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Orca lifespan: I fixed the source as the original one I provided is no longer in service. This is still the same research paper by the same authors.
 
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'''Granny''' (born c. 1936-1951 or earlierapprox, died c. 2016), also known as '''J2''', was a female [[orca]] of the J pod of [[southern resident orcas]] notable for her long life. Early estimates placed her birth in 1911, putting her at 105 years old at the time of her death. However, this estimate was later revealedtheorized to have been based on mistaken information and more recent studies put her at 65-8065–80 years old.<ref>{{cite web|url=https://abcnews.go.com/US/wireStory/oldest-puget-sound-orca-granny-missing-presumed-dead-44530922|title=Oldest Puget Sound Orca, 'Granny,' Missing and Presumed Dead |publisher= ABC News|website= abcnews.go.com|access-date=January 3, 2017}}</ref><ref name=King>{{Cite news|url=http://www.king5.com/news/local/oldest-southern-resident-killer-whale-considered-dead/381349614|title=Oldest Southern Resident killer whale considered dead| last=Tegna| website= KING5.com|language=en-US|access-date=January 3, 2017}}</ref><ref name="Orcazine">{{cite web|url=http://orcazine.com/granny-j2/|title=Orca Granny: was she really 105?| last=Podt| first=Annemieke|date=December 31, 2016 |language=en-US|access-date=September 11, 2017}}</ref> If she was 105, she would have been the oldest known orca at the time of her death.<ref>{{cite web|last=Bender |first=Kelli |url=http://www.people.com/people/mobile/article/0,,20817101,00.html |title=Granny, World's Oldest Orca, Returns to Home Waters at 103 Years Old : People.com Mobile |website= [[People (magazine)|People.com]] |access-date=May 16, 2014}}</ref><ref name="theprovince1">{{cite web |last=Luba |first=Frank |url=https://theprovince.com/news/Great+grandmother+whale+still+going+strong+delight+experts/9828660/story.html |title=B.C.'s matriarch orca 'Granny' is still going strong at 103 |publisher=Theprovince.com |date=May 12, 2014 |access-date=May 16, 2014 |archive-date=May 15, 2014 |archive-url=https://web.archive.org/web/20140515194710/http://www.theprovince.com/news/Great+grandmother+whale+still+going+strong+delight+experts/9828660/story.html |url-status=dead }}</ref><ref>{{cite web |last=Ellison |first=Jake |url=http://blog.seattlepi.com/bigscience/2014/05/12/oldest-living-orca-granny-visits-nw-over-mothers-day-weekend/#9102105=0&9399107=0&12348103=0&23307101=0 |title=Oldest living orca 'Granny' visits NW over Mother's Day weekend – The Big Science Blog |publisher=Blog.seattlepi.com |date=May 12, 2014 |access-date=May 16, 2014 |archive-date=December 12, 2016 |archive-url=https://web.archive.org/web/20161212002903/http://blog.seattlepi.com/bigscience/2014/05/12/oldest-living-orca-granny-visits-nw-over-mothers-day-weekend/#9102105=0&9399107=0&12348103=0&23307101=0 |url-status=dead }}</ref><ref>{{cite web|url=http://www.cbc.ca/news/canada/british-columbia/orca-granny-103-comes-home-for-mother-s-day-1.2641231 |title=Orca 'Granny,' 103, comes home for Mother's Day – British Columbia |publisher= CBC News |website= CBC.ca |access-date=May 16, 2014}}</ref> Granny lived in the northeast [[Pacific Ocean]] and coastal bays of [[Washington (state)|Washington state]] and [[British Columbia]]. She was last seen on October 12, 2016, and was considered deceased by The Center for Whale Research in January 2017.<ref>{{Cite web|url=http://www.whaleresearch.com/j2|title=Center for Whale Research|website=Center for Whale Research|access-date=January 3, 2017}}</ref><ref name=King />
 
==Description==
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==J pod==
Granny, along with several of her descendants, travelled in the J pod, a group of about 25 orcas.<ref name=Genealogy>{{cite book | url=https://books.google.com/books?id=D8cw6_K4MHQC&pg=PA84 | title=Killer Whales: The Natural History and Genealogy of Orcinus Orca in British Columbia| isbn=9780774844307| last1=Ford| first1=John K. B.| last2=Ellis| first2=Graeme M.| last3=Balcomb| first3=Kenneth C.| year=1996| publisher=UBC Press}}</ref> J pod, along with Pods K and L, are the "J clan", which constitute the entire southern resident population. They frequent the inland waters of British Columbia and Washington state in the summer months, but roam from southeast [[Alaska]] to central [[California]]. They have completed a journey as far as {{convert|800|mi}} in a week.<ref name=Genealogy/> As the oldest female in J pod, Granny would have been considered its [[matriarch#Animals|matriarch]].<ref name="OrcaAdventures">{{cite web|url=https://orcaspirit.com/2012/05/the-resident-orcas-of-j-pod/|title=The Resident Orcas Of J-Pod|first= John| last= Douglas|date=2012}}</ref>
 
A well-known male orca previously thought to be Granny's son is J1 Ruffles. He was last seen in late 2010. As of 2012, none of Granny's immediate children are known to be living.<ref name="OrcaAdventures"/> However, Granny had multiple grandchildren and great-grandchildren who travelled in the pod with her.<ref name=Genealogy/>
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==Legacy==
As one of the longest living known orcas, Granny is employed as an example in arguments against keeping orcas in captivity, referencing the allegedly reduced lifespan of captive animals.<ref name="DeathAtSeaworld">{{cite book | url=https://books.google.com/books?id=N0JHLOQKomAC&pg=PA462 | title=Death at SeaWorld: Shamu and the Dark Side of Killer Whales in Captivity | author=Kirby, David| date=July 17, 2012 | publisher=Macmillan | isbn=9781250002020 }}</ref> As of 2023, the oldest [[captive orca]] is [[Corky (killer whale)|Corky]], who has been in captivity since 1969. The average lifespan for a captive orca is 20 to 30 years.<ref>{{cite news| work= [[Calgary Herald]]| agency= [[Times Colonist]]| first= Sarah| last= Petrescu| url= https://calgaryherald.com/news/national/worlds-oldest-known-killer-whale-seen-off-b-c-coast| title= World's oldest known killer whale seen off B.C. coast| date= May 14, 2014| access-date= January 5, 2017}}</ref>
 
Granny was also used as a focal point of environmental efforts that resulted in the Endangered Species Act protections for orca. Environmentalists estimate that Granny may have had a [[Polychlorinated biphenyl|PCB]] load of up to 100 parts per million, and that her descendants' reproductive systems may have been damaged by exposure to pollution.<ref name=Lyke6514>{{Cite webnews
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==Orca lifespan==
{{main|Orca#Life cycle}}
Estimates of lifespans for wild orcas vary. [[SeaWorld]] says wild lifespans are 30–50 years for females, and 19–30 years for males.<ref>{{cite web|url=http://seaworldcares.com/killer-whales/lifespan/|title=Lifespan|work=SeaWorld Cares}}</ref> These estimatesThis departis fromconsistent thewith findingsa 2021 study of athe 2005lifespans study,of whichwild peggednorth thePacific meankiller agewhales, ofwith resident females atliving 45.8from 34-52 years and malesmale atresidents 31.0living duringfrom the19-34 periodyears between 1973 andon 1996average.<ref>{{cite web |first1=Peter FM.L.K. |last1=OlesiukNielsen |first2=Graeme MS. |last2=Ellis |first3=John KJ.BR. |last3=FordTowers |title=LifeA Historylong andpost-reproductive Populationlifespan Dynamicsis ofa Northernshared Residenttrait Killeramong Whalesgenetically (Orcinusdistinct orca)killer inwhale British Columbiapopulations |url=httphttps://wwwonlinelibrary.dfo-mpo.gcwiley.cacom/CSASdoi/Csasfull/DocREC10.1002/2005/RES2005_045_eece3.pdf |date7756#:~:text=2005 |publisher=Canadian Science Advisory Secretariat}}</ref> Marine conservation groups argue that even these estimates are low due to the effects of huntingWe%20show%20that%20three%20sympatric%20killer%20whale, pollution, and capture on the wild populations, and that natural wild orca lifespans are equivalent to that of humans, with male orcas living up to 75 years and female orcas living up to 80 yearshave%20comparably%20long%20postreproductive%20female%20life%20spans.<ref>{{cite web|url&text=http://www.orcanetwork.org/Main/index.php?categories_file=OrcaThis%20is%20the%20Lifespans20first%20evidence%20showing%20a,might%20be%20present%20in%20other%20discrete%20populations.|titledate=Welcome2021 to Orca Network|authorpublisher=HowardEcology and GarrettEvolution}}</ref>
 
These estimates depart from the findings of a 2005 study, which pegged the mean age of females at 45.8 years and males at 31.0 during the period between 1973 and 1996.<ref>{{cite web |first1=Peter F. |last1=Olesiuk |first2=Graeme M. |last2=Ellis |first3=John K.B. |last3=Ford |title=Life History and Population Dynamics of Northern Resident Killer Whales (Orcinus orca) in British Columbia |url=http://www.dfo-mpo.gc.ca/CSAS/Csas/DocREC/2005/RES2005_045_e.pdf |date=2005 |publisher=Canadian Science Advisory Secretariat}}</ref> Marine conservation groups argue that even these estimates are low due to the effects of hunting, pollution, and capture on the wild populations, and that natural wild orca lifespans are equivalent to that of humans, with male orcas living up to 75 years and female orcas living up to 80 years.{{Citation needed|date=September 2023}}
 
Orcas are one of the few species to exhibit [[Menopause#Other_animals|menopause]] and Granny's great age gave her a chance to use her skills to enhance the reproductive success of her offspring (see [[Grandmother hypothesis]]).{{Citation needed|date=June 2023}}