preference) on this species for Ghana. Other biological and ecological data are not available for the species, but see also general comments in Schein (1954a) and the remarks under the genus<i> Coenochilus</i>. It is a new species record for the Cunene Province.
<i> Mecynorhina</i> (<i> Mecynorhina</i>)<i> polyphemus polyphemus&... more <i> Mecynorhina</i> (<i> Mecynorhina</i>)<i> polyphemus polyphemus</i> (Fabricius, 1781)<b> Distribution</b>: ANG, BEN, CAM, CON, DOC, GHA, RCA, RCI, TOG.<b> Distribution in Angola (Provinces):</b> 1) CAB, CNO, LNO.<b> Historic records:</b> — Cabinda (CAB), Cazengo (CNO) ( Paulino de Oliveira 1882 sub<i> Ceratorrhina polyphemus</i> F.); —Dundo (LNO) ( Ferreira 1965 sub<i> Chelorrhina polyphemus</i> F.).<b> Material examined:</b> none.<b> Remarks.</b> There is some controversy on the distribution of this species (see Beinhundner 2017a; Schoolmeesters 2018). It seems represented in the northern territory of Angola ( Paulino de Oliveira 1882; Ferreira 1965). The species is well-known and biological data were recently provided by Šípek (2010) and Christiansen (2013).
<i> Phonotaenia hassoni</i> ( Antoine &amp; Legrand, 2005)<b> Distribution&... more <i> Phonotaenia hassoni</i> ( Antoine &amp; Legrand, 2005)<b> Distribution</b>: ANG, CAM, DOC, GAB, RCA.<b> Distribution in Angola (Provinces):</b> 1) CUS.<b> Historic records:</b> — Kwanza Sul (without locality) (CUS) ( Antoine &amp; Legrand 2005, Antoine 2005a).<b> Material examined:</b> none.<b> Remarks.</b> The species mimics<i> Phonotaenia scalaris scalaris</i> Gory &amp; Percheron, 1833 with which can be mistaken ( Antoine &amp; Legrand 2005). Only the holotype specimen of the typical series is known from Angola. Biological and ecological data are not available for this species.
<i> Dolichostethus angolensis</i> Antoine, 1991 ( Fig. 2i)<b> Distribution</... more <i> Dolichostethus angolensis</i> Antoine, 1991 ( Fig. 2i)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) BGO; 2) LUA.<b> Historic records:</b> —Ambriz ( BENGO) ( Antoine 1991a).<b> Material examined:</b> Mussulo Island ( 8° 53' 9.13" S, 13° 8' 4.62" E, 5m alt., 89) ( LUA), 31.III.1975, 2♂, DO, A. Serrano leg., ASC.<b> Remarks.</b> The species is an endemism of Angola ( Antoine 1991a). It was known only from the Ambriz specimens (typical series). The adult specimens of Mussulo ( Fig. 2i) were collected on flowers (unknown herbaceous plant species). It is a new species record for the Luanda Province.
<i> Pachnoda viridana karinae</i> Antoine, 2000 ( Fig. 6h)<b> Distribution</... more <i> Pachnoda viridana karinae</i> Antoine, 2000 ( Fig. 6h)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) CUS; 2) HUA.<b> Historic records:</b> — Kwanza Sul (without locality) (CUS) ( Antoine 2000).<b> New localities reported:</b> Conda ( 5 km W) (Serra Cumbira) ( 11º 8´ 21.3´´ S, 14º 17´ 27.2´´ E, 856 m alt., 185) ( CUS), 19-20.XI.2017, 2♂, S. Naumann, E. Ott and H. Sulak leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> Huambo ( 12º 46´S, 15º 44´E, 1650 m alt., 256) ( HUA), 1 ♂, IX. 1934, Nº 1984 (blue label), J. Pimentel leg., box nº I-70, MZUC.<b> Remarks.</b> There are four recognized subspecies within<i> P. viridana</i> Blanchard, 1850. This subspecies was described by Antoine (2000) and is endemic of Angola ( Beinhundner 2017a; Schoolmeesters 2018). The two specimens of Conda were captured in a coffee plantation (P. Malec, personal communication). Biological and other ecological data are not available for this subspecies. It is a new species record for the Huambo Province.
<i> Pileotrichius tristiculus</i> ( Kraatz, 1900)<b> Distribution</b>: AN... more <i> Pileotrichius tristiculus</i> ( Kraatz, 1900)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) LUA.<b> Historic records:</b> —Loanda (= Luanda) (LUA) ( Kraatz 1900 and Ferreira 1965 sub<i> Trichius tristiculus</i> Kraatz).<b> Material examined:</b> none.<b> Remarks.</b> The species is endemic of Angola ( Kraatz 1900; Schoolmeesters 2018). Burgeon (1946) stated that the species of this genus are diurnal and flower visitors. Other biological and ecological data are not available.
<i> Stephanorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis tibialis&... more <i> Stephanorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis tibialis</i> ( Waterhouse, 1879) ( Figs. 24g, h)<b> Distribution</b>: ANG, CAM, KEN, UGA.<b> Distribution in Angola (Provinces):</b> 1)?<b> Historic records:</b> — Angola (without locality) [ Waterhouse 1879 sub<i> Cerathorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis</i>)]; — Angola (without locality) ( Ferreira 1965).<b> Material examined:</b> Angola, 1 ♂, Nº 1907 (blue label), nº 47 (white label), old collection, box nº IX-78, MZUC.<b> Remarks.</b> A polytypic species with three recognized subspecies ( Krajcik 2008a; Beinhundner 2017a). It was described originally from Angola and presents a scattered distribution throughout Africa ( Krajcik 2008a; Beinhundner 2017a; Schoolmeesters 2018). The male specimen of MZUC ( Fig. 24g) fits well this species description. Ecological data are not available for this species.
<i> Incala</i> (<i> Incala</i>)<i> calabarina</i> (Westwood, ... more <i> Incala</i> (<i> Incala</i>)<i> calabarina</i> (Westwood, 1873)<b> Distribution</b>: ANG, CON, DOC, NIG, RCA, RCI.<b> Distribution in Angola (Provinces):</b> 1) LNO.<b> Historic records:</b> —Quango (=Cuango) (LNO) ( Quedenfeldt 1884 sub<i> Incala calabariea</i> Westw.).<b> Material examined:</b> none.<b> Remarks.</b> The species is widespread throughout West and Central Africa ( Schoolmeesters 2018). Biological and ecological data are not available.
The effects of land-use management and environmental features at different scales on carabid beet... more The effects of land-use management and environmental features at different scales on carabid beetle diversity and trait structure were assessed across olive groves in northeastern Portugal. We selected organic and integrated olive groves that were distinct in terms of specific management practices, local linear features and landscape configurations. Besides the management intensification levels, differences in carabid diversity and community traits were mainly due to local habitat and ecological linear structures at a finer spatial scale. Carabid community traits related to disturbance, namely traits of body size and species dispersal ability, responded to land-use intensity and particular olive grove features were influencing diversity patterns. Within the olive grove patches, larger and brachypterous species were associated to plots with more dense vegetation cover while macropterous and small-sized species were more associated to open areas. Also, larger carabid species benefitted from higher patch size heterogeneity within the landscape mosaics. Our findings indicate that the effects of farming system is contingent on the specific management practices, local and linear features present in agroecosystems such as olive groves. Particularly, the influence of local features on carabid diversity patterns and community traits linked to dispersal and movement may be crucial in maintaining pest control at a landscape scale.
Genus<i> Dicronorhina</i> Hope, 1837 The genus was revised by Lekkerkerk & Krikken (1... more Genus<i> Dicronorhina</i> Hope, 1837 The genus was revised by Lekkerkerk & Krikken (1986). Beinhundner (2017a) recognized five valid species, two of them recorded for Angola ( Ferreira 1965; Lekkerkerk & Krikken 1986).
<i> Chlorocala smaragdina angolensis</i> ( Schauer, 1941)<b> Distribution</b... more <i> Chlorocala smaragdina angolensis</i> ( Schauer, 1941)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) BGO. Historic records: —Dembos (BGO) ( Schauer 1941 and Ferreira 1965 sub<i> Smaragdesthes smaragdina angolensis</i> Schauer).<b> Material examined:</b> none.<b> Remarks.</b> This is an enigmatic taxon. We did not find any record in Marais &amp; Holm (1992), Sakai &amp; Nagai (1998), Krajcik (2008a) or Beinhundner (2017a). The subspecies was briefly described by Schauer (1941) in a key of the representatives of the genus<i> Smaragdesthes</i>. While Beinhundner (2017a) considers<i> C. smaragdina</i> as a subspecies of<i> C. africana</i>, Schoolmeesters (2018) lists it as a valid species.<i> Chlorocala smaragdina angolensis</i> seems to be an endemic subspecies of Angola ( Schauer 1941), but the validity of the taxon is dubious. The larva of the third instar of<i> C</i>.<i> smaragdina</i> was described by Jerath &amp; Unny (1965). Ecological data are not available for this subspecies.
<i> Leucocelis</i> (<i> Leucocelis</i>)<i> cincticollis</i> M... more <i> Leucocelis</i> (<i> Leucocelis</i>)<i> cincticollis</i> Moser, 1908<b> Distribution</b>: ANG, CON, DOC, TAN.<b> Distribution in Angola (Provinces):</b> 1)?<b> Historic records:</b> — Angola (without locality) ( Beinhundner 2017a).<b> Material examined:</b> none.<b> Remarks.</b> The species was originally described from "Deutsch Ost-Afrika" ( Moser 1908), but curiously Schoolmeesters (2018) refers only East Africa for its distribution. Beinhundner (2017a) indicates Angola in its distribution without reference to any locality. Biological and ecological data are not available for this species.
<i> Tephraea leucomelona</i> ( Gory &amp; Percheron, 1833)<b> Distribution&... more <i> Tephraea leucomelona</i> ( Gory &amp; Percheron, 1833)<b> Distribution</b>: ANG, BOW, MOZ, NAB, RSA, TAN, ZIM.<b> Distribution in Angola (Provinces):</b> 1) BIE, CNN.<b> Historic records:</b> —Bihe (= Bié) (without locality) ( BIE) ( Gory &amp; Percheron 1833); —Merero (CNN) ( Paulino de Oliveira 1882 sub<i> Cetonia leucomelana</i> Gory &amp; Perch.).<b> Material examined:</b> none.<b> Remarks.</b> The species was originally described from Angola ( Gory &amp; Percheron 1833). The larvae instars of<i> T. leucomelona</i> were described by Donaldson (1987) and oviposition by Donaldson &amp; Todner (1992). Biological and ecological data were provided by Holm (1992) and Holm &amp; Marais (1992).
<i>Trechus isabelae</i> Borges &amp; Serrano, new species (Figs. 1–3) <b>Ty... more <i>Trechus isabelae</i> Borges &amp; Serrano, new species (Figs. 1–3) <b>Type material:</b> Holotype, male, Azores, São Jorge, Algar do Morro Pelado (= Algar do Montoso), 5– 11.VII. 2004. (P.A.V. Borges &amp; F. Pereira leg.). Paratypes 1 female, same locality as holotype, 7–11.V. 2004 (P.A.V. Borges &amp; F. Pereira leg.); 2 males and 1 female, same locality as holotype, 5–11.VII. 2004 (P.A.V. Borges &amp; F. Pereira leg.); The Holotype and two paratypes (1 male, 1 female) are deposited at the University of the Azores, Terceira ("Arruda Furtado" Collection); one paratype (female) is deposited at the Faculty of Sciences of the University of Lisbon, and the other one (male) at the University of La Laguna, Dep. of Animal Biology. <b>Additional material:</b> One male was collected in the same locality (7–11.V. 2004) for molecular analysis. <b>Etymology.</b> The species is named after Isabel Amorim who has discovered several new species of <i>Trechus</i> in the Azores. <b>Diagnosis.</b> Large species (4.8–5.38 mm) (Fig. 1), dark brown pigmentation, with some lighter parts; appendages less pigmented (light brown), tibiae and tarsomeres with the same colour patterns as the femora; pronotum as dark as elytra. Eyes slightly convex; 3 rd antennomere longer than 2 nd; pronotum rather cordiform and transverse, with lateral furrows larger than in <i>T. terrabravensis</i>; wingless. <b>Description.</b> Length of holotype: 5.18 mm. Length of female specimens: 4.80–5.38 mm. Large size, convex and ovate. Head almost 1.30 x as wide as long (females: 1.29–1.42), 0.78 x narrower than pronotum (females: 0.74–0.78); eyes well-developed (0.025 mm), moderately convex, diameter of eyes as long as temples; temples with microchaetae. Frons slightly convex, the frontal furrows deeply curved and rounded. Anterior margin of labrum very concave. Maxillary palpi long and slender. Chaetotaxy: two pairs of supraocular setae; four setae in clypeus (outer ones larger); six setae on labrum, the outer ones being the largest. Antennae 2.99 mm long (f [...]
Genus<i> Oplostomus</i> MacLeay, 1838 The genus was not revised. There are ten recogn... more Genus<i> Oplostomus</i> MacLeay, 1838 The genus was not revised. There are ten recognized species ( Holm & Marais 1992, Krajcik 2008a), three of them already recorded for Angola ( Ferreira 1965).
Genus<i> Mausoleopsis</i> Lansberg, 1882 Holm & Marais (1992) and Antoine (1997) refe... more Genus<i> Mausoleopsis</i> Lansberg, 1882 Holm & Marais (1992) and Antoine (1997) refer that the genus is justified itself by the prominently protruding last abdominal spiracles (="élévation des stigmates propygidiaux"). Among the twelve species recognized in the nominotypical subgenus ( Beinhundner 2017a), there are one species and one subspecies recorded for Angola ( Paulino de Oliveira 1882; Antoine 1989a).
<i> Scaptobius kochi</i> Schein, 1957<b> Distribution</b>: ANG.<b> ... more <i> Scaptobius kochi</i> Schein, 1957<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) NAM; 2) HUI.<b> Historic records:</b> —Ongueria (= Namibe) (NAM) ( Schein 1957); —Hunguéria (= Namibe) (NAM) ( Ferreira 1965).<b> New localities reported</b>: Kipungo ( 40 km W) (coord. not available,? alt., 338) ( HUI), 26.XI.2013, 1 ex., P. Schüle leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> none<b> Remarks.</b> The species is endemic of Angola ( Schein 1957). Biological and ecological data are not available. Presumably it presents the same myrmecophilic habits of the other species of this genus. It is a new species record for the Huíla Province.
<i> Gnathocera cruda gracilis</i> (Janson, 1877) ( Fig. 22a)<b> Distribution<... more <i> Gnathocera cruda gracilis</i> (Janson, 1877) ( Fig. 22a)<b> Distribution</b>: ANG, CON, DOC.<b> Distribution in Angola (Provinces):</b> 1) LNO, UIG.<b> Historic records:</b> — Angola (without locality), Bembe (UIG) ( Janson 1877b); —Dundo (LNO) ( Gomes Alves 1959 sub<i> Gnathocera lurida</i> Janson); —Dundo, Chicapa (LNO), Angola (LNO, without locality) ( Antoine 2002a).<b> Material examined:</b> Angola, 2 ♂, Nº 1907 (blue labels), nº 146a (white labels), 1 ♀, Nº 1907 (blue label), nº 147a (white label), 1 ♂, Nº 1907 (blue label), nº 148a (white label), old collection, box nº IX-78, MZUC; Dundo ( 7º 22´S, 20º 50´E, 1470 m alt., 51) ( LNO), -. III.1949, 5 ♂, 4 ♀, Nº 2132, 2 ♀, Nº 2151, Junta de Investigações Coloniais (Entomologia), IICT, MUHNAC.<b> Remarks.</b> The status of this taxon is very controversial. Its validity was established by Antoine (2002a), a criterion followed by Beinhundner (2017a). However, Krajcik (2008a) as well as Schoolmeesters (2018) keep it as a synonym of<i> G. lurida</i> Janson, 1877 [the name<i> lucida</i> used by Krajcik (2008a) is a misspelling of<i> lurida</i>, since Janson (1877b) did not describe any taxon named<i> lucida</i>].<i> Gnathocera cruda gracilis</i> was described based on one specimen from Angola, but Janson (1877b) referred that "there are also specimens in Mr. Higgins's collection found by Monteiro, at Bembe", a municipality within the Uíge Province. The adult specimens of IICT were collected by light (registered information) probably at the same time of<i> G. trivittata ardoini</i> (see remarks under this taxon). Ecological data are not available for this subspecies.
<i> Pseudinca incoides</i> (J. Thomson, 1860)<b> Distribution</b>: ANG, B... more <i> Pseudinca incoides</i> (J. Thomson, 1860)<b> Distribution</b>: ANG, BEN, CAM, CON, DOC, EQG, GAB, GHA, RCA, RCI, NRE, TOG.<b> Distribution in Angola (Provinces):</b> 1) CAB, ZAI.<b> Historic records:</b> — Cabinda (CAB) ( Paulino de Oliveira 1882 sub<i> Diplognatha incoides</i> Thoms.); —S. Salvador (ZAI) ( Valck Lucassen 1933).<b> Material examined:</b> none.<b> Remarks.</b> The species in Angola seems restricted to its northwesternmost territory. Tchibozo<i> et al</i>. (2008) and Joly (2018) provided biodiversity information (abundance, habitat preference) of this species for Benin and Ghana, respectively.
<i> Coenochilus ventricosus</i> (Gyllenhal, 1817) ( Fig. 12f)<b> Distribution&l... more <i> Coenochilus ventricosus</i> (Gyllenhal, 1817) ( Fig. 12f)<b> Distribution</b>: ANG, CAM, DOC, EQG, GAB, GAM, KEN, MOZ, NAB, RCI, RSA, SEN, SLN, TAN, TOG, ZIM.<b> Distribution in Angola (Provinces):</b> 1) LNO, MAL; 2) CNN.<b> Historic records:</b> —Quango (=Cuango) (LNO) ( Quedenfeldt 1884 and Ferreira 1965); —Malange (MAL) ( Schein 1954a).<b> New localities reported</b>: Cahama ( 20 km NE) (coord. not available, 1315 m alt., 399) ( CNN), 8.XII.2013, 1♂, P. Schüle leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> Cacuso ( 9º 25´ 28.03´´S, 15º 44´ 37.63´´ E, 1078m alt., 112) ( MAL), 16–18.XI.2017, 1♂, DO, A. Serrano leg., ASC; Quitota ( 9º 37´ 32.39´´ S, 15º 25´ 22.57´´ E, 898m alt., 129) ( MAL), 20.XI.2017, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC.<b> Remarks.</b> It is the type species of a group with its own name (<i> C. ventricosus</i> group<i> sensu</i> Schein 1954a). The Cacuso adult specimen was found on the ground during night near public lights. Like some<i> C. procerus</i> specimens, the Cacuso specimen had phoretic acarina on some parts of the body (<i> e.g.</i> legs). The Quitota adult specimen was collected by LT (<i> e.g.</i> Fig. 1c) in an open secondary forest. Joly (2018) provided biodiversity information (abundance, habitat preference) on this species for Ghana. Other biological and ecological data are not available for the species, but see also general comments in Schein (1954a) and the remarks under the genus<i> Coenochilus</i>. It is a new species record for the Cunene Province.
<i> Mecynorhina</i> (<i> Mecynorhina</i>)<i> polyphemus polyphemus&... more <i> Mecynorhina</i> (<i> Mecynorhina</i>)<i> polyphemus polyphemus</i> (Fabricius, 1781)<b> Distribution</b>: ANG, BEN, CAM, CON, DOC, GHA, RCA, RCI, TOG.<b> Distribution in Angola (Provinces):</b> 1) CAB, CNO, LNO.<b> Historic records:</b> — Cabinda (CAB), Cazengo (CNO) ( Paulino de Oliveira 1882 sub<i> Ceratorrhina polyphemus</i> F.); —Dundo (LNO) ( Ferreira 1965 sub<i> Chelorrhina polyphemus</i> F.).<b> Material examined:</b> none.<b> Remarks.</b> There is some controversy on the distribution of this species (see Beinhundner 2017a; Schoolmeesters 2018). It seems represented in the northern territory of Angola ( Paulino de Oliveira 1882; Ferreira 1965). The species is well-known and biological data were recently provided by Šípek (2010) and Christiansen (2013).
<i> Phonotaenia hassoni</i> ( Antoine &amp; Legrand, 2005)<b> Distribution&... more <i> Phonotaenia hassoni</i> ( Antoine &amp; Legrand, 2005)<b> Distribution</b>: ANG, CAM, DOC, GAB, RCA.<b> Distribution in Angola (Provinces):</b> 1) CUS.<b> Historic records:</b> — Kwanza Sul (without locality) (CUS) ( Antoine &amp; Legrand 2005, Antoine 2005a).<b> Material examined:</b> none.<b> Remarks.</b> The species mimics<i> Phonotaenia scalaris scalaris</i> Gory &amp; Percheron, 1833 with which can be mistaken ( Antoine &amp; Legrand 2005). Only the holotype specimen of the typical series is known from Angola. Biological and ecological data are not available for this species.
<i> Dolichostethus angolensis</i> Antoine, 1991 ( Fig. 2i)<b> Distribution</... more <i> Dolichostethus angolensis</i> Antoine, 1991 ( Fig. 2i)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) BGO; 2) LUA.<b> Historic records:</b> —Ambriz ( BENGO) ( Antoine 1991a).<b> Material examined:</b> Mussulo Island ( 8° 53' 9.13" S, 13° 8' 4.62" E, 5m alt., 89) ( LUA), 31.III.1975, 2♂, DO, A. Serrano leg., ASC.<b> Remarks.</b> The species is an endemism of Angola ( Antoine 1991a). It was known only from the Ambriz specimens (typical series). The adult specimens of Mussulo ( Fig. 2i) were collected on flowers (unknown herbaceous plant species). It is a new species record for the Luanda Province.
<i> Pachnoda viridana karinae</i> Antoine, 2000 ( Fig. 6h)<b> Distribution</... more <i> Pachnoda viridana karinae</i> Antoine, 2000 ( Fig. 6h)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) CUS; 2) HUA.<b> Historic records:</b> — Kwanza Sul (without locality) (CUS) ( Antoine 2000).<b> New localities reported:</b> Conda ( 5 km W) (Serra Cumbira) ( 11º 8´ 21.3´´ S, 14º 17´ 27.2´´ E, 856 m alt., 185) ( CUS), 19-20.XI.2017, 2♂, S. Naumann, E. Ott and H. Sulak leg., PMC ( P. Malec, personal communication).<b> Material examined:</b> Huambo ( 12º 46´S, 15º 44´E, 1650 m alt., 256) ( HUA), 1 ♂, IX. 1934, Nº 1984 (blue label), J. Pimentel leg., box nº I-70, MZUC.<b> Remarks.</b> There are four recognized subspecies within<i> P. viridana</i> Blanchard, 1850. This subspecies was described by Antoine (2000) and is endemic of Angola ( Beinhundner 2017a; Schoolmeesters 2018). The two specimens of Conda were captured in a coffee plantation (P. Malec, personal communication). Biological and other ecological data are not available for this subspecies. It is a new species record for the Huambo Province.
<i> Pileotrichius tristiculus</i> ( Kraatz, 1900)<b> Distribution</b>: AN... more <i> Pileotrichius tristiculus</i> ( Kraatz, 1900)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) LUA.<b> Historic records:</b> —Loanda (= Luanda) (LUA) ( Kraatz 1900 and Ferreira 1965 sub<i> Trichius tristiculus</i> Kraatz).<b> Material examined:</b> none.<b> Remarks.</b> The species is endemic of Angola ( Kraatz 1900; Schoolmeesters 2018). Burgeon (1946) stated that the species of this genus are diurnal and flower visitors. Other biological and ecological data are not available.
<i> Stephanorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis tibialis&... more <i> Stephanorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis tibialis</i> ( Waterhouse, 1879) ( Figs. 24g, h)<b> Distribution</b>: ANG, CAM, KEN, UGA.<b> Distribution in Angola (Provinces):</b> 1)?<b> Historic records:</b> — Angola (without locality) [ Waterhouse 1879 sub<i> Cerathorrhina</i> (<i> Aphelorrhina</i>)<i> tibialis</i>)]; — Angola (without locality) ( Ferreira 1965).<b> Material examined:</b> Angola, 1 ♂, Nº 1907 (blue label), nº 47 (white label), old collection, box nº IX-78, MZUC.<b> Remarks.</b> A polytypic species with three recognized subspecies ( Krajcik 2008a; Beinhundner 2017a). It was described originally from Angola and presents a scattered distribution throughout Africa ( Krajcik 2008a; Beinhundner 2017a; Schoolmeesters 2018). The male specimen of MZUC ( Fig. 24g) fits well this species description. Ecological data are not available for this species.
<i> Incala</i> (<i> Incala</i>)<i> calabarina</i> (Westwood, ... more <i> Incala</i> (<i> Incala</i>)<i> calabarina</i> (Westwood, 1873)<b> Distribution</b>: ANG, CON, DOC, NIG, RCA, RCI.<b> Distribution in Angola (Provinces):</b> 1) LNO.<b> Historic records:</b> —Quango (=Cuango) (LNO) ( Quedenfeldt 1884 sub<i> Incala calabariea</i> Westw.).<b> Material examined:</b> none.<b> Remarks.</b> The species is widespread throughout West and Central Africa ( Schoolmeesters 2018). Biological and ecological data are not available.
The effects of land-use management and environmental features at different scales on carabid beet... more The effects of land-use management and environmental features at different scales on carabid beetle diversity and trait structure were assessed across olive groves in northeastern Portugal. We selected organic and integrated olive groves that were distinct in terms of specific management practices, local linear features and landscape configurations. Besides the management intensification levels, differences in carabid diversity and community traits were mainly due to local habitat and ecological linear structures at a finer spatial scale. Carabid community traits related to disturbance, namely traits of body size and species dispersal ability, responded to land-use intensity and particular olive grove features were influencing diversity patterns. Within the olive grove patches, larger and brachypterous species were associated to plots with more dense vegetation cover while macropterous and small-sized species were more associated to open areas. Also, larger carabid species benefitted from higher patch size heterogeneity within the landscape mosaics. Our findings indicate that the effects of farming system is contingent on the specific management practices, local and linear features present in agroecosystems such as olive groves. Particularly, the influence of local features on carabid diversity patterns and community traits linked to dispersal and movement may be crucial in maintaining pest control at a landscape scale.
Genus<i> Dicronorhina</i> Hope, 1837 The genus was revised by Lekkerkerk & Krikken (1... more Genus<i> Dicronorhina</i> Hope, 1837 The genus was revised by Lekkerkerk & Krikken (1986). Beinhundner (2017a) recognized five valid species, two of them recorded for Angola ( Ferreira 1965; Lekkerkerk & Krikken 1986).
<i> Chlorocala smaragdina angolensis</i> ( Schauer, 1941)<b> Distribution</b... more <i> Chlorocala smaragdina angolensis</i> ( Schauer, 1941)<b> Distribution</b>: ANG.<b> Distribution in Angola (Provinces):</b> 1) BGO. Historic records: —Dembos (BGO) ( Schauer 1941 and Ferreira 1965 sub<i> Smaragdesthes smaragdina angolensis</i> Schauer).<b> Material examined:</b> none.<b> Remarks.</b> This is an enigmatic taxon. We did not find any record in Marais &amp; Holm (1992), Sakai &amp; Nagai (1998), Krajcik (2008a) or Beinhundner (2017a). The subspecies was briefly described by Schauer (1941) in a key of the representatives of the genus<i> Smaragdesthes</i>. While Beinhundner (2017a) considers<i> C. smaragdina</i> as a subspecies of<i> C. africana</i>, Schoolmeesters (2018) lists it as a valid species.<i> Chlorocala smaragdina angolensis</i> seems to be an endemic subspecies of Angola ( Schauer 1941), but the validity of the taxon is dubious. The larva of the third instar of<i> C</i>.<i> smaragdina</i> was described by Jerath &amp; Unny (1965). Ecological data are not available for this subspecies.
<i> Leucocelis</i> (<i> Leucocelis</i>)<i> cincticollis</i> M... more <i> Leucocelis</i> (<i> Leucocelis</i>)<i> cincticollis</i> Moser, 1908<b> Distribution</b>: ANG, CON, DOC, TAN.<b> Distribution in Angola (Provinces):</b> 1)?<b> Historic records:</b> — Angola (without locality) ( Beinhundner 2017a).<b> Material examined:</b> none.<b> Remarks.</b> The species was originally described from "Deutsch Ost-Afrika" ( Moser 1908), but curiously Schoolmeesters (2018) refers only East Africa for its distribution. Beinhundner (2017a) indicates Angola in its distribution without reference to any locality. Biological and ecological data are not available for this species.
<i> Tephraea leucomelona</i> ( Gory &amp; Percheron, 1833)<b> Distribution&... more <i> Tephraea leucomelona</i> ( Gory &amp; Percheron, 1833)<b> Distribution</b>: ANG, BOW, MOZ, NAB, RSA, TAN, ZIM.<b> Distribution in Angola (Provinces):</b> 1) BIE, CNN.<b> Historic records:</b> —Bihe (= Bié) (without locality) ( BIE) ( Gory &amp; Percheron 1833); —Merero (CNN) ( Paulino de Oliveira 1882 sub<i> Cetonia leucomelana</i> Gory &amp; Perch.).<b> Material examined:</b> none.<b> Remarks.</b> The species was originally described from Angola ( Gory &amp; Percheron 1833). The larvae instars of<i> T. leucomelona</i> were described by Donaldson (1987) and oviposition by Donaldson &amp; Todner (1992). Biological and ecological data were provided by Holm (1992) and Holm &amp; Marais (1992).
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