Papers by Khuat Dang Long
Zootaxa, Sep 21, 2021
The relatively small genus Colastomion Baker, 1917 is newly recorded for Vietnam’s braconid fauna... more The relatively small genus Colastomion Baker, 1917 is newly recorded for Vietnam’s braconid fauna and six new species are described and fully illustrated, viz. Colastomion asperum Long, sp. n; Colastomion cephonodes Long & PVLam, sp. n.; Colastomion citrum Long, sp. n.; Colastomion neuterum Long, sp. n.; Colastomion nigroconfusum Long & Pham, sp. n. and Colastomion vietnamenese Long, sp. n. Based on both females and males, a key to the new species from Vietnam is given. Diagnostic characters of the new species are also analyzed and compared with other Colastomion species from the Oriental region. Additionally, the first host record for the new species, C. cephonodes sp. n., is provided as this new species is reared from caterpillars of the coffee hawkmoth, Cephonodes hylas (Linnaeus, 1771).
Zoologische Mededelingen, 2008
Key to species of the genus <i>Darnilia</i> van Achterberg 1. Length of eye 3.5 × as ... more Key to species of the genus <i>Darnilia</i> van Achterberg 1. Length of eye 3.5 × as long as temple in dorsal view; occipital carina complete, but medio-dorsally weakly developed; basal half of third metasomal tergite distinctly striate medially; hind wing vein 2-SC+R quadrate (Fig. 319 in Chen & He, 1997); Eastern-central China (Jiangsu)............................................ <i>Darnilia chinensis</i> Chen & He, 1997 - Length of eye 5.5–11.0 × as long as temple in dorsal view (Figs 1, 13, 24); occipital carina interrupted medio-dorsally (Figs 13, 24); basal half of third metasomal tergite only finely granulate-coriaceous; hind wing vein 2-SC+R slightly longer than wide (Fig. 22)............................................................................................ 2 2. Eye large, length of eye approx. 11.0 × as long as temple in dorsal view (Fig. 42 in van Achterberg, 1989); first metasomal tergite with median carina in basal 0.3 of tergite (Fig. 4...
<i>Zombrus pallidiventris</i> Cameron, 1910 (Figs 33–47) <i>Zombrus pallidivent... more <i>Zombrus pallidiventris</i> Cameron, 1910 (Figs 33–47) <i>Zombrus pallidiventris</i> Cameron, 1910: 94; Shenefelt &amp; Marsh, 1976: 1370; Fischer, 1983: 340 –342. <b>Material.</b> 1 female (RMNH), "S. Vietnam: Dông Nai, Cát Tiên N.P., Bird trail, Mal. Trap 30–35, c. 100 m, 15– 20.v.2007, C. v. Achterberg &amp; R. de Vries, RMNH'07". <b>Distribution.</b> Indonesia (Kalimantan, * Java [RMNH]) and * Vietnam. * = new record. <b>Notes.</b> The specimen from Java has the pterostigma entirely yellowish as does the female from Vietnam. However, he holotype from Borneo has the pterostigma bicoloured: basally narrowly and apical 0.6 dark brown (see Fig. 47 in Fischer 1983). For the time being, the Bornean, Javanese, and Vietnamese specimens are considered to be colour variation of the same species.
<i>Zombrus tricoloricorpus</i> Long, sp. n. (Figs 22–32) <b>Material.</b>... more <i>Zombrus tricoloricorpus</i> Long, sp. n. (Figs 22–32) <b>Material.</b> Holotype, female, ' Doryc.759 '(IEBR), NW. Vietnam: Phu Tho, Tan Son, Xuan Dai, orchard, MT, 21°07'N, 103°00'E, 120m, 5.v.2009, KDLong. Paratypes: 4 females, 'Doryc.388' (IEBR), NC Vietnam: Nghe An, Con Cuong, Binh Chuan, Pu Huong, 6-9.viii.2005, NT Huong; 'Doryc.603' (RMNH), N. Vietnam: Ha Noi, Gia Lam, Da Ton, M[alaise] T[rap], orchard, 20°59'13''N, 105°55'59''E, 8m, 15-30.viii.2001, KD Long; 'Doryc.616' (VNMN), NE. Vietnam: Vinh Phuc, Me Linh, MT, forest, 21°23'N, 105°42'E, 60m, 4-14.iv.2001, KD Long; 'Doryc.674' (IEBR), NW. Vietnam: Phu Tho, Tan Son, Kiet Son, 21°10'N, 104°57'E, 140m, 10-15.ix.2009, KD Long, NH Thao; 1 male, 'Doryc.603' (IEBR), SC. Vietnam: Quang Nam, Phuoc Son, 15°26'N, 107°53'E, 260m, 19.vii.2009, KD Long. Holotype, female. Body length 8.7 mm; fore wing length 6.0 mm; ovipositor sheath 1.8 mm (Fig. 22). <i>Head</i>. Antenna with 46 segments; scapus 1.55 × as long as its maximum width (14:9); third antennal segment 2.6 × as long as its apical width (13:5), 1.2 × as long as fourth segment (13:11); palpi long, maxillary palp slightly longer than height of head (without mandible) (51:49). In dorsal view, head width 1.9 × its median length (58:31) (Fig. 23); length of head 2.0 × as long as temple (22:11); frons with distinct median protuberance; occipital carina complete; temple roundly narrowed behind eyes; temple rather short in lateral view, transverse diameter of eye 1.5 × length of temple (20:13) (Fig. 24); ocelli small, basal side of ocellar triangle 1.3 × as long as lateral side (4:3); POL 0.8 × as long as OD (4:8) and 0.45 × OOL (5:11) (Fig. 23) (POL:OD:OOL = 4:5:11); in frontal view, eye 1.9 × as high as broad (25:13) (Fig. 25); malar space height 0.6 × height of eye (16:25), 1.3 × as long as basal width of mandible (16:12); face reticulate-punctate (Fig. 25) and its width 1.3 × height of eye (34:26) and 1.1 × height of face and clypeus combined (34:32); width of hypoclypeal depression 0.7 × distance from edge of [...]
Key to Oriental and Palaearctic species of <i>Zombrus</i> Marshall 1 Occipital carina... more Key to Oriental and Palaearctic species of <i>Zombrus</i> Marshall 1 Occipital carina at least latero-dorsally present (Figs 23, 24, 44); bristles of fourth hind tarsal segment 0.5–0.6 × as long as hind telotarsus; subgenus <i>Zombrus</i> Marshall, 1897.............................................................. 7 <b>2</b> - Occipital carina completely absent (Fig. 11); bristles of fourth hind tarsal segment approx. 0.4 × as long as hind telotarsus; [hind coxa and femur with dense and long setae; metasomal tergites 4-6 shiny and entirely smooth (Figs 8, 18)]; subgenus <i>Trichiobracon</i> Cameron, 1905............................................................................. <b>7</b> 2 Legs and palpi black or dark reddish brown; head and mesosoma with long and conspicuous setae; frons with small medial protuberance; middle tarsus not or slightly shorter than tibia; face sparsely punctate to reticulate-punctate; [long s...
<i>WROUGHTONIA</i> CAMERON (based on females) 1. Mandible bent ventrally (fig. 3); ma... more <i>WROUGHTONIA</i> CAMERON (based on females) 1. Mandible bent ventrally (fig. 3); malar suture almost absent; occipital carina faint mediodorsally (fig. 2); vein 2-SC+R hind wing vertical (fig. 13); subgenus <i>Neowroughtonia,</i>................................................................. <i>W. angularis</i> – Mandible evenly curved ventrally (figs. 16, 40, 50, 61); malar suture present; occipital carina complete mediodorsally; vein 2-SC+R of hind wing horizontal (figs. 22, 37, 125), but vertical in <i>W. vietnamica</i> (fig. 135); subgenus <i>Wroughtonia</i> Cameron............2 2. Frontal protuberance between antennal sockets with distinct ledge halfway and blunt dorsally (fig. 127)..............................................................3 – Frontal protuberance between antennal sockets lamelliform, acute dorsally, without ledge (figs. 41, 66)...............................................................10 3. Hind femur comparatively short and robust, 3.2–3.8 <i>×</i> as long as its maximum width (measured without tooth or serrations), and rugose-punctate dorsolaterally (figs. 36, 133)... 4 – Hind femur longer and slenderer, 4.5–5.0 <b>×</b> as long as its maximum width (measured without tooth or serrations), and finely punctate dorsolaterally (figs. 140)...............6 4. Hind femur without ventral tooth-shaped protuberance (fig. 36); second metasomal tergite sculptured (fig. 34)................................................. <i>W. coffeana</i> – Hind femur with ventral tooth-shaped protuberance (figs. 79, 133); second metasomal tergite smooth (figs. 81, 132)..................................................5 5. Ventral tooth-shaped protuberance of hind femur blunt (fig. 133); hind femur 3.2 <b>×</b> longer than its maximum width (measured without tooth); medially second tergite distinctly shorter than third tergite (fig. 132); vein 2-SC+R of hind wing vertical (fig. 135); antennal segments 10–24 cream white. Body length 8.6 mm. Vietnam. <i>W. vietnamica</i> – Ventral tooth-shaped protuberance [...]
<i>Wroughtonia similis</i>, sp. nov. Figures 82–91 TYPE MATERIAL: Holotype, &amp;... more <i>Wroughtonia similis</i>, sp. nov. Figures 82–91 TYPE MATERIAL: Holotype, &amp;female;, "Hel.009" (IEBR), C Vietnam: Thua Thien-Hue, A Luoi, A Roang, 600–900 m, 29.v.2006, H.V. Tru. DISTRIBUTION: C Vietnam: Thua Thien-Hue. BIOLOGY: Unknown. DIAGNOSIS: Antenna with 41 segments; antennal segments 12–23 cream white; maxillary palp 1.7 <b>×</b> as long as head; in frontal view width of face 1.8 <i>×</i> length of face and clypeus combined; frontal protuberance with distinct ledge halfway and blunt dorsally; in dorsal view width of head 1.4 <i>×</i> its median length; height of eye 1.5 <i>×</i> as long as temple; in lateral view height of eye 1.4 <i>×</i> as long as its transverse width; transverse width of eye 1.4 <i>×</i> temple. Mesosoma 2.4 <i>×</i> as long as high; notauli crenulated anteriorly, fused with coarsely rugose area posteriorly; scutellar sulcus 0.5 <i>×</i> as long as scutellum; propodeum with basal median carinae and narrow areola; propodeum almost areolate-rugulose. Length of forewing 3.5 <i>×</i> its maximum width; vein 3-SR 2.0 <i>×</i> vein r; vein 2-M 2.0 <b>×</b> as long as vein 3-SR; hind wing vein 1-M 0.5 <i>×</i> vein 1r-m; vein 2-SC+R long, 0.3 <i>×</i> as long as vein 1r-m; hind wing with 5 hamuli. Foretarsus 1.5 <i>×</i> longer than foretibia; hind femur with ventral serrations and stout protuberance; length of hind femur (without protuberance or serrations) 4.4 <i>×</i> its maximum width; hind coxa sparsely punctate; hind femur largely rugose-punctate. First metasomal tergite 1.8 <i>×</i> as long as its apical width; dorsal carinae in basal 0.6 of tergite; first tergite smooth basally and apically, punctate laterally, punctate-reticulate medially between dorsal carinae; median length of second tergite 0.7 <i>×</i> its basal width, and as long as third tergite; second suture distinct; second tergite smooth with sparse punctures laterobasally; third tergite smooth. NOTES: <i>W. similis,</i> sp. nov., runs to <i>W. eurygenys</i> Yan and Chen, 2017, from China in couplet 3b of the key by Y [...]
<i>Wroughtonia laevis</i>, sp. nov. Figures 59–70 TYPE MATERIAL: Holotype, &amp;f... more <i>Wroughtonia laevis</i>, sp. nov. Figures 59–70 TYPE MATERIAL: Holotype, &amp;female;, "Hel.002" (IEBR), NW Vietnam: Phu Tho, Xuan Son NP, xom Lap, 9.v.2005, PT Nhi. DISTRIBUTION: NW Vietnam: Phu Tho (Xuan Son NP). BIOLOGY: Unknown. DIAGNOSIS: Antenna with 42 segments; 14th–22nd antennal segments cream white; frontal protuberance lamelliform, blunt apically; maxillary palp long, 1.7 <b>×</b> height of head; malar space 0.4 <i>×</i> height of eye; in dorsal view width of head 1.35 <i>×</i> its median length; height of eye 1.2 <i>×</i> temple; in lateral view height of eye 1.75 <i>×</i> its transverse width; transverse width of eye 0.95 <i>×</i> temple. Mesosoma 2.3 <i>×</i> longer than high; notauli narrow, crenulated anteriorly, rugose posteriorly, divided by median posterior carina; scutellar sulcus 0.4 <i>×</i> scutellum; propodeum with basal carina and areola; propodeum rugose-punctate. Length of forewing 3.5 <i>×</i> its maximum width; vein 3-SR 1.45 <i>×</i> vein r; vein 2-M 1.9 <b>×</b> 3-SR; hind wing vein 1-M 0.6 <i>×</i> vein 1r-m; hind wing with 4 hamuli. Foretarsus 1.4 <i>×</i> longer than foretibia; hind femur only with serrations; hind coxa sparsely punctate; hind femur largely rugose-punctate; length of hind femur (without serrations) 4.35× its maximum width. First metasomal tergite 2.0 <i>×</i> longer than its apical width; dorsal carinae in basal 0.75 of tergite; first tergite smooth basally and apically; mediolateral and median area between dorsal carinae of first tergite sparsely foveolate-punctate; median length of second tergite 0.8 <i>×</i> its basal width; second tergite 1.2 <i>×</i> third tergite; second and third tergites shiny and smooth. NOTES: <i>W. laevis,</i> sp. nov., is similar to <i>W. eurygenys</i> Yan and Chen, 2017, from China according to the key by Yan et al. (2017), but differs as follows: (1) first metasomal tergite 2 <i>×</i> as long as its apical width (1.4 <b>×</b> in <i>W. eurygenys</i>); (2) scapus densely setose ventrally (scapus with few medium-sized setae in <i>W. [...]
Figures 77–87. Eodendrus petiolatus sp. nov. (77) Head, frontal view. (78) Head, dorsal view. (79... more Figures 77–87. Eodendrus petiolatus sp. nov. (77) Head, frontal view. (78) Head, dorsal view. (79) Head, lateral view. (80) Basal five segments of antenna. (81) Fore wing. (82) Hind wing. (83) Hind coxa. (84) Hind femur. (85) Hind tibia. (86) First tergite, lateral view. (87) Metasoma, dorsal view.
Figures 66–76. Eodendrus hoabinicus sp. nov. (66) Head, frontal view. (67) Head, dorsal view. (68... more Figures 66–76. Eodendrus hoabinicus sp. nov. (66) Head, frontal view. (67) Head, dorsal view. (68) Head, lateral view. (69) Basal five segments of antenna. (70) Fore wing. (71) Hind wing. (72) Hind coxa. (73) Hind femur. (74) Hind tibia. (75) First tergite, lateral view. (76) Metasoma, dorsal view.
Figures 32–43. Eodendrus elongatus sp. nov. (32) Head, frontal view. (33) Head, dorsal view. (34)... more Figures 32–43. Eodendrus elongatus sp. nov. (32) Head, frontal view. (33) Head, dorsal view. (34) Head, lateral view. (35) Basal and apical segments of antenna. (36) Mesosoma, lateral view. (37) Hind coxa. (38) Hind femur. (39) Hind tibia. (40) Fore wing. (41) Hind wing. (42) First tergite, lateral view. (43) Metasoma, dorsal view.
Figures 21–31. Eodendrus convergens sp. nov. (21) Head, frontal view. (22) Head, dorsal view. (23... more Figures 21–31. Eodendrus convergens sp. nov. (21) Head, frontal view. (22) Head, dorsal view. (23) Head, lateral view. (24) Basal and apical segments of antenna. (25) Fore wing. (26) Hind wing. (27) Hind tibia. (28) Hind coxa. (29) Hind femur. (30) First tergite, lateral view. (31) Metasoma, dorsal view.
FIGURE 14. Austerocardiochiles simulatus, sp. n. Habitus (holotype, female, lateral view).
FIGURES 2–13. Austerocardiochiles melossus, sp. n. (holotype, female) 2. Head (dorsal view), 3. H... more FIGURES 2–13. Austerocardiochiles melossus, sp. n. (holotype, female) 2. Head (dorsal view), 3. Head (frontal view), 4. Head (lateral view), 5. Mesonotum, 6. Mesopleuron, 7. Propodeum, 8. Metasoma, 9. Fore wing, 10. Apex of fore wing, 11. Hind wing hamuli, 12. Hind tarsal claws, 13. Hind wing.
FIGURES 137–141. Wroughtonia cornuta Cameron, 1899, female. 137. Habitus (lateral). 138. Metasoma... more FIGURES 137–141. Wroughtonia cornuta Cameron, 1899, female. 137. Habitus (lateral). 138. Metasoma. 139. Head (lateral) and mesopleuron. 140. Hind femur, lateral. 141. Hind tibia, lateral.
FIGURES 127–136. Wroughtonia vietnamica holotype, female. 127. Head, dorsal. 128. Head, frontal. ... more FIGURES 127–136. Wroughtonia vietnamica holotype, female. 127. Head, dorsal. 128. Head, frontal. 129. Head, lateral. 130. Mesoscutum. 131. Mesopleuron. 132. Metasoma. 133. Hind coxa and femur, lateral. 134. Propodeum. 135. Hind wing. 136. Forewing.
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Papers by Khuat Dang Long