Papers by Idris ABD GHANI
<i>Alyculus malaypeninsularis</i> sp. nov. (Figs. 1–5) <b>Type material.</b&... more <i>Alyculus malaypeninsularis</i> sp. nov. (Figs. 1–5) <b>Type material.</b> Holotype (deposited in UKMB), male, Malaysia, Pahang, Cameron Highlands, Gunung Brinchang, 13.iii.2005, N. Takahashi leg. Paratypes, 1 male (ELKU), same locality and collector as holotype, 13.vii.2003; 1 male (LMBC), 11.iii.2005. 1 male (ELKU), Malaysia, Pahang, Cameron Highlands, Gunung Berembun, 19.i.2006, N. Takahashi leg. 1 male (UKMB), Malaysia, Pahang, Genting Highland, 12.i.2006, N. Takahashi leg. 1 male (UKMB), Malaysia, Pahang, Bukit Fraser, Abu Suradi Trail—Mager Trail, 15.iii.2005, N. Takahashi leg. <b>Diagnosis.</b> Body dark brown. Antennomere 3 slightly compressed and 1.75–2.30 times longer than wide. Pronotum without longitudinal carina on surface. Parameres of aedeagus widest at about basal 3/8 of length and hooked at apices. <b>Description.</b> MALE. Body tiny and elongate (Fig. 1), almost uniform dark brown and covered with brownish pubescence; cranium slightly dark reddish and sometimes faintly lighter around prominent frontal part (Fig. 2); eyes black; pronotum and prosternum feebly dark reddish in some cases; antennomere 2 yellowish (Figs 2–3); pterothoracic ventrites and pleura sometimes faintly lighter; hindwings slightly dusky (Fig. 1); apical portions of tarsomere 5 with claws of legs more or less yellowish. Head (Fig. 2) small, hypognathous, quadrately rounded and widest at eyes in dorsal view; cranium with prominent frontal part, gently convex dorsad though depressed just behind prominent frontal part and slightly flattened behind vertex, lustrous without apparent microstructure and sparsely covered with long pubescence. Eyes with granular facets, large, semiglobularly prominent, shallowly and obtusely emarginate at posterior margin in profile, and diameter of an eye in profile 0.87–1.18 times as large as interocular distance. Antennae (Figs 1–3) inserted anterolaterally on prominent frontal part, moderately robust, densely covered with pubescence though less so on antennomeres 1–2, extending over elytral apice [...]
The <i>krucilense</i> group (new species group) This group is so far represented by a... more The <i>krucilense</i> group (new species group) This group is so far represented by a single species, <i>T. krucilense</i>, known only from males. Forewing not extensively covered with small spots (Fig. 1U). Eighth sternum with single sclerite separated from hypandrium (Fig. 29 AC). Epiproct chair-shaped, epiproct lobe long, protruding over clunium (Fig. 29 AB). Hypandrium symmetrical, median tongue absent (Fig. 29C). Phallosome closed apically, with pseudoparameres (Fig. 29D).
<i>Trichadenotecnum pardus</i> Badonnel (Fig 1E) <i>Trichadenotecnum pardus<... more <i>Trichadenotecnum pardus</i> Badonnel (Fig 1E) <i>Trichadenotecnum pardus</i> Badonnel, 1955: 231. <i>Trichadenotecnum pardidum</i>: New, 1975: 258. [Misidentification] [Not <i>Trichadenotecnum pardidum</i> Thornton, 1961: 16.] <b>Specimen examined</b>. 1 female, Singapore, Botanic Gardens, Rain Forest, beating, 25m, 16.xii.1987, CL (MHNG). <b>Remarks</b>. See Yoshizawa (2001) and Yoshizawa &amp; Lienhard (2004) for diagnosis of this species.
FIGURE 2. Male terminalia of Trichadenotecnum cinnamonum. A: terminalia, lateral view; B: epiproc... more FIGURE 2. Male terminalia of Trichadenotecnum cinnamonum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
FIGURE 4. Zorotypus impolitus sp. n.: A: abdomen of male, ventral view; B: terminalia of male, po... more FIGURE 4. Zorotypus impolitus sp. n.: A: abdomen of male, ventral view; B: terminalia of male, posterior view; C: terminalia of male, posterolateral view; D: abdomen of female, ventral view; E: terminalia of female, posterior view, as for the asterisk and stars, see the description. S 8, 9 and 11, abdominal sterna 8, 9 and 11; T 8 - 11, abdominal terga 8 to 11.
FIGURE 3. Zorotypus impolitus sp. n.: A: head of apteron male, dorsal view; B: thorax of apteron ... more FIGURE 3. Zorotypus impolitus sp. n.: A: head of apteron male, dorsal view; B: thorax of apteron male, dorsal view; C: thorax of alate female, dorsal view; D: abdomen of male, dorsal view, as for the asterisk and star, see the description.
FIGURE 7. Zorotypus cervicornis sp. n.: A: antenna; B: left mandible, anterior view; B': righ... more FIGURE 7. Zorotypus cervicornis sp. n.: A: antenna; B: left mandible, anterior view; B': right mandible, anterior view; C: right metaleg, anterior view; D: male genitalia, posterior to the bottom; E: abdominal sternum 9 of female. AH, aedeagus hook; BP, basal plate; F, flagellum.
FIGURE 6. Zorotypus magnicaudelli sp. n.: A: antenna; B: left mandible, anterior view; B': ri... more FIGURE 6. Zorotypus magnicaudelli sp. n.: A: antenna; B: left mandible, anterior view; B': right mandible, anterior view; C: forewing; C': hindwing; D: right metaleg, anterior view; E: male genitalia, posterior to the bottom. AH, aedeagus hook; BP, basal plate; F, flagellum.
FIGURE 5. Zorotypus caudelli: A: antenna; B: left mandible, anterior view; B': right mandible... more FIGURE 5. Zorotypus caudelli: A: antenna; B: left mandible, anterior view; B': right mandible, anterior view; C: wings; D: right metaleg, anterior view; E: male genitalia, posterior to the bottom. AH, aedeagus hook; BP, basal plate; F, flagellum.
FIGURE 2. Zorotypus caudelli: A: abdomen of male, dorsal view; B: terminalia of male, posterior v... more FIGURE 2. Zorotypus caudelli: A: abdomen of male, dorsal view; B: terminalia of male, posterior view; C: terminalia of female, posterior view; D: abdomen of male, ventral view; E: abdomen of female, ventral view; F: abdominal sternum 9 of female. S 9, 11, abdominal sterna 9 and 11; T 9 - 11, abdominal terga 9 to 11.
<i>Trichadenotecnum yatai</i> Yoshizawa &amp; Lienhard, sp.n. (Figs 1F, 6, 7) <... more <i>Trichadenotecnum yatai</i> Yoshizawa &amp; Lienhard, sp.n. (Figs 1F, 6, 7) <b>Holotype.</b> Male, Malaysia, Gunung Jasar, Cameron Highland, Pahang, 9.iii.2003, K. Yoshizawa (UKM). <b>Paratypes</b>: Malaysia, 3 females, same data as holotype (UKM); 1 male, Gunung Berembun, Cameron Highland, Pahang, 10.iii.2003, NT (UKM); 1 male 3 females, type locality, 14.iii.2003, KY (SEHU &amp; MHNG); 1 male 2 females, same locality, 14.vii.2003, KY (SEHU &amp; MHNG). <b>Description</b>. <b>Male</b>. <b>Head.</b> Yellowish white in ground color; vertical markings blackish brown, each marking touching with neighbors; with pair of blackish brown markings anterior to vertical markings; orbital markings blackish brown; coronal suture black; epicranial suture blackish brown, laterally with brackish brown marking dorsally; frons with central pair of brown bands reaching to oceller region dorsally, and with lateral pair of blackish brown broader band not reaching to epicranial suture dorsally, central and lateral bands fused with each other ventrally; eye dark gray, IO/D = 0.8; ocelli white, ocellar field black; gena with transversal blackish brown band medially, ventral and eye margins blackish brown; postclypeus mostly blackish brown ventrally except ventrolateral corner white, dorsal region white with four rows of blackish brown spots; anteclypeus blackish brown. Antenna brown, scape and pedicel blackish brown. Mouthparts brown. <b>Thorax</b>. Prothorax blackish brown. Mesonotum mostly brown, scutum with yellowish white bands along posterolateral margins of anterior lobe, at middle from anterior to posterior ends of scutum, and along anterior margin of lateral lobe. Metanotum brown, median part and anterior region of lateral lobe of scutum yellowish white. Meso- and metapleuron brown except membranous regions. <b>Legs</b>. Mostly blackish brown, ventral surface of distal half of fore femur and tip of hind femur white, tibiae paler. <b>Forewing</b> (Fig. 1F). Extensively and densely covered with tiny spots. Distal spot in cell a1 [...]
FIGURE 8. Male terminalia of Trichadenotecnum kojimai. A: terminalia, lateral view; B: epiproct, ... more FIGURE 8. Male terminalia of Trichadenotecnum kojimai. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: ep... more FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
<i>Trichadenotecnum krucilense</i> Endang, Thornton &amp; New (Figs 1U, 29) <i... more <i>Trichadenotecnum krucilense</i> Endang, Thornton &amp; New (Figs 1U, 29) <i>Trichadenotecnum krucilense</i> Endang, Thornton &amp; New, 2002: 165; Endang &amp; New, 2005: 23. <b>Specimen examined.</b> [Malaysia] 1 male, Gnung Berembun, Path 3, Cameron Highlands, Pahang, 14.vii.2003, HK.etal (canopy fogging, St. 6). <b>Remarks</b>. The species was originally described from Java, Indonesia, later also recorded from Sumatra, Indonesia, and is here recorded from Peninsula Malaysia for the first time. The variation of the hypandrial structure, three (Javanese) or four (Sumatran) hypandrial spikes, has been pointed out by Endang &amp; New (2005). The presently examined specimen has four hypandrial spikes. Judging from the original hypandrial illustration published by Endang <i>et al.</i> (2002: fig. 333), missing of some hypandrial parts on the left side of the specimen seems evident; on that side only one spike is figured whereas two on the other side. It is likely that the variation pointed out by Endang &amp; New (2005) is actually an artifact caused by dissection or preparation. The hypandrium of this species (Fig. 29C) is unique, very different from that of any other known species. Absence of the hypandrial median tongue may indicate its close affinity with <i>T. apertum</i> and its close relatives. In the original description of this species, the phallosome is illustrated in highly deformed condition, but the present examination clearly shows that this species has the pseudo-parameres on the phallosome (Fig. 29D). The presence of this structure is a synapomorphy of the <i>sexpunctatum</i>, <i>medium</i> and New World species groups (see Yoshizawa <i>et al</i>., 2008) so that assignment of <i>T. krucilense</i> into the <i>majus</i> group is excluded. However, this species cannot be assigned to the <i>sexpunctatum</i>, <i>medium</i> or New World species groups because of lack of apomorphies characterizing any of these species group so that a new species group for its own is proposed here. Females of this species [...]
<i>Trichadenotecnum anomalum</i> Yoshizawa &amp; Lienhard, sp.n. (Figs 1R, 25) &l... more <i>Trichadenotecnum anomalum</i> Yoshizawa &amp; Lienhard, sp.n. (Figs 1R, 25) <b>Holotype.</b> Male, Singapore, Bukit Timah forest, 24 hr survey, section trap 5–9 am, 26.x.1969, D.H. Murphy (MHNG). <b>Description</b> (note that the condition of the specimen is bad). <b>Male</b>. <b>Head.</b> Yellowish in ground color; vertical markings and orbital markings obscure; coronal suture black; epicranial suture faintly bordered with brown band dorsally; frons with faint.markings; eye black, IO/D= 1.1; ocelli white, ocellar field brown; gena mostly yellowish; postclypeus with seven rows of spots dorsomedially; anteclypeus brown. Antenna pale brown. Mouthparts brown. <b>Thorax</b>. Prothorax brown. Mesonotum yellowish, lateral lobe of scutum with faint marking, anterolateral corners of scutellum brown. Metanotum yellowish, lateral lobe of scutum with faint marking, lateral corner of scutellum brown. Meso- and metapleuron pale brown except for white membranous regions. <b>Legs</b>. Mostly pale brown, foretibia and tarsi of all legs darker. <b>Forewing</b> (Fig. 1R). Hyaline, lacking almost all marking elements characteristics for <i>Trichadenotecnum</i> except following: opposing spots in cell r distinct, fused to single marking; basal band only distinct around Rs-M fusion and in cell cua. <b>Terminalia</b>. 8th sternum with pair of sclerotized portions (Fig. 25 AC). Clunium (Fig. 25A) with broad lateral arms, about parallel sided and with strong dorsal expansion apically. Epiproct (Fig. 25 AB) strongly expanded dorsally and anteriorly over clunium, posteriorly with small conical projection medially; dorsal margin of epiproct lobe broad and posterior surface convex. Paraproct (Fig. 25A) with small distal lobe and with short and almost straight distal process directed posteriorly. Hypandrium (Fig. 25C) asymmetrical, without median tongue but with tongue-shaped projection medially near anterior margin, anterolaterally with pair of areas densely covered with tiny denticles, distally with two processes, one arises medially (= [...]
FIGURE 27. Male terminalia of Trichadenotecnum kerinciense. A: terminalia, lateral view; B: epipr... more FIGURE 27. Male terminalia of Trichadenotecnum kerinciense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
FIGURE 17. Female genitalia of Trichadenotecnum malayense, ventral view. A: subgenital plate; B: ... more FIGURE 17. Female genitalia of Trichadenotecnum malayense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate.
FIGURE 1.2. Forewings of Trichadenotecnum spp.: N: T. sibolangitense, male; O: T. santosai, femal... more FIGURE 1.2. Forewings of Trichadenotecnum spp.: N: T. sibolangitense, male; O: T. santosai, female; P: T. soenarti, male; Q: T. cornutum, male; R: T. anomalum, holotype male; S: T. rachimi, female; T: T. kerinciense, female; U: T. krucilense, male.
<i>Trichadenotecnum sibolangitense</i> Endang &amp; New (Figs 1N, 18, 19) <i&g... more <i>Trichadenotecnum sibolangitense</i> Endang &amp; New (Figs 1N, 18, 19) <i>Trichadenotecnum sibolangitense</i> Endang &amp; New, 2005: 27. <b>Specimens examined</b>. [Malaysia] 1 female, Gunung Jasar, Cameron Highlands, Pahang, 14.iii.2003, KY; 1 male (KY345) Gunung Berembun, Cameron Highlands, Pahang, 10.iii.2005, NT. [Singapore] 1 female, Botanic Gardens, Forest (low foliage), 6.xi.1981, DHM (MHNG). <b>Description of female genitalia</b>. Egg guide (Fig. 19A) short, shorter than basal width, ventrally membranous and dorsally widely sclerotized, distal margin concave; body of subgenital plate broadly sclerotized, anteromedially with wide membranous region, posterior margin with pair of long, narrow and sharply pointed lateral processes. Gonapophyses (Fig. 19B). Ventral valve not exceeding external valve; dorsal valve broad, with deep pouch posteromedially, distal process long; external valve with broad membranous region anteriorly and with flap-like expansion posteriorly, posterior lobe broad, internal lobe narrow. Spermapore plate (Fig. 19C) symmetrical, pigmented around spermapore. <b>Remarks</b>. This species was originally described from Sumatra, Indonesia based on a single male and is here recorded from Peninsula Malaysia for the first time. The male specimens examined here agree exactly with the original description except for the hypandrial structures being in mirror image (Fig. 18C). Given the exact similarity of each structure, we concluded that the illustration of the hypandrium of this species presented in Endang &amp; New (2005) is drawn in dorsal (internal) view or reproduced in reversed position. This species is apparently closely related to <i>T. alinguum</i> Endang, Thornton &amp; New, 2005 and, in <i>T. sibolangitense</i>, the right side of the hypandrium is strongly expanded, as illustrated in Fig. 18C. Given the lack of the hypandrial median tongue and the triangular shape of the hypandrium, this species is considered to be closely related to <i>T. arciforme</i> (Thornton, 1961; Yoshizawa &a [...]
<i>Trichadenotecnum endangae</i> Yoshizawa &amp; Lienhard, sp. n. (Figs 1I, 11, 1... more <i>Trichadenotecnum endangae</i> Yoshizawa &amp; Lienhard, sp. n. (Figs 1I, 11, 12) <b>Holotype.</b> Male, Malaysia, Gunung Brinchang, Cameron Highlands, Pahang, 11.iii.2003, K. Yoshizawa (UKM). <b>Paratype.</b> 1 female, Malaysia, Gunung Jasar, Cameron Highlands, Pahang, 9.iii.2003, KY (UKM). <b>Description</b>. <b>Male</b>. <b>Head.</b> Yellowish white in ground color; vertical markings blackish brown, each marking fused with neighbors; with pair of blackish brown markings anterior to vertical markings; orbital markings blackish brown; coronal suture black; epicranial suture broadly bordered with blackish brown band dorsally; frons mostly blackish brown, with pair of dorsal white spots and pair of lateral white bands; eye black, IO/D= 1.3; ocelli white, ocellar field black; gena mostly blackish brown; postclypeus mostly blackish brown, dorsally with six white bands; anteclypeus blackish brown. Antenna brown, scape and pedicel blackish brown. Mouthparts brown. <b>Thorax</b>. Prothorax blackish brown. Mesonotum blackish brown, medially with trifurcated white marking. Metanotum blackish brown, medially with white spot. Meso- and metapleuron brown except for white membranous regions. <b>Legs</b>. Mostly blackish brown, distal half of ventral surface of fore femur and distal tip of hind femur white, tibiae paler. <b>Forewing</b> (Fig. 1I). Extensively covered with tiny spots, dense basally from basal band, sparse distally from basal band. Distal spot in cell a1 distinct, basal spots obscure. Opposing spots in cell r obscure. Basal band narrow but distinct, broadly interrupted above M-Cu fork, narrowly interrupted above CuP vein. Distal band narrow but distinct. Spot on roof of cell m3 faint, only distinct at top of areola postica. Submarginal spots distinct in cells r3 to m2, obscure in other cells. Marginal clouds almost absent. Hindwing hyaline; veins brown. <b>Abdomen</b> dark brown, with irregular white spots. <b>Terminalia</b>. 8th sternum with single short triangular sclerite fused to hypandrium (Fig. 11 AC). [...]
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